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''Silene dioica'' ( syn. ''Melandrium rubrum''), known as red campion and red catchfly, is a
herbaceous Herbaceous plants are vascular plants that have no persistent woody stems above ground. This broad category of plants includes many perennials, and nearly all annuals and biennials. Definitions of "herb" and "herbaceous" The fourth edition of ...
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Ancient Greek, Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed with ...
in the family
Caryophyllaceae Caryophyllaceae, commonly called the pink family or carnation family, is a family (biology), family of flowering plants. It is included in the dicotyledon order Caryophyllales in the APG III system, alongside 33 other families, including Amaranth ...
, native to Europe and introduced to the Americas.


Description

It is a biennial or
perennial plant In horticulture, the term perennial (''wikt:per-#Prefix, per-'' + ''wikt:-ennial#Suffix, -ennial'', "through the year") is used to differentiate a plant from shorter-lived annual plant, annuals and biennial plant, biennials. It has thus been d ...
, The plant grows to tall, with branching stems. The deep green
leaves A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, ...
are in
opposite In lexical semantics, opposites are words lying in an inherently incompatible binary relationship. For example, something that is ''even'' entails that it is not ''odd''. It is referred to as a 'binary' relationship because there are two members i ...
and
decussate Decussation is used in biological contexts to describe a crossing (due to the shape of the Roman numeral for ten, an uppercase 'X' (), ). In Latin anatomical terms, the form is used, e.g. . Similarly, the anatomical term chiasma is named aft ...
pairs, simple acute ovate, long with an untoothed margin; both the leaves and stems of the plant are hairy and slightly sticky. The upper leaves are stalkless. Blooming from May to October, the unscented
flower Flowers, also known as blooms and blossoms, are the reproductive structures of flowering plants ( angiosperms). Typically, they are structured in four circular levels, called whorls, around the end of a stalk. These whorls include: calyx, m ...
s are dark pink to red, each across. There are five
petals Petals are modified leaves that form an inner whorl surrounding the reproductive parts of flowers. They are often brightly coloured or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''coroll ...
which are deeply notched at the end, narrowed at the base and all go into an urn-shaped calyx. As indicated by the specific name, male and female flowers are borne on separate plants (
dioecious Dioecy ( ; ; adj. dioecious, ) is a characteristic of certain species that have distinct unisexual individuals, each producing either male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproduction is ...
), the male with 10
stamens The stamen (: stamina or stamens) is a part consisting of the male reproductive organs of a flower. Collectively, the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filamen ...
and a 10-veined calyx, the female with 5 styles and a 20-veined calyx. The fruit, produced from July onwards, is an ovoid capsule containing numerous seeds, opening at the apex by 10 teeth which curve back.


Taxonomy

Plants of ''Silene latifolia'' × ''Silene dioica'' = ''Silene'' × ''hampeana'' that are fertile hybrids with the closely related white campion (''
Silene latifolia ''Silene latifolia'', commonly known as white campion, is a dioecious flowering plant in the family Caryophyllaceae, native to most of Europe, Western Asia and northern Africa. It is a herbaceous annual, occasionally biennial or a short-lived ...
'') are common in some areas. They may have paler pink flowers and be intermediate between the two species in other characters.


Distribution and habitat

''Silene dioica'' is native to northern and central Europe and is locally abundant throughout the
British Isles The British Isles are an archipelago in the Atlantic Ocean, North Atlantic Ocean off the north-western coast of continental Europe, consisting of the islands of Great Britain, Ireland, the Isle of Man, the Inner Hebrides, Inner and Outer Hebr ...
. It is generally common in
Northern Ireland Northern Ireland ( ; ) is a Countries of the United Kingdom, part of the United Kingdom in the north-east of the island of Ireland. It has been #Descriptions, variously described as a country, province or region. Northern Ireland shares Repub ...
, but rare elsewhere in
Ireland Ireland (, ; ; Ulster Scots dialect, Ulster-Scots: ) is an island in the North Atlantic Ocean, in Northwestern Europe. Geopolitically, the island is divided between the Republic of Ireland (officially Names of the Irish state, named Irelan ...
. It is common on the
Isle of Man The Isle of Man ( , also ), or Mann ( ), is a self-governing British Crown Dependency in the Irish Sea, between Great Britain and Ireland. As head of state, Charles III holds the title Lord of Mann and is represented by a Lieutenant Govern ...
. It has been introduced in Iceland, Canada, the US, and Argentina. Red campion grows in roadsides,
woodland A woodland () is, in the broad sense, land covered with woody plants (trees and shrubs), or in a narrow sense, synonymous with wood (or in the U.S., the '' plurale tantum'' woods), a low-density forest forming open habitats with plenty of sunli ...
s, and rocky slopes. It prefers to grow on damp, non-acid soils.


Ecology

The flowers are frequently visited by flies such as ''
Rhingia campestris ''Rhingia campestris'' is a species of hoverfly, long, with a wingspan of . It is common across the Palearctic from March until November. It has a broad orange abdomen with a black line along the sides (the black line is absent along the sides o ...
''. The
nectar Nectar is a viscous, sugar-rich liquid produced by Plant, plants in glands called nectaries, either within the flowers with which it attracts pollination, pollinating animals, or by extrafloral nectaries, which provide a nutrient source to an ...
of the flowers is utilised by
bumblebees A bumblebee (or bumble bee, bumble-bee, or humble-bee) is any of over 250 species in the genus ''Bombus'', part of Apidae, one of the bee families. This genus is the only Extant taxon, extant group in the tribe Bombini, though a few extinct r ...
and
butterflies Butterflies are winged insects from the lepidopteran superfamily Papilionoidea, characterized by large, often brightly coloured wings that often fold together when at rest, and a conspicuous, fluttering flight. The oldest butterfly fossi ...
, and several species of
moth Moths are a group of insects that includes all members of the order Lepidoptera that are not Butterfly, butterflies. They were previously classified as suborder Heterocera, but the group is Paraphyly, paraphyletic with respect to butterflies (s ...
feed on the
foliage A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, f ...
. The flowers of red-campion along with a number of other Caryophyllaceae members, are very susceptible to a
smut (fungus) The smuts are multicellular fungi characterized by their large numbers of teliospores. The smuts get their name from a Germanic word for 'dirt' because of their dark, thick-walled, and dust-like teliospores. They are mostly Ustilaginomycetes (ph ...
infection. In this case by ''
Microbotryum silenes-dioicae ''Microbotryum silenes-dioicae'' is a species of fungus first isolated from Brittany, France. Its name refers to its host species, ''Silene dioica''. The fungus is the cause of anther-smut disease, which results in fungal spores replacing the pol ...
'' known as anther-smut which appears as a mass of brown spores in the mouth of the flower where the anthers would normally be.


Cultivation

This plant is used as an ornamental perennial flower for the perennial border. One particularly notable variety is a hot pink, double flowered variety with deep green leaves called 'Firefly'.


In culture

On the Isle of Man, it is known as "blaa ny ferrishyn" or "fairy flower", and there is a local taboo against picking it.


References

dioica Dioecious plants {{Caryophyllaceae-stub