''Tilia tomentosa'', known as silver linden in the US and silver lime in the UK, is a

species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...

flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Ancient Greek, Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed with ...

in the

family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...

Malvaceae,

native Native may refer to: People * '' Jus sanguinis'', nationality by blood * '' Jus soli'', nationality by location of birth * Indigenous peoples, peoples with a set of specific rights based on their historical ties to a particular territory ** Nat ...

to southeastern Europe and southwestern Asia, from Romania and the Balkans east to western Turkey, occurring at moderate altitudes.Rushforth, K. (1999). ''Trees of Britain and Europe''. Collins .Flora Europaea
''Tilia tomentosa''
/ref>

Description

''Tilia tomentosa'' is a

deciduous In the fields of horticulture and botany, the term deciduous () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed Leaf, leaves, usually in the autumn; to the shedding of petals, aft ...

tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, e.g., including only woody plants with secondary growth, only ...

growing to tall, with a trunk up to in diameter. The

leaves A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, ...

are alternately arranged, rounded to triangular-ovate, 4–13 cm long and broad with a 2.5–4 cm petiole, green and mostly hairless above, densely white tomentose with white hairs below, and with a coarsely toothed margin. The

flower Flowers, also known as blooms and blossoms, are the reproductive structures of flowering plants ( angiosperms). Typically, they are structured in four circular levels, called whorls, around the end of a stalk. These whorls include: calyx, m ...

s are pale yellow,

hermaphrodite A hermaphrodite () is a sexually reproducing organism that produces both male and female gametes. Animal species in which individuals are either male or female are gonochoric, which is the opposite of hermaphroditic. The individuals of many ...

, produced in

cymes In botany, an inflorescence is a group or cluster of flowers arranged on a plant's stem that is composed of a main branch or a system of branches. An inflorescence is categorized on the basis of the arrangement of flowers on a main axis ( ped ...

of three to ten in mid to late summer with a pale green subtending leafy

bract In botany, a bract is a modified or specialized leaf, associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves in size, color, shape or texture. They also lo ...

; they have a strong scent and are pollinated by

honeybee A honey bee (also spelled honeybee) is a eusocial flying insect within the genus ''Apis'' of the bee clade, all native to mainland Afro-Eurasia. After bees spread naturally throughout Africa and Eurasia, humans became responsible for the c ...

s. The

fruit In botany, a fruit is the seed-bearing structure in flowering plants (angiosperms) that is formed from the ovary after flowering. Fruits are the means by which angiosperms disseminate their seeds. Edible fruits in particular have long propaga ...

is a dry nut-like

drupe In botany, a drupe (or stone fruit) is a type of fruit in which an outer fleshy part (exocarp, or skin, and mesocarp, or flesh) surrounds a single shell (the ''pip'' (UK), ''pit'' (US), ''stone'', or ''pyrena'') of hardened endocarp with a seed ...

8–10 mm long, downy, and slightly ribbed.Mitchell, A. F. (1974). ''A Field Guide to the Trees of Britain and Northern Europe''. Collins

Cultivation and uses

It is widely grown as an

ornamental tree Ornamental plants or ''garden plants'' are plants that are primarily grown for their beauty but also for qualities such as scent or how they shape physical space. Many flowering plants and garden varieties tend to be specially bred cultivars th ...

throughout Europe. The

cultivar A cultivar is a kind of Horticulture, cultivated plant that people have selected for desired phenotypic trait, traits and which retains those traits when Plant propagation, propagated. Methods used to propagate cultivars include division, root a ...

'Brabant' has a strong central stem and a symmetrical conic crown. The cultivar 'Petiolaris' (pendent or weeping silver lime) differs in longer leaf petioles 4–8 cm long and drooping leaves; it is of unknown origin and usually sterile, and may be a

hybrid Hybrid may refer to: Science * Hybrid (biology), an offspring resulting from cross-breeding ** Hybrid grape, grape varieties produced by cross-breeding two ''Vitis'' species ** Hybridity, the property of a hybrid plant which is a union of two diff ...

with another ''Tilia'' species. It is very tolerant of urban pollution, soil compaction, heat, and drought, and would be a good street tree in urban areas.Mitchell, A. F. (1996). ''Alan Mitchell's Trees of Britain''. HarperCollins. . In cultivation in the UK, ''T. tomentosa'' 'Petiolaris' has gained the

Royal Horticultural Society The Royal Horticultural Society (RHS), founded in 1804 as the Horticultural Society of London, is the UK's leading gardening charity. The RHS promotes horticulture through its five gardens at Wisley (Surrey), Hyde Hall (Essex), Harlow Carr ...

Award of Garden Merit The Award of Garden Merit (AGM) is a long-established award for plants by the British Royal Horticultural Society (RHS). It is based on assessment of the plants' performance under UK growing conditions. It includes the full range of cultivated p ...

. An

infusion Infusion is the process of extracting chemical compounds or flavors from plant material in a solvent such as water, oil or alcohol, by allowing the material to remain suspended in the solvent over time (a process often called steeping). An inf ...

made from the flowers of ''T. tomentosa'' is

antispasmodic An antispasmodic (synonym: spasmolytic) is a pharmaceutical drug or other agent that suppresses muscle spasms. Smooth muscle spasm One type of antispasmodics is used for smooth muscle relaxation, especially in tubular organs of the gastrointestina ...

diaphoretic Perspiration, also known as sweat, is the fluid secreted by sweat glands in the skin of mammals. Two types of sweat glands can be found in humans: eccrine glands and Apocrine sweat gland, apocrine glands. The eccrine sweat glands are distribu ...

and

sedative A sedative or tranquilliser is a substance that induces sedation by reducing irritability or Psychomotor agitation, excitement. They are central nervous system (CNS) Depressant, depressants and interact with brain activity, causing its decelera ...

.Plants For A Future
''Tilia tomentosa''
which cites Lauriault, J. (1989). ''Identification Guide to the Trees of Canada''. Fitzhenry and Whiteside, Ontario. This may be attributable to the presence of pharmacologically active ligands of

benzodiazepine Benzodiazepines (BZD, BDZ, BZs), colloquially known as "benzos", are a class of central nervous system (CNS) depressant, depressant drugs whose core chemical structure is the fusion of a benzene ring and a diazepine ring. They are prescribed t ...

receptor Receptor may refer to: * Sensory receptor, in physiology, any neurite structure that, on receiving environmental stimuli, produces an informative nerve impulse *Receptor (biochemistry), in biochemistry, a protein molecule that receives and respond ...

. A widespread belief is that the nectar of this species contains

mannose Mannose is a sugar with the formula , which sometimes is abbreviated Man. It is one of the monomers of the aldohexose series of carbohydrates. It is a C-2 epimer of glucose. Mannose is important in human metabolism, especially in the glycosylatio ...

, which can be toxic to some

bee Bees are winged insects closely related to wasps and ants, known for their roles in pollination and, in the case of the best-known bee species, the western honey bee, for producing honey. Bees are a monophyletic lineage within the superfamil ...

s. This is incorrect; the sight of numerous comatose bees found on the ground at flowering time is rather a result of the paucity of nectar sources in late summer in urban areas. The evidence against a toxin in the nectar being responsible for mass bee deaths under Tilia trees is supported further by Koch and Stevenson (2017) who also suggest that the presence of caffeine in linden nectar may mean that linden trees can chemically deceive foraging bees to make sub-optimal foraging decisions, in some cases leading to their starvation. This species, while fragrant in spring, drops buds and pollen during the spring and fall.

Notable trees

Eminescu's Linden Tree Eminescu's Linden Tree () is a 500-year-old silver lime (''Tilia tomentosa Moench'') in Copou Park, Iași, Romania. Mihai Eminescu reportedly wrote some of his best works underneath this lime, rendering the tree one of Romania's most important ...

() is a 500-year-old silver lime situated in the Copou Public Garden,

Iași Iași ( , , ; also known by other #Etymology and names, alternative names), also referred to mostly historically as Jassy ( , ), is the Cities in Romania, third largest city in Romania and the seat of Iași County. Located in the historical ...

Romania Romania is a country located at the crossroads of Central Europe, Central, Eastern Europe, Eastern and Southeast Europe. It borders Ukraine to the north and east, Hungary to the west, Serbia to the southwest, Bulgaria to the south, Moldova to ...

Mihai Eminescu Mihai Eminescu (; born Mihail Eminovici; 15 January 1850 – 15 June 1889) was a Romanians, Romanian Romanticism, Romantic poet, novelist, and journalist from Moldavia, generally regarded as the most famous and influential Romanian poet. Emin ...

reportedly wrote some of his best works underneath this silver lime, rendering the tree one of Romania's most important natural monuments and an Iași landmark.Pettersen, L. & Baker, M. . Romania. Lonely Planet Travel Guide. p. 262.

References

External links

''Tilia tomentosa''
- information, genetic conservation units and related resources.

European Forest Genetic Resources Programme European Forest Genetic Resources Programme (EUFORGEN) is an international network that promotes the conservation and sustainable use of forest genetic resources in Europe as an integral part of sustainable forest management. It was established ...

(EUFORGEN) {{Authority control Flora of Europe tomentosa Taxa named by Conrad Moench