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''Sinornithomimus'' is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of ornithomimid that lived in
Asia Asia ( , ) is the largest continent in the world by both land area and population. It covers an area of more than 44 million square kilometres, about 30% of Earth's total land area and 8% of Earth's total surface area. The continent, which ...
during the
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
period. The first remains were found in 1997, in the
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
strata In geology and related fields, a stratum (: strata) is a layer of Rock (geology), rock or sediment characterized by certain Lithology, lithologic properties or attributes that distinguish it from adjacent layers from which it is separated by v ...
of the
Ulansuhai Formation The Ulansuhai Formation () is a geological Formation (geology), formation in Inner Mongolia, north China. Dinosaur remains are among the fossils that have been recovered from the formation.Weishampel, David B; et al. (2004). "Dinosaur distribution ...
located at Alshanzuo Banner, Inner Mongolia Autonomous Region, Northern
China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
.


Discovery and naming

The first
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
remains of ''Sinornithomimus'' were uncovered by
Dong Zhiming Dong Zhiming (Chinese language, Chinese: 董枝明, Pinyin: ''Dǒng Zhimíng''; January 1937 – 20 October 2024) was a Chinese vertebrate paleontologist formerly employed at the Institute of Vertebrate Paleontology and Paleoanthropology (IVPP) ...
in the
Ulansuhai Formation The Ulansuhai Formation () is a geological Formation (geology), formation in Inner Mongolia, north China. Dinosaur remains are among the fossils that have been recovered from the formation.Weishampel, David B; et al. (2004). "Dinosaur distribution ...
as part of the Mongol Highland International Dinosaur Project in 1997. They contained at least fourteen
skeleton A skeleton is the structural frame that supports the body of most animals. There are several types of skeletons, including the exoskeleton, which is a rigid outer shell that holds up an organism's shape; the endoskeleton, a rigid internal fra ...
s found in close association, nine of which are nearly complete and relatively uncrushed. The find consisted of three sub-adult to adult specimens and eleven juveniles. The unweathered state of the bones, preserved in
siltstone Siltstone, also known as aleurolite, is a clastic sedimentary rock that is composed mostly of silt. It is a form of mudrock with a low clay mineral content, which can be distinguished from shale by its lack of fissility. Although its permeabil ...
interspersed with layers of
clay Clay is a type of fine-grained natural soil material containing clay minerals (hydrous aluminium phyllosilicates, e.g. kaolinite, ). Most pure clay minerals are white or light-coloured, but natural clays show a variety of colours from impuriti ...
and the absence of evidence for post-mortem movement, argue for a catastrophic event that killed all the individuals present in the find simultaneously and instantaneously. The
type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
''Sinornithomimus dongi'' was named and described by Yoshitsugu Kobayashi and
Lü Junchang Lü Junchang (; 1965 – 9 October 2018) was a Chinese palaeontologist and professor at the Institute of Geology, Chinese Academy of Geological Sciences. An expert on Mesozoic reptiles, he described and named dozens of dinosaur and pterosaur taxa ...
in 2003. The generic name means “Chinese bird mimic” while the specific descriptor honours Dong as the discoverer of the fossils. The
holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
, IVPP-V11797-10, is one of the subadult skeletons. The other skeletons have been assigned as
paratype In zoology and botany, a paratype is a specimen of an organism that helps define what the scientific name of a species and other taxon actually represents, but it is not the holotype (and in botany is also neither an isotype (biology), isotype ...
s. A second expedition in 2001 at the same site led to the discovery of another fossilized herd of thirteen juveniles and subadults of ''Sinornithomimus''. Their positioning suggest that they died together and over a short interval, likely after having become mired in the mud of a drying waterhole. The second discovery also largely consisted of nearly intact exemplars making ''Sinornithomimus'' the most completely known ornithomimid.


Description

''Sinornithomimus'' was a small ornithomimid measuring in length and weighing about with a relatively short neck and head for a member of that group. Autapomorphies (unique derived traits) included the possession of a quadrate with a depression having within a smaller
opening Opening may refer to: Types of openings * Hole * A title sequence or opening credits * Grand opening of a business or other institution * Inauguration * Keynote * Opening sentence * Opening sequence * Opening statement, a beginning statemen ...
which is divided by a vertical sheet of bone; and a depression on the lateral side of the posterior
process A process is a series or set of activities that interact to produce a result; it may occur once-only or be recurrent or periodic. Things called a process include: Business and management * Business process, activities that produce a specific s ...
of the
parietal bone The parietal bones ( ) are two bones in the skull which, when joined at a fibrous joint known as a cranial suture, form the sides and roof of the neurocranium. In humans, each bone is roughly quadrilateral in form, and has two surfaces, four bord ...
.


Classification

''Sinornithomimus'' was by the describers assigned to the Ornithomimidae. It was a basal ornithomimid that was considered by its describers a more derived form than '' Archaeornithomimus'', though more recent analyses reverse the situation. The structure of the hand is similar to that of ''Archaeornithomimus'' representing thus an intermediate between the "primitive" condition of the ornithomimosaur ''
Harpymimus ''Harpymimus'' is a basal ornithomimosaurian theropod dinosaur from the Early Cretaceous Period (geology), Period of what is now Mongolia. Unlike later, more derived ornithomimosaurs, ''Harpymimus'' still possessed teeth, although they appear to ...
'' and that of the more derived ornithomimids. ''Sinornithomimus'' renders some
synapomorphies In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to ...
of ornithomimids
plesiomorphic In phylogenetics, a plesiomorphy ("near form") and symplesiomorphy are synonyms for an ancestral character shared by all members of a clade, which does not distinguish the clade from other clades. Plesiomorphy, symplesiomorphy, apomorphy, an ...
, while also differentiating Asian ornithomimid rhamphothecae from North American ones, based on
maxilla In vertebrates, the maxilla (: maxillae ) is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The two maxil ...
ry
vascular Vascular can refer to: * blood vessels, the vascular system in animals * vascular tissue Vascular tissue is a complex transporting tissue, formed of more than one cell type, found in vascular plants. The primary components of vascular tissue ...
foramina In anatomy and osteology, a foramen (; : foramina, or foramens ; ) is an opening or enclosed gap within the dense connective tissue (bones and deep fasciae) of extant and extinct amniote animals, typically to allow passage of nerves, arter ...
found in the latter. The following cladogram follows that of Xu and colleagues in 2011:


Paleobiology

The
herbivory A herbivore is an animal anatomically and physiologically evolved to feed on plants, especially upon vascular tissues such as foliage, fruits or seeds, as the main component of its diet. These more broadly also encompass animals that eat n ...
of this genus is supported by the rare presence of
gastrolith A gastrolith, also called a stomach stone or gizzard stone, is a rock held inside a gastrointestinal tract. Gastroliths in some species are retained in the muscular gizzard and used to grind food in animals lacking suitable grinding teeth. In ...
s, as such stones were found forming conspicuous masses in the stomach areas of the fossil skeletons.Kobayashi, Y., Lü, J.−C., Dong, Z.−M., Barsbold, R., Azuma, Y., and Tomida, Y. (1999) Herbivorous diet in an ornithomimid dinosaur. ''Nature 402:'' 480–481. This species was gregarious, which is corroborated by the arrangement of the fossil remains in a small bonebed with the juvenile individuals being approximately of the same age. This is furthermore supported by the increase of running ability as the animal progressed in its
ontogeny Ontogeny (also ontogenesis) is the origination and development of an organism (both physical and psychological, e.g., moral development), usually from the time of fertilization of the ovum, egg to adult. The term can also be used to refer to t ...
, thus shown in the more
cursorial A cursorial organism is one that is adapted specifically to run. An animal can be considered cursorial if it has the ability to run fast (e.g. cheetah) or if it can keep a constant speed for a long distance (high endurance). "Cursorial" is often ...
proportions of the adults which possess relatively longer lower legs. It was assumed by the describers that adults protected themselves and their juveniles from predators by forming familial groups; in 2008 however a study concluded that the all juvenile herds suggest that immature individuals were left to fend for themselves in juvenile groups while adults preoccupied themselves with nesting or brooding.


See also

*
Timeline of ornithomimosaur research This timeline of ornithomimosaur research is a chronological listing of events in the History of paleontology, history of paleontology focused on the ornithomimosaurs, a group of bird-like theropods popularly known as the ostrich dinosaurs. Altho ...


References

{{Portalbar, Dinosaurs Ornithomimidae Dinosaur genera Turonian dinosaurs Taxa named by Lü Junchang Fossil taxa described in 2003 Dinosaurs of China