Sir Francis Darwin
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Sir Francis Darwin (16 August 1848 – 19 September 1925) was a British
botanist Botany, also called plant science, is the branch of natural science and biology studying plants, especially Plant anatomy, their anatomy, Plant taxonomy, taxonomy, and Plant ecology, ecology. A botanist or plant scientist is a scientist who s ...
. He was the third son of the naturalist and scientist
Charles Darwin Charles Robert Darwin ( ; 12 February 1809 – 19 April 1882) was an English Natural history#Before 1900, naturalist, geologist, and biologist, widely known for his contributions to evolutionary biology. His proposition that all speci ...
.


Biography

Francis Darwin was born at
Down House Down House is the former home of the English Natural history, naturalist Charles Darwin and his family. It was in this house and garden that Darwin worked on his theory of evolution by natural selection, which he had conceived in London befor ...
,
Downe Downe, formerly Down (), is a village in Greater London, England, located within the London Borough of Bromley, which formed part of the historical county of Kent until 1965, and is beyond London's contiguous urban area. Charles Darwin lived ...
,
Kent Kent is a Ceremonial counties of England, ceremonial county in South East England. It is bordered by Essex across the Thames Estuary to the north, the Strait of Dover to the south-east, East Sussex to the south-west, Surrey to the west, and Gr ...
in 1848. He was the third son and seventh child of Charles Darwin and his wife
Emma Wedgwood Emma Darwin (; 2 May 1808 – 2 October 1896) was an English woman who was the cousin marriage, wife and first cousin of Charles Darwin. They were married on 29 January 1839 and were the parents of ten children, seven of whom survived to adulth ...
. He was educated at
Clapham Clapham () is a district in south London, south west London, England, lying mostly within the London Borough of Lambeth, but with some areas (including Clapham Common) extending into the neighbouring London Borough of Wandsworth. History Ea ...
Grammar School. He then went to
Trinity College, Cambridge Trinity College is a Colleges of the University of Cambridge, constituent college of the University of Cambridge. Founded in 1546 by King Henry VIII, Trinity is one of the largest Cambridge colleges, with the largest financial endowment of any ...
, initially studying
mathematics Mathematics is a field of study that discovers and organizes methods, Mathematical theory, theories and theorems that are developed and Mathematical proof, proved for the needs of empirical sciences and mathematics itself. There are many ar ...
, then changing to
natural science Natural science or empirical science is one of the branches of science concerned with the description, understanding and prediction of natural phenomena, based on empirical evidence from observation and experimentation. Mechanisms such as peer ...
s, graduating in 1870. He then went to study medicine at St George's Medical School, London, earning an MB in 1875, but did not practice medicine. Darwin was married three times and widowed twice. First, he married Amy Richenda Ruck in 1874, but she died in 1876 four days after the birth of their son
Bernard Darwin Bernard Richard Meirion Darwin CBE JP (7 September 1876 − 18 October 1961) was a golf writer and high-standard amateur golfer. A grandson of the British naturalist Charles Darwin, he was inducted into the World Golf Hall of Fame. Biography B ...
, who was later to become a golf writer. In September 1883, he married Ellen Wordsworth Crofts (1856–1903) and they had a daughter Frances Crofts Darwin (1886–1960), a poet who married the poet
Francis Cornford Francis Macdonald Cornford (27 February 1874 – 3 January 1943) was an English classical scholar and translator known for work on ancient philosophy, notably Plato, Parmenides, Thucydides, and ancient Greek religion. Frances Cornford, his w ...
and became known under her married name. His third wife was Florence Henrietta Fisher, daughter of
Herbert William Fisher Herbert William Fisher (30 July 1826 – 17 January 1903) was a British historian, best known for his book ''Considerations on the Origin of the American War'' (1865). Life He was born at Poulshot, Wiltshire, the eldest son of Rev. William Fis ...
and widow of
Frederic William Maitland Frederic William Maitland (28 May 1850 – ) was an English historian and jurist who is regarded as the modern father of English legal history. From 1884 until his death in 1906, he was reader in English law, then Downing Professor of the Laws ...
, whom he married in 1913, the year in which he was
knighted A knight is a person granted an honorary title of a knighthood by a head of state (including the pope) or representative for service to the monarch, the church, or the country, especially in a military capacity. The concept of a knighthood ...
. Her sister Adeline Fisher was the first wife of Darwin's double first cousin once removed
Ralph Vaughan Williams Ralph Vaughan Williams ( ; 12 October 1872– 26 August 1958) was an English composer. His works include operas, ballets, chamber music, secular and religious vocal pieces and orchestral compositions including nine symphonies, written over ...
. Francis Darwin worked with his father on experiments dealing with
plant movements Plant physiology is a subdiscipline of botany concerned with the functioning, or physiology, of plants. Plant physiologists study fundamental processes of plants, such as photosynthesis, respiration, plant nutrition, plant hormone functions, tropi ...
, specifically
phototropism In biology, phototropism is the growth of an organism in response to a light stimulus. Phototropism is most often observed in plants, but can also occur in other organisms such as fungi. The cells on the plant that are farthest from the ligh ...
. They co-authored ''
The Power of Movement in Plants ''The Power of Movement in Plants'' is a book by Charles Darwin on phototropism and other types of movement in plants. This book continues his work in producing evidence for his theory of natural selection. As it was one of his last books, fol ...
'' (1880) and Francis Darwin published a second expanded edition of '' Insectivorous Plants'' (1888) after his father's death. Their experiments showed that the
coleoptile Coleoptile is the pointed protective sheath covering the emerging shoot in monocotyledons such as grasses in which few leaf primordia and shoot apex of monocot embryo remain enclosed. The coleoptile protects the first leaf as well as the growing ...
of a young grass seedling directs its growth toward the light by comparing the responses of seedlings with covered and uncovered coleoptiles. These observations would later lead to the discovery of
auxin Auxins (plural of auxin ) are a class of plant hormones (or plant-growth regulators) with some morphogen-like characteristics. Auxins play a cardinal role in coordination of many growth and behavioral processes in plant life cycles and are essent ...
. Darwin was nominated by his father to the
Linnean Society of London The Linnean Society of London is a learned society dedicated to the study and dissemination of information concerning natural history, evolution, and Taxonomy (biology), taxonomy. It possesses several important biological specimen, manuscript a ...
in 1875, and was elected as a Fellow of the Society on 2 December 1875. He was elected a
Fellow of the Royal Society Fellowship of the Royal Society (FRS, ForMemRS and HonFRS) is an award granted by the Fellows of the Royal Society of London to individuals who have made a "substantial contribution to the improvement of natural science, natural knowledge, incl ...
on 8 June 1882, the same year in which his father died. Darwin edited ''
The Autobiography of Charles Darwin ''The Autobiography of Charles Darwin'' is an autobiography by the English naturalist Charles Darwin. Darwin wrote the text, which he entitled ''Recollections of the Development of my Mind and Character'', for his family. He states that he start ...
'' (1887), and produced some books of letters from the
correspondence of Charles Darwin The British naturalist Charles Darwin corresponded with his extended family and with an extraordinarily wide range of people from all over the world. The letters, over 15,000 in all, provide many insights on issues ranging from the origins of k ...
; ''The Life and Letters of Charles Darwin'' (1887) and ''More Letters of Charles Darwin'' (1905). He also edited
Thomas Huxley Thomas Henry Huxley (4 May 1825 – 29 June 1895) was an English biologist and anthropologist who specialized in comparative anatomy. He has become known as "Darwin's Bulldog" for his advocacy of Charles Darwin's theory of evolution. The stor ...
's ''On the Reception of the Origin of Species'' (1887). Darwin was elected to the
American Philosophical Society The American Philosophical Society (APS) is an American scholarly organization and learned society founded in 1743 in Philadelphia that promotes knowledge in the humanities and natural sciences through research, professional meetings, publicat ...
in 1909. That same year,
Cambridge University The University of Cambridge is a Public university, public collegiate university, collegiate research university in Cambridge, England. Founded in 1209, the University of Cambridge is the List of oldest universities in continuous operation, wo ...
awarded him an honorary doctorate (DSc). He also received honorary doctorates from
Dublin Dublin is the capital and largest city of Republic of Ireland, Ireland. Situated on Dublin Bay at the mouth of the River Liffey, it is in the Provinces of Ireland, province of Leinster, and is bordered on the south by the Dublin Mountains, pa ...
,
Liverpool Liverpool is a port City status in the United Kingdom, city and metropolitan borough in Merseyside, England. It is situated on the eastern side of the River Mersey, Mersey Estuary, near the Irish Sea, north-west of London. With a population ...
,
Sheffield Sheffield is a city in South Yorkshire, England, situated south of Leeds and east of Manchester. The city is the administrative centre of the City of Sheffield. It is historically part of the West Riding of Yorkshire and some of its so ...
,
Brussels Brussels, officially the Brussels-Capital Region, (All text and all but one graphic show the English name as Brussels-Capital Region.) is a Communities, regions and language areas of Belgium#Regions, region of Belgium comprising #Municipalit ...
,
St Andrews St Andrews (; ; , pronounced ʰʲɪʎˈrˠiː.ɪɲ is a town on the east coast of Fife in Scotland, southeast of Dundee and northeast of Edinburgh. St Andrews had a recorded population of 16,800 , making it Fife's fourth-largest settleme ...
, Upsala, and
Prague Prague ( ; ) is the capital and List of cities and towns in the Czech Republic, largest city of the Czech Republic and the historical capital of Bohemia. Prague, located on the Vltava River, has a population of about 1.4 million, while its P ...
. He was knighted in 1913. He is buried in Cambridge. His daughter,
Frances Cornford Frances Crofts Cornford (née Darwin; 30 March 1886 – 19 August 1960) was an English poet. Biography She was the daughter of the botanist Francis Darwin and Newnham College, Cambridge, Newnham College fellow Ellen Wordsworth Darwin, Ellen ...
, was later buried with him.


Family

His first wife, Amy Ruck, died in 1876, a few days after the birth of her son
Bernard Bernard ('' Bernhard'') is a French and West Germanic masculine given name. It has West Germanic origin and is also a surname. The name is attested from at least the 9th century. West Germanic ''Bernhard'' is composed from the two elements ''be ...
, and was buried in Holy Trinity Church,
Corris Corris is a village in the county of Gwynedd, Wales, about north of the town of Machynlleth. The village lies on the west bank of the Afon Dulas (which here forms the boundary with Powys), around that river's confluence with the Afon Deri. ...
, North Wales. According to a letter written by Charles Darwin to his close friend,
Joseph Dalton Hooker Sir Joseph Dalton Hooker (30 June 1817 – 10 December 1911) was a British botanist and explorer in the 19th century. He was a founder of geographical botany and Charles Darwin's closest friend. For 20 years he served as director of the Ro ...
: " I never saw anyone suffer so much as poor Frank. He has gone to N. Wales to bury the body in a little church-yard amongst the mountains". He married his second wife, Ellen Wordsworth Crofts, in 1883. She was a Fellow and lecturer at
Newnham College Newnham College is a women's constituent college of the University of Cambridge. The college was founded in 1871 by a group organising Lectures for Ladies, members of which included philosopher Henry Sidgwick and suffragist campaigner Millicen ...
. She was also a member of the Ladies Dining Society in Cambridge. She died in 1903, and was buried in the churchyard of St. Andrew's Church, Girton. In 1913, he married his third wife, Lady Florence Henrietta Fisher, the widow of
Frederic William Maitland Frederic William Maitland (28 May 1850 – ) was an English historian and jurist who is regarded as the modern father of English legal history. From 1884 until his death in 1906, he was reader in English law, then Downing Professor of the Laws ...
. She died in 1920 and is interred in the Parish of the Ascension Burial Ground, Cambridge, opposite the grave of Sir Francis Darwin and his daughter Frances Cornford.


Publications

*''Life and Letters of Charles Darwin'' (1880); (revised 2nd edition, 1887);2nd printing of revised edition, 1888
*''The Power of Movement in Plants'' (1880) *''The Practical Physiology of Plants'' (1894) *''Elements of Botany'' (1895)
''Rustic Sounds and Other Studies in Literature and Natural History''
(1917)


See also

*
Dorothea Pertz Dorothea Frances Matilda Pertz FLS (14 March 1859 – 6 March 1939) was a British botanist. She co-authored five papers with Francis Darwin, Charles Darwin's son. She was made a Fellow of the Linnean Society, among the first women admitted to ...
with whom he coauthored five papers


References

*


Further reading

* Ayres, Peter. "The Aliveness of Plants: The Darwins at the Dawn of Plant Science" London: PIckering & Chatto, 2008. * Darwin, Francis Sacheverell. (1927). ''Travels in Spain and the East, 1808-1810''. Cambridge University press (reissued by
Cambridge University Press Cambridge University Press was the university press of the University of Cambridge. Granted a letters patent by King Henry VIII in 1534, it was the oldest university press in the world. Cambridge University Press merged with Cambridge Assessme ...
, 2009; )


External links

* * * * {{DEFAULTSORT:Darwin, Francis English biologists English botanists Fellows of the Royal Society Fellows of the Linnean Society of London Alumni of Trinity College, Cambridge Knights Bachelor Charles Darwin biographers Darwin–Wedgwood family People from Downe 1848 births 1925 deaths Burials in Cambridgeshire Presidents of the British Science Association Members of the American Philosophical Society Presidents of the Cambridge Philosophical Society