The shadow darner (''Aeshna umbrosa'') is a species of

dragonfly A dragonfly is a flying insect belonging to the infraorder Anisoptera below the order Odonata. About 3,000 extant species of dragonflies are known. Most are tropical, with fewer species in temperate regions. Loss of wetland habitat threat ...

in the

family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...

Aeshnidae Aeshnidae, also called aeshnids, hawkers, or darners, is a family of dragonflies, found nearly worldwide, with more than 50 genera and over 450 species. The family includes some of the largest dragonflies. Description Common worldwide or nearl ...

. It is found in almost all of Canada and most states in the

United States The United States of America (USA), also known as the United States (U.S.) or America, is a country primarily located in North America. It is a federal republic of 50 U.S. state, states and a federal capital district, Washington, D.C. The 48 ...

Identification

The shadow darner is a large dragonfly with a length of . The base is brownish black in color. Greenish crescent-shaped spots are found at the top of the thorax. The sides of the thorax are marked with two yellowish to yellowish-green diagonal stripes. Its abdomen is marked with bluish green spots. The male shadow darner has paddle-shaped anal appendages. The

naiad In Greek mythology, the naiads (; ), sometimes also hydriads, are a type of female spirit, or nymph, presiding over fountains, wells, springs, streams, brooks and other bodies of fresh water. They are distinct from river gods, who embodied ...

of the shadow darner is large in size, with a length of . This naiad is long and slender, which is the typical shape of immature darners. It is mottled green and brown. The shadow darner has a vertically flattened cerci with a spike at the end, which is much brighter than the lance-tipped darner.

Distribution

This dragonfly is found in most of the United States except the dry Southwest and in all of the provinces and territories of Canada.

Habitat and diet

The shadow darner patrols along small marshy streams. It is often found feeding along woodland edges or even in deep shadow in full forest. Shadow darners can also be found near ditches, slow streams, and ponds. This darner has a long flight season of late April to November. The shadow darner naiad feeds on a wide variety of aquatic insects, such as

mosquito Mosquitoes, the Culicidae, are a Family (biology), family of small Diptera, flies consisting of 3,600 species. The word ''mosquito'' (formed by ''Musca (fly), mosca'' and diminutive ''-ito'') is Spanish and Portuguese for ''little fly''. Mos ...

larvae A larva (; : larvae ) is a distinct juvenile form many animals undergo before metamorphosis into their next life stage. Animals with indirect developmental biology, development such as insects, some arachnids, amphibians, or cnidarians typical ...

, other aquatic

fly Flies are insects of the order Diptera, the name being derived from the Greek δι- ''di-'' "two", and πτερόν ''pteron'' "wing". Insects of this order use only a single pair of wings to fly, the hindwings having evolved into advanced ...

larvae,

mayfly Mayflies (also known as shadflies or fishflies in Canada and the upper Midwestern United States, as Canadian soldiers in the American Great Lakes region, and as up-winged flies in the United Kingdom) are aquatic insects belonging to the orde ...

larvae, and freshwater

shrimp A shrimp (: shrimp (American English, US) or shrimps (British English, UK)) is a crustacean with an elongated body and a primarily Aquatic locomotion, swimming mode of locomotion – typically Decapods belonging to the Caridea or Dendrobranchi ...

. They also feed on small fish and

tadpoles A tadpole or polliwog (also spelled pollywog) is the larval stage in the biological life cycle of an amphibian. Most tadpoles are fully aquatic, though some species of amphibians have tadpoles that are terrestrial. Tadpoles have some fish-li ...

. This adult will eat almost any soft-bodied flying insect, including mosquitoes, flies,

butterflies Butterflies are winged insects from the lepidopteran superfamily Papilionoidea, characterized by large, often brightly coloured wings that often fold together when at rest, and a conspicuous, fluttering flight. The oldest butterfly fossi ...

moths Moths are a group of insects that includes all members of the order Lepidoptera that are not butterflies. They were previously classified as suborder Heterocera, but the group is paraphyletic with respect to butterflies (suborder Rhopalocera) a ...

mayflies Mayflies (also known as shadflies or fishflies in Canada and the upper Midwestern United States, as Canadian soldiers in the American Great Lakes region, and as up-winged flies in the United Kingdom) are aquatic insects belonging to the order ...

, and

stoneflies Plecoptera is an order of insects commonly known as stoneflies. Some 3,500 species are described worldwide, with new species still being discovered. Stoneflies are found worldwide, except Antarctica. Stoneflies are believed to be one of the most ...

Ecology

The naiad is an active predator and are able to swim by jet propulsion. They squirt water out from the ends of their abdomens. They will generally take several years to mature, and when the immature changes into a dragonfly, it does so at night. This behavior probably was evolved to avoid being eaten be daytime predators. The adult generally flies from late April to November, and does all of its hunting while on the wing. The shadow darner is able to regulate its body temperature which enables it to fly in temperatures too cold for most dragonflies. This dragonfly species in particular seems to be extremely cold tolerant. This dragonfly flies at dusk and in shaded areas, and it flies later into the fall than any species other than the yellow-legged meadowhawk ('' Sympetrum vicinum'').

Reproduction

Male shadow darners establish and defend territories along the shores of slow streams and ponds. After mating, females fly singly, without the male attached, to lay their eggs in the stems and leaves of aquatic plants.

Subspecies

*''Aeshna umbrosa occidentalis'' *''Aeshna umbrosa umbrosa''

References

Ordonates FrameShadow Darner - Aeshna umbrosa''Aeshna umbrosa'' (Shadow Darner)Catalogue of Life : 2009 Annual Checklist : Search
* Kurt Mead. (2009) Dragonflies of the North Woods. Second edition. Duluth, MN:Kollath+Stensaas, p. 38-39. * Dunkle, Sidney W. (2000) Dragonflies through Binoculars. New York: Oxford University Press. pp. 176. . {{Taxonbar, from=Q1836401 Aeshnidae Odonata of North America Insects described in 1908