Segestria Florentina
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''Segestria florentina'' is the largest European segestriid
spider Spiders (order (biology), order Araneae) are air-breathing arthropods that have eight limbs, chelicerae with fangs generally able to inject venom, and spinnerets that extrude spider silk, silk. They are the largest order of arachnids and ran ...
. Some vernacular names are green-fanged tube web spider and cellar spider, although the latter is not exclusive to this species.


Description

Females can reach a body length of 22 mm, males up to 15 mm. This species is much darker than others of the same genus. While subadult spiders have a greyish
opisthosoma The opisthosoma is the posterior part of the body in some arthropods, behind the prosoma ( cephalothorax). It is a distinctive feature of the subphylum Chelicerata (arachnids, horseshoe crabs and others). Although it is similar in most respects ...
with a marking similar to '' Segestria senoculata'', adults are of a uniform black, sometimes with a green
iridescent Iridescence (also known as goniochromism) is the phenomenon of certain surfaces that appear gradually to change colour as the angle of view or the angle of illumination changes. Iridescence is caused by wave interference of light in microstruc ...
shine, especially on the
chelicerae The chelicerae () are the arthropod mouthparts, mouthparts of the subphylum Chelicerata, an arthropod group that includes arachnids, horseshoe crabs, and sea spiders. Commonly referred to as "jaws", chelicerae may be shaped as either articulated ...
,Bellmann, H. (1997). Kosmos-Atlas Spinnentiere Europas. ''Kosmos''. which reflect with a striking green. The sexes are similar. Adults occur from June to November.


Habits

They spin a tubular web, often in cracks of buildings. Six or more silken lines radiate from it, and the spider waits in the entrance, touching the lines with the frontal six legs. Prey triggering these lines get caught, and the spider immediately retreats again to eat it. However, in some cases the spider will kill and begin to consume its prey in the opening of the tube, but will retreat further if disturbed. They hunt nocturnal insects such as
moth Moths are a group of insects that includes all members of the order Lepidoptera that are not Butterfly, butterflies. They were previously classified as suborder Heterocera, but the group is Paraphyly, paraphyletic with respect to butterflies (s ...
s and
cockroach Cockroaches (or roaches) are insects belonging to the Order (biology), order Blattodea (Blattaria). About 30 cockroach species out of 4,600 are associated with human habitats. Some species are well-known Pest (organism), pests. Modern cockro ...
es.
Bee Bees are winged insects closely related to wasps and ants, known for their roles in pollination and, in the case of the best-known bee species, the western honey bee, for producing honey. Bees are a monophyletic lineage within the superfamil ...
s and
wasp A wasp is any insect of the narrow-waisted suborder Apocrita of the order Hymenoptera which is neither a bee nor an ant; this excludes the broad-waisted sawflies (Symphyta), which look somewhat like wasps, but are in a separate suborder ...
s are always bitten at the head end, so the sting will face away from the spider. The female lays her eggs inside the tube web. Sometimes she will die after the spiderlings have hatched, and they will eat their mother. The spider can be lured to the entrance by gently touching the triplines with a stick in the evening or at night.


Distribution

Originally a species of the Mediterranean region as far east as
Georgia Georgia most commonly refers to: * Georgia (country), a country in the South Caucasus * Georgia (U.S. state), a state in the southeastern United States Georgia may also refer to: People and fictional characters * Georgia (name), a list of pe ...
, it can now be found in many towns in southern Britain, where they were probably introduced via seaports at least since 1845. It has been found in several places in
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(North Devon) ,
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(Mid Devon),
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,
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, North Somerset, Swindon,
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, Ashford,
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,
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Dover Dover ( ) is a town and major ferry port in Kent, southeast England. It faces France across the Strait of Dover, the narrowest part of the English Channel at from Cap Gris Nez in France. It lies southeast of Canterbury and east of Maidstone. ...
, Hayes,
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,
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,
Whitstable Whitstable () is a town on the north coast of Kent, England, at the convergence of the The Swale, Swale and the Greater Thames Estuary, north of Canterbury and west of Herne Bay, Kent, Herne Bay. The town, formerly known as Whitstable-on-Se ...
,
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, South East London, Vinters Park, Maidstone,
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Exeter Exeter ( ) is a City status in the United Kingdom, cathedral city and the county town of Devon in South West England. It is situated on the River Exe, approximately northeast of Plymouth and southwest of Bristol. In Roman Britain, Exeter w ...
,
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,
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,
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,
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,
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and Newport, where it prefers south-facing walls. Specimens have been found in
Exeter Cathedral Exeter Cathedral, properly known as the Cathedral Church of Saint Peter in Exeter, is an Anglican cathedral, and the seat of the Bishop of Exeter, in the city status in the United Kingdom, city of Exeter, Devon, in South West England. The presen ...
as far back as 1890. In 2023, it was reported that a population was established in
Limerick Limerick ( ; ) is a city in western Ireland, in County Limerick. It is in the Provinces of Ireland, province of Munster and is in the Mid-West Region, Ireland, Mid-West which comprises part of the Southern Region, Ireland, Southern Region. W ...
in western Ireland. It has also been found in
Argentina Argentina, officially the Argentine Republic, is a country in the southern half of South America. It covers an area of , making it the List of South American countries by area, second-largest country in South America after Brazil, the fourt ...
,
Australia Australia, officially the Commonwealth of Australia, is a country comprising mainland Australia, the mainland of the Australia (continent), Australian continent, the island of Tasmania and list of islands of Australia, numerous smaller isl ...
and several
Atlantic The Atlantic Ocean is the second largest of the world's five oceanic divisions, with an area of about . It covers approximately 17% of Earth's surface and about 24% of its water surface area. During the Age of Discovery, it was known for se ...
islands, where it was probably also introduced. Many specimens inhabit
Westminster Abbey Westminster Abbey, formally titled the Collegiate Church of Saint Peter at Westminster, is an Anglican church in the City of Westminster, London, England. Since 1066, it has been the location of the coronations of 40 English and British m ...
in an isolated population group that has been proposed as a new subspecies due to their fangs being brighter green and their size being significantly smaller to other European populations. Egg-laying females are known to prefer the habitation of brick structured buildings. During winter months, females will lay eggs within the bricks of south facing structures. If a female is found between the months of September and October, it is highly likely that she has laid eggs in preparation for the next spring. Infestation of buildings is possible if not found and treated early.


Bite

Its bite is reported to be extremely painful. It has been compared to a "deep injection", and the pain can last for several hours. Two
neurotoxin Neurotoxins are toxins that are destructive to nervous tissue, nerve tissue (causing neurotoxicity). Neurotoxins are an extensive class of exogenous chemical neurological insult (medical), insultsSpencer 2000 that can adversely affect function ...
s and one
insecticide Insecticides are pesticides used to kill insects. They include ovicides and larvicides used against insect eggs and larvae, respectively. The major use of insecticides is in agriculture, but they are also used in home and garden settings, i ...
were found in the venom. The venom reduces the rate and amount of
sodium Sodium is a chemical element; it has Symbol (chemistry), symbol Na (from Neo-Latin ) and atomic number 11. It is a soft, silvery-white, highly reactive metal. Sodium is an alkali metal, being in group 1 element, group 1 of the peri ...
inactivation. Bites are reported to feel like a bee sting, which would make it a 2 on the
Schmidt sting pain index The Schmidt sting pain index is a pain scale rating the relative pain caused by different hymenopteran stings. It is mainly the work of Justin O. Schmidt, who was an entomologist at the Carl Hayden Bee Research Center in Arizona. Schmidt's or ...
, but do not have any lasting effects.


References

{{Taxonbar, from=Q141264 Segestriidae Spiders of Europe Spiders of Asia Spiders of Africa Spiders described in 1790