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The Dixie chub (''Semotilus thoreauianus'') is a species of freshwater
ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fishes, is a class of bony fish. They comprise over 50% of living vertebrate species. The ray-finned fishes are so called because their fins are webs of skin supported by bony or h ...
from the
carp Carp are various species of oily freshwater fish from the family Cyprinidae, a very large group of fish native to Europe and Asia. While carp is consumed in many parts of the world, they are generally considered an invasive species in parts of ...
and
minnow Minnow is the common name for a number of species of small freshwater fish, belonging to several genera of the families Cyprinidae and Leuciscidae. They are also known in Ireland as pinkeens. Smaller fish in the subfamily Leusciscidae are ...
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Cyprinidae Cyprinidae is a family of freshwater fish commonly called the carp or minnow family. It includes the carps, the true minnows, and relatives like the barbs and barbels. Cyprinidae is the largest and most diverse fish family and the largest v ...
. It is
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found els ...
to southeastern United States.


Description

The Dixie chub is similar in shape to the creek chub, (''
Semotilus atromaculatus ''Semotilus atromaculatus'', known as the creek chub or the common creek chub, is a small minnow, a freshwater fish found in the eastern US and Canada. Differing in size and color depending on origin of development, the creek chub can usually be ...
'') although it is typically a more robust and shorter than the creek chub. It also has larger and fewer lateral line scales, the origin of the
dorsal fin A dorsal fin is a fin located on the back of most marine and freshwater vertebrates within various taxa of the animal kingdom. Many species of animals possessing dorsal fins are not particularly closely related to each other, though through c ...
is to the rear of that of the
pelvic fin Pelvic fins or ventral fins are paired fins located on the ventral surface of fish. The paired pelvic fins are homologous to the hindlimbs of tetrapods. Structure and function Structure In actinopterygians, the pelvic fin consists of two e ...
whereas in ''S. atromaculatus'' it sits directly over the pelvic fin origin, and the spots on the dorsal fin and
caudal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as s ...
are less distinct. In the breeding season the males develop four large, hooked tubercles on either side of the head, with those nearest the nares frequently becoming fused. They are generally dark on the back, with the underside being colored from orange to pink on the underside while its fins are yellow-orange. Compared to ''S. atromaculatus'' the Dixie chub has a smaller number of head tubercles on its gill covers and caudal fin. The spot on the caudal peduncle is more diffuse than in ''S. atromaculatus'' which is normally wedge-shaped and distinct from the lateral stripe, while the equivalent spot in ''S. atromaculatus'' is quadrilateral and is joined to the lateral stripe. The lateral strip is dark and rather wide but not very distinct. It grows to a length of .


Distribution

The Dixie chub is endemic to rivers that flow into the
Gulf of Mexico The Gulf of Mexico ( es, Golfo de México) is an ocean basin and a marginal sea of the Atlantic Ocean, largely surrounded by the North American continent. It is bounded on the northeast, north and northwest by the Gulf Coast of the United S ...
in the south-eastern United States from the
Tombigbee River The Tombigbee River is a tributary of the Mobile River, approximately 200 mi (325 km) long, in the U.S. states of Mississippi and Alabama. Together with the Alabama, it merges to form the short Mobile River before the latter empties int ...
in
Alabama (We dare defend our rights) , anthem = " Alabama" , image_map = Alabama in United States.svg , seat = Montgomery , LargestCity = Huntsville , LargestCounty = Baldwin County , LargestMetro = Greater Birmingham , area_total_km2 = 135,7 ...
to the
Ochlockonee River The Ochlockonee River ( ) is a fast running river, except where it has been dammed to form Lake Talquin in Florida, originating in Georgia and flowing for before terminating in Florida. Background The name is from the Hitchiti language word ...
drainage in
Georgia Georgia most commonly refers to: * Georgia (country), a country in the Caucasus region of Eurasia * Georgia (U.S. state), a state in the Southeast United States Georgia may also refer to: Places Historical states and entities * Related to t ...
and
Florida Florida is a state located in the Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia, to the east by the Bahamas and Atlantic Ocean, a ...
. It has also been recorded from three tributaries of Bear Creek in Colbert County, Alabama, the first records from the drainage of the
Tennessee River The Tennessee River is the largest tributary of the Ohio River. It is approximately long and is located in the southeastern United States in the Tennessee Valley. The river was once popularly known as the Cherokee River, among other names ...
. The Dixie chub is
sympatric In biology, two related species or populations are considered sympatric when they exist in the same geographic area and thus frequently encounter one another. An initially interbreeding population that splits into two or more distinct species s ...
with ''S. atromaculatus'' in the rivers draining into
Mobile Bay Mobile Bay ( ) is a shallow inlet of the Gulf of Mexico, lying within the state of Alabama in the United States. Its mouth is formed by the Fort Morgan Peninsula on the eastern side and Dauphin Island, a barrier island on the western side. Th ...
where it is found in the coastal reaches up to the Fall Line but replaces it eastwards from the Conecuh River.


Habitat and ecology

The Dixie chub is found in small, clear headwater streams in the pools of creeks and small rivers with sand or gravel substrates. The fish aggregate to spawn in April. As in ''S. atromaculatus'' and ''S. lumbee'' this species creates breeding structures known as pit/ridge nests. These are constructed in the flowing stretches near pools which have a suitable refuge for the male to escape to, e.g. an undercut bank. The nests starts out as a pit excavated by a dominant male, the male mates with a female and then begins to cover the eggs with small pebbles taken from the substrate starting from the upstream end of the pit. Subsequent matings lead to the eggs being laid down stream of each previous batch of eggs and these are in turn covered by more pebbles and a new pit is excavated downstream, lengthening the ridge. The males guard the nests and display at other similar sized males, by swimming parallel to them. It will also aggessivley lunge at other species which approach the nest. The saffron shiner (''Notropis rubricroceus'') has been recorded associating with nesting Dixie chubs. The Dixie chub is omnivorous and has been recorded eating variety of animal and plant material, including insects, worms, fishes, mollusks, crayfishes, and plant material.


Naming

The Dixie chub was described by the American ichthyologist David Starr Jordan in 1877. The generic name ''Semotilus'' means "spotted banner" and refers to the dorsal fin while the specific name ''thoreauianus'' was given to honor of Henry David Thoreau.


References

{{Taxonbar, from=Q5530913 Semotilus Fish described in 1877