
''Sinornithomimus'' is a
genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
of
ornithomimid
Ornithomimidae (meaning " bird-mimics") is a family of theropod dinosaurs which bore a superficial resemblance to modern ostriches. Ornithomimids were fast, omnivorous or herbivorous dinosaurs known mainly from the Late Cretaceous Period of ...
that lived in
Asia
Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an ...
during the
Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', ...
period. The first remains were found in 1997, in the
Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', ...
strata
In geology and related fields, a stratum ( : strata) is a layer of rock or sediment characterized by certain lithologic properties or attributes that distinguish it from adjacent layers from which it is separated by visible surfaces known as e ...
of the
Ulansuhai Formation located at Alshanzuo Banner,
Inner Mongolia Autonomous Region
Inner Mongolia, officially the Inner Mongolia Autonomous Region, is an autonomous region of the People's Republic of China. Its border includes most of the length of China's border with the country of Mongolia. Inner Mongolia also accounts for a ...
, Northern
China.
Discovery and naming

The first
fossil
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
remains of ''Sinornithomimus'' were uncovered by
Dong Zhiming
Dong Zhiming ( Chinese: 董枝明, Pinyin: ''Dǒng Zhimíng''; born January 1937) is a Chinese vertebrate paleontologist formerly employed at the Institute of Vertebrate Paleontology and Paleoanthropology (IVPP) in Beijing. He began working at th ...
in the
Ulansuhai Formation as part of the Mongol Highland International Dinosaur Project in 1997. They contained at least fourteen
skeleton
A skeleton is the structural frame that supports the body of an animal. There are several types of skeletons, including the exoskeleton, which is the stable outer shell of an organism, the endoskeleton, which forms the support structure inside ...
s found in close association, nine of which are nearly complete and relatively uncrushed. The find consisted of three sub-adult to adult specimens and eleven juveniles. The unweathered state of the bones, preserved in
siltstone
Siltstone, also known as aleurolite, is a clastic sedimentary rock that is composed mostly of silt. It is a form of mudrock with a low clay mineral content, which can be distinguished from shale by its lack of fissility.Blatt ''et al.'' 1980, ...
interspersed with layers of
clay
Clay is a type of fine-grained natural soil material containing clay minerals (hydrous aluminium phyllosilicates, e.g. kaolin, Al2 Si2 O5( OH)4).
Clays develop plasticity when wet, due to a molecular film of water surrounding the clay part ...
and the absence of evidence for post-mortem movement, argue for a catastrophic event that killed all the individuals present in the find simultaneously and instantaneously.
The
type species
In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen( ...
''Sinornithomimus dongi'' was named and described by
Yoshitsugu Kobayashi and
Lü Junchang in 2003. The generic name means “Chinese bird mimic” while the
specific descriptor honours Dong as the discoverer of the fossils. The
holotype
A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of seve ...
, IVPP-V11797-10, is one of the subadult skeletons. The other skeletons have been assigned as
paratype
In zoology and botany, a paratype is a specimen of an organism that helps define what the scientific name of a species and other taxon actually represents, but it is not the holotype (and in botany is also neither an isotype nor a syntype). O ...
s.
A second expedition in 2001 at the same site led to the discovery of another fossilized herd of thirteen juveniles and subadults of ''Sinornithomimus''. Their positioning suggest that they died together and over a short interval, likely after having become mired in the mud of a drying waterhole. The second discovery also largely consisted of nearly intact exemplars making ''Sinornithomimus'' the most completely known ornithomimid.
Description

''Sinornithomimus'' was a small ornithomimid measuring in length and weighing about with a relatively short neck and head for a member of that group.
[ ]Autapomorphies
In phylogenetics, an autapomorphy is a distinctive feature, known as a derived trait, that is unique to a given taxon. That is, it is found only in one taxon, but not found in any others or outgroup taxa, not even those most closely related to t ...
(unique derived traits) included the possession of a quadrate with a depression having within a smaller opening which is divided by a vertical sheet of bone; and a depression on the lateral side of the posterior process
A process is a series or set of activities that interact to produce a result; it may occur once-only or be recurrent or periodic.
Things called a process include:
Business and management
*Business process, activities that produce a specific se ...
of the parietal bone
The parietal bones () are two bones in the skull which, when joined at a fibrous joint, form the sides and roof of the cranium. In humans, each bone is roughly quadrilateral in form, and has two surfaces, four borders, and four angles. It is named ...
.[
]
Classification
''Sinornithomimus'' was by the describers assigned to the Ornithomimidae. It was a basal
Basal or basilar is a term meaning ''base'', ''bottom'', or ''minimum''.
Science
* Basal (anatomy), an anatomical term of location for features associated with the base of an organism or structure
* Basal (medicine), a minimal level that is nec ...
ornithomimid that was considered by its describers a more derived form than ''Archaeornithomimus
''Archaeornithomimus'' (meaning "ancient bird mimic") is a genus of ornithomimosaurian theropod dinosaur that lived in Asia during the Late Cretaceous period, around 96 million years ago in the Iren Dabasu Formation.
Discovery and naming
In 1 ...
'',[ though more recent analyses reverse the situation. The structure of the hand is similar to that of ''Archaeornithomimus'' representing thus an intermediate between the "primitive" condition of the ornithomimosaur '']Harpymimus
''Harpymimus'' is a basal ornithomimosaurian theropod dinosaur from the Early Cretaceous Period of what is now Mongolia. Unlike later, more derived ornithomimosaurs, ''Harpymimus'' still possessed teeth, although they appear to have been re ...
'' and the one of the more derived ornithomimids. ''Sinornithomimus'' renders some synapomorphies
In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to hav ...
of ornithomimids plesiomorphic
In phylogenetics, a plesiomorphy ("near form") and symplesiomorphy are synonyms for an ancestral character shared by all members of a clade, which does not distinguish the clade from other clades.
Plesiomorphy, symplesiomorphy, apomorphy, an ...
, while also differentiating Asian ornithomimid rhamphothecae from North American ones, based on maxilla
The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The ...
ry vascular
Blood vessels are the structures of the circulatory system that transport blood throughout the human body. These vessels transport blood cells, nutrients, and oxygen to the tissues of the body. They also take waste and carbon dioxide away from ...
found in the latter.[
The following cladogram follows that of Xu and colleagues in 2011:]
Paleobiology
The herbivory
A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthp ...
of this genus is supported by the rare presence of gastrolith
A gastrolith, also called a stomach stone or gizzard stone, is a rock held inside a gastrointestinal tract. Gastroliths in some species are retained in the muscular gizzard and used to grind food in animals lacking suitable grinding teeth. In oth ...
s, as such stones were found forming conspicuous masses in the stomach areas of the fossil skeletons.[Kobayashi, Y., Lü, J.−C., Dong, Z.−M., Barsbold, R., Azuma, Y., and Tomida, Y. (1999) Herbivorous diet in an ornithomimid dinosaur. ''Nature 402:'' 480–481.] This species was gregarious, which is corroborated by the arrangement of the fossil remains in a small bonebed with the juvenile individuals being approximately of the same age. This is furthermore supported by the increase of running ability as the animal progressed in its ontogeny
Ontogeny (also ontogenesis) is the origination and development of an organism (both physical and psychological, e.g., moral development), usually from the time of fertilization of the egg to adult. The term can also be used to refer to the stu ...
, thus shown in the more cursorial
A cursorial organism is one that is adapted specifically to run. An animal can be considered cursorial if it has the ability to run fast (e.g. cheetah) or if it can keep a constant speed for a long distance (high endurance). "Cursorial" is often u ...
proportions of the adults which possess relatively longer lower legs. It was assumed by the describers that adults protected themselves and their juveniles from predators by forming familial groups; in 2008 however a study concluded that the all juvenile herds suggest that immature individuals were left to fend for themselves in juvenile groups while adults preoccupied themselves with nesting or brooding.[
]
See also
* Timeline of ornithomimosaur research
References
{{Portalbar, Dinosaurs, Cretaceous
Ornithomimids
Late Cretaceous dinosaurs of Asia
Fossil taxa described in 2003
Taxa named by Lü Junchang