Rye (''Secale cereale'') is a
grass
Poaceae ( ), also called Gramineae ( ), is a large and nearly ubiquitous family (biology), family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos, the grasses of natural grassland and spe ...
grown extensively as a
grain
A grain is a small, hard, dry fruit (caryopsis) – with or without an attached husk, hull layer – harvested for human or animal consumption. A grain crop is a grain-producing plant. The two main types of commercial grain crops are cereals and ...
, a
cover crop and a
forage
Forage is a plant material (mainly plant leaves and stems) eaten by grazing livestock. Historically, the term ''forage'' has meant only plants eaten by the animals directly as pasture, crop residue, or immature cereal crops, but it is also used m ...
crop. It is grown principally in an area from Eastern and Northern Europe into Russia. It is much more tolerant of cold weather and poor soil than other cereals, making it useful in those regions; its vigorous growth suppresses weeds and provides abundant forage for animals early in the year. It is a member of the wheat tribe (
Triticeae) which includes the cereals
wheat
Wheat is a group of wild and crop domestication, domesticated Poaceae, grasses of the genus ''Triticum'' (). They are Agriculture, cultivated for their cereal grains, which are staple foods around the world. Well-known Taxonomy of wheat, whe ...
and
barley
Barley (), a member of the grass family, is a major cereal grain grown in temperate climates globally. It was one of the first cultivated grains; it was domesticated in the Fertile Crescent around 9000 BC, giving it nonshattering spikele ...
. It is likely that rye arrived in Europe as a
secondary crop, meaning that it was a minor admixture in wheat as a result of
Vavilovian mimicry, and was only later cultivated in its own right.
Rye grain is used for
bread
Bread is a baked food product made from water, flour, and often yeast. It is a staple food across the world, particularly in Europe and the Middle East. Throughout recorded history and around the world, it has been an important part of many cu ...
,
beer
Beer is an alcoholic beverage produced by the brewing and fermentation of starches from cereal grain—most commonly malted barley, although wheat, maize (corn), rice, and oats are also used. The grain is mashed to convert starch in the ...
,
rye whiskey, and animal
fodder
Fodder (), also called provender (), is any agriculture, agricultural foodstuff used specifically to feed domesticated livestock, such as cattle, domestic rabbit, rabbits, sheep, horses, chickens and pigs. "Fodder" refers particularly to food ...
. In Scandinavia, rye was a staple food in the Middle Ages, and rye
crispbread remains a popular food in the region. Europe produces around half of the world's rye; relatively little is traded between countries. A wheat-rye hybrid,
triticale, combines the qualities of the two parent crops and is produced in large quantities worldwide. In European folklore, the ("rye wolf") is a carnivorous corn demon or .
Origins

The rye genus ''
Secale
''Secale'' is a genus of the grass
Poaceae ( ), also called Gramineae ( ), is a large and nearly ubiquitous family (biology), family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos, t ...
'' is in the
grass
Poaceae ( ), also called Gramineae ( ), is a large and nearly ubiquitous family (biology), family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos, the grasses of natural grassland and spe ...
tribe
Triticeae, which contains other
cereal
A cereal is a grass cultivated for its edible grain. Cereals are the world's largest crops, and are therefore staple foods. They include rice, wheat, rye, oats, barley, millet, and maize ( Corn). Edible grains from other plant families, ...
s such as
barley
Barley (), a member of the grass family, is a major cereal grain grown in temperate climates globally. It was one of the first cultivated grains; it was domesticated in the Fertile Crescent around 9000 BC, giving it nonshattering spikele ...
(''Hordeum'') and
wheat
Wheat is a group of wild and crop domestication, domesticated Poaceae, grasses of the genus ''Triticum'' (). They are Agriculture, cultivated for their cereal grains, which are staple foods around the world. Well-known Taxonomy of wheat, whe ...
(''Triticum'').
The generic name ''Secale'', related to Italian and French meaning "rye", is of unknown origin but may derive from a Balkan language. The English name rye derives from Old English , related to Dutch , German , and Russian , again all with the same meaning.
Rye is one of several
cereals that grow wild in the
Levant
The Levant ( ) is the subregion that borders the Eastern Mediterranean, Eastern Mediterranean sea to the west, and forms the core of West Asia and the political term, Middle East, ''Middle East''. In its narrowest sense, which is in use toda ...
, central and eastern
Turkey
Turkey, officially the Republic of Türkiye, is a country mainly located in Anatolia in West Asia, with a relatively small part called East Thrace in Southeast Europe. It borders the Black Sea to the north; Georgia (country), Georgia, Armen ...
and adjacent areas. Evidence uncovered at the
Epipalaeolithic
In archaeology, the Epipalaeolithic or Epipaleolithic (sometimes Epi-paleolithic etc.) is a period occurring between the Upper Paleolithic and Neolithic during the Stone Age. Mesolithic also falls between these two periods, and the two are someti ...
site of
Tell Abu Hureyra in the
Euphrates
The Euphrates ( ; see #Etymology, below) is the longest and one of the most historically important rivers of West Asia. Tigris–Euphrates river system, Together with the Tigris, it is one of the two defining rivers of Mesopotamia (). Originati ...
valley of northern
Syria
Syria, officially the Syrian Arab Republic, is a country in West Asia located in the Eastern Mediterranean and the Levant. It borders the Mediterranean Sea to the west, Turkey to Syria–Turkey border, the north, Iraq to Iraq–Syria border, t ...
suggests that rye was among the first cereal crops to be systematically cultivated, around 13,000 years ago.
However, that claim remains controversial; critics point to inconsistencies in the
radiocarbon dates, and identifications based solely on grain, rather than on
chaff
Chaff (; ) is dry, scale-like plant material such as the protective seed casings of cereal grains, the scale-like parts of flowers, or finely chopped straw. Chaff cannot be digested by humans, but it may be fed to livestock, ploughed into soil ...
.
Domesticated rye occurs in small quantities at a number of
Neolithic
The Neolithic or New Stone Age (from Ancient Greek, Greek 'new' and 'stone') is an archaeological period, the final division of the Stone Age in Mesopotamia, Asia, Europe and Africa (c. 10,000 BCE to c. 2,000 BCE). It saw the Neolithic Revo ...
sites in Asia Minor (
Anatolia
Anatolia (), also known as Asia Minor, is a peninsula in West Asia that makes up the majority of the land area of Turkey. It is the westernmost protrusion of Asia and is geographically bounded by the Mediterranean Sea to the south, the Aegean ...
, now Turkey), such as the
Pre-Pottery Neolithic B
Pre-Pottery Neolithic B (PPNB) is part of the Pre-Pottery Neolithic, a Neolithic culture centered in upper Mesopotamia and the Levant, dating to years ago, that is, 8800–6500 BC. It was Type site, typed by British archaeologist Kathleen Kenyon ...
Can Hasan III near
Çatalhöyük,
but is otherwise absent from the archaeological record until the
Bronze Age
The Bronze Age () was a historical period characterised principally by the use of bronze tools and the development of complex urban societies, as well as the adoption of writing in some areas. The Bronze Age is the middle principal period of ...
of central Europe, c. 1800–1500 BCE.
It is likely that rye was brought westwards from Asia Minor as a
secondary crop, meaning that it was a minor admixture in wheat as a result of
Vavilovian mimicry, and was only later cultivated in its own right.
Archeological evidence of this grain has been found in
Roman contexts along the
Rhine
The Rhine ( ) is one of the List of rivers of Europe, major rivers in Europe. The river begins in the Swiss canton of Graubünden in the southeastern Swiss Alps. It forms part of the Swiss-Liechtenstein border, then part of the Austria–Swit ...
and the
Danube
The Danube ( ; see also #Names and etymology, other names) is the List of rivers of Europe#Longest rivers, second-longest river in Europe, after the Volga in Russia. It flows through Central and Southeastern Europe, from the Black Forest sou ...
and in Ireland and Britain.
[ page 26.] The Roman naturalist
Pliny the Elder
Gaius Plinius Secundus (AD 23/24 79), known in English as Pliny the Elder ( ), was a Roman Empire, Roman author, Natural history, naturalist, and naval and army commander of the early Roman Empire, and a friend of the Roman emperor, emperor Vesp ...
was dismissive of a grain that may have been rye, writing that it "is a very poor food and only serves to avert starvation".
He said it was mixed with
spelt
Spelt (''Triticum spelta''), also known as dinkel wheat is a species of wheat. It is a relict crop, eaten in Central Europe and northern Spain. It is high in protein and may be considered a health food.
Spelt was cultivated from the Neolit ...
"to mitigate its bitter taste, and even then is most unpleasant to the stomach".
Description
Rye is a tall grass grown for its seeds; it can be an
annual or a
biennial. Depending on environmental conditions and variety it reaches in height. Its leaves are blue-green, long, and pointed. The seeds are carried in a curved head or spike some long. The head is composed of many
spikelet
A spikelet, in botany, describes the typical arrangement of the inflorescences of grasses, sedges and some other monocots.
Each spikelet has one or more florets. The spikelets are further grouped into panicles or spikes. The part of the sp ...
s, each of which holds two small flowers; the spikelets alternate left and right up the head.
File:Illustration Secale cereale0.jpg, Botanical illustration
File:Secale cereale - cereal rye 2 - Steve Hurst USDA-NRCS PLANTS Database.jpg, Seed in husk
Husk (or hull) in botany is the outer shell or coating of a seed. In the United States, the term husk often refers to the leafy outer covering of an Ear (botany), ear of maize (corn) as it grows on the plant. Literally, a husk or hull includes t ...
File:Secale cereale - cereal rye - Steve Hurst USDA-NRCS PLANTS Database.jpg, Different types of grains
File:Rye grains rotated (cropped).jpg, The seeds of rye are some 7 or 8 mm long, much larger and less round than wheat
Wheat is a group of wild and crop domestication, domesticated Poaceae, grasses of the genus ''Triticum'' (). They are Agriculture, cultivated for their cereal grains, which are staple foods around the world. Well-known Taxonomy of wheat, whe ...
.
Cultivation
Since the
Middle Ages
In the history of Europe, the Middle Ages or medieval period lasted approximately from the 5th to the late 15th centuries, similarly to the post-classical period of global history. It began with the fall of the Western Roman Empire and ...
, people have cultivated rye widely in
Central and
Eastern Europe
Eastern Europe is a subregion of the Europe, European continent. As a largely ambiguous term, it has a wide range of geopolitical, geographical, ethnic, cultural and socio-economic connotations. Its eastern boundary is marked by the Ural Mountain ...
. It serves as the main
bread
Bread is a baked food product made from water, flour, and often yeast. It is a staple food across the world, particularly in Europe and the Middle East. Throughout recorded history and around the world, it has been an important part of many cu ...
cereal in most areas east of the
France–Germany border
The international border between the modern states of France and Germany has a length of . The southern portion of the border, between Saint-Louis at the border with Switzerland and Lauterbourg, follows the River Rhine (Upper Rhine) in a so ...
and north of
Hungary
Hungary is a landlocked country in Central Europe. Spanning much of the Pannonian Basin, Carpathian Basin, it is bordered by Slovakia to the north, Ukraine to the northeast, Romania to the east and southeast, Serbia to the south, Croatia and ...
. In Southern Europe, it was cultivated on marginal lands.
Rye grows well in much poorer soils than those necessary for most cereal grains. Thus, it is an especially valuable crop in regions where the soil has
sand
Sand is a granular material composed of finely divided mineral particles. Sand has various compositions but is usually defined by its grain size. Sand grains are smaller than gravel and coarser than silt. Sand can also refer to a textural ...
or
peat
Peat is an accumulation of partially Decomposition, decayed vegetation or organic matter. It is unique to natural areas called peatlands, bogs, mires, Moorland, moors, or muskegs. ''Sphagnum'' moss, also called peat moss, is one of the most ...
. Rye plants withstand cold better than other small grains, surviving snow cover that would kill winter wheat. Winter rye is the most popular: it is planted and begins to grow in autumn. In spring, the plants develop rapidly.
This allows it to provide spring grazing, at a time when spring-planted wheat has only just germinated.
The physical properties of rye affect attributes of the final food product such as seed size, surface area, and porosity. The surface area of the seed directly correlates to the drying and heat transfer time.
Smaller seeds have increased heat transfer, which leads to lower drying time. Seeds with lower porosity lose water more slowly during the process of drying.
Rye is
harvest
Harvesting is the process of collecting plants, animals, or fish (as well as fungi) as food, especially the process of gathering mature crops, and "the harvest" also refers to the collected crops. Reaping is the cutting of grain or pulses fo ...
ed like wheat with a
combine harvester
The modern combine harvester, also called a combine, is a machine designed to harvest a variety of cultivated seeds. Combine harvesters are one of the most economically important labour-saving inventions, significantly reducing the fraction of ...
, which cuts the plants,
threshes and
winnows the grain, and releases the
straw
Straw is an agricultural byproduct consisting of the dry wikt:stalk, stalks of cereal plants after the grain and chaff have been removed. It makes up about half of the crop yield, yield by weight of cereal crops such as barley, oats, rice, ry ...
to the field where it is later pressed into bales or left as soil amendment. The resultant grain is stored in local
silo
A silo () is a structure for storing Bulk material handling, bulk materials.
Silos are commonly used for bulk storage of grain, coal, cement, carbon black, woodchips, food products and sawdust. Three types of silos are in widespread use toda ...
s or transported to regional
grain elevators and combined with other lots for storage and distant shipment. Before the era of
mechanised agriculture
Mechanised agriculture or agricultural mechanization is the use of machinery and equipment, ranging from simple and basic hand tools to more sophisticated, motorized equipment and machinery, to perform agricultural operations. In modern times, po ...
, rye harvesting was a
manual task performed with
scythe
A scythe (, rhyming with ''writhe'') is an agriculture, agricultural hand-tool for mowing grass or Harvest, harvesting Crop, crops. It was historically used to cut down or reaping, reap edible grain, grains before they underwent the process of ...
s or
sickle
A sickle, bagging hook, reaping-hook or grasshook is a single-handed agricultural tool designed with variously curved blades and typically used for harvesting or reaping grain crops, or cutting Succulent plant, succulent forage chiefly for feedi ...
s.
Agroecology
Winter rye is any breed of rye planted in the autumn to provide ground cover for the winter. It grows during warmer days of the winter when sunlight temporarily warms the plant above freezing, even while there is general snow cover. It can be used as a cover crop to prevent the growth of
winter-hardy weeds.
Rye grows better than any other cereal in heavy
clay
Clay is a type of fine-grained natural soil material containing clay minerals (hydrous aluminium phyllosilicates, e.g. kaolinite, ). Most pure clay minerals are white or light-coloured, but natural clays show a variety of colours from impuriti ...
and light sandy soil, and infertile or drought-affected soils. It can tolerate
pH between 4.5 and 8.0, but soils having pH 5.0 to 7.0 are best suited for rye cultivation. Rye grows best in fertile, well-drained
loam
Loam (in geology and soil science) is soil composed mostly of sand (particle size > ), silt (particle size > ), and a smaller amount of clay (particle size < ). By weight, its mineral composition is about 40–40–20% concentration of sand–si ...
or clay-loam soils. As for temperature, the crop can thrive in subzero environments, assisted by the production of
antifreeze polypeptides (different from those produced by some fish and insects) by the leaves of winter rye.
Rye is a common,
unwanted invader of
winter wheat fields. If allowed to grow and mature, it may cause substantially reduced prices (docking) for harvested wheat.
Pests and diseases

Pests including the nematode ''
Ditylenchus dipsaci
''Ditylenchus'' is a genus of plant pathogenic nematodes.
References
Tylenchida
Secernentea genera
Agricultural pest nematodes
{{agri-stub ...
'' and a variety of herbivorous insects can seriously affect plant health.
Rye is highly susceptible to the
ergot fungus. Consumption of ergot-infected rye by humans and animals results in
ergotism, which causes convulsions, miscarriage, necrosis of digits, hallucinations and death. Historically, damp northern countries that depended on rye as a staple crop were subject to periodic epidemics.
Modern grain-cleaning and milling methods have practically eliminated ergotism, but it remains a risk if food safety vigilance breaks down.
After an absence of 60 years,
stem rust (''Puccinia graminis'' f. sp. ''tritici'') has returned to
Europe
Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...
in the 2020s.
Areas affected include
Germany
Germany, officially the Federal Republic of Germany, is a country in Central Europe. It lies between the Baltic Sea and the North Sea to the north and the Alps to the south. Its sixteen States of Germany, constituent states have a total popu ...
,
Russia
Russia, or the Russian Federation, is a country spanning Eastern Europe and North Asia. It is the list of countries and dependencies by area, largest country in the world, and extends across Time in Russia, eleven time zones, sharing Borders ...
(
Western Siberia
Western Siberia or West Siberia ( rus, Западная Сибирь, p=ˈzapədnəjə sʲɪˈbʲirʲ; , ) is a region in North Asia. It is part of the wider region of Siberia that is mostly located in the Russia, Russian Federation, with a Sout ...
),
Spain
Spain, or the Kingdom of Spain, is a country in Southern Europe, Southern and Western Europe with territories in North Africa. Featuring the Punta de Tarifa, southernmost point of continental Europe, it is the largest country in Southern Eur ...
, and
Sweden
Sweden, formally the Kingdom of Sweden, is a Nordic countries, Nordic country located on the Scandinavian Peninsula in Northern Europe. It borders Norway to the west and north, and Finland to the east. At , Sweden is the largest Nordic count ...
.
Production and consumption

Rye is grown primarily in Eastern, Central and Northern Europe. The main rye belt stretches from northern Germany through Poland, Ukraine, and eastwards into central and northern Russia. Rye is also grown in North America, in South America including Argentina, in Oceania (Australia and New Zealand), in Turkey, and in northern China. Production levels of rye have fallen since 1992 in most of the producing nations, ; for instance, production of rye in Russia fell from 13.9 metric tons in 1992 to 2.2 metric tons in 2022.
World trade of rye is low compared with other grains such as wheat. The total export of rye for 2016 was $186 million compared with $30.1 billion for wheat.
Poland consumes the most rye per person at per capita (2009), followed by the Nordic and Baltic countries. The EU in general is around per capita. The entire world only consumes per capita.
Nutritional value
Raw rye contains 11% water, 76%
carbohydrate
A carbohydrate () is a biomolecule composed of carbon (C), hydrogen (H), and oxygen (O) atoms. The typical hydrogen-to-oxygen atomic ratio is 2:1, analogous to that of water, and is represented by the empirical formula (where ''m'' and ''n'' ...
s, 10%
protein
Proteins are large biomolecules and macromolecules that comprise one or more long chains of amino acid residue (biochemistry), residues. Proteins perform a vast array of functions within organisms, including Enzyme catalysis, catalysing metab ...
, and 2%
fat
In nutrition science, nutrition, biology, and chemistry, fat usually means any ester of fatty acids, or a mixture of such chemical compound, compounds, most commonly those that occur in living beings or in food.
The term often refers specif ...
(table). A reference amount of rye provides of
food energy
Food energy is chemical energy that animals and humans derive from food to sustain their metabolism and muscular activity.
Most animals derive most of their energy from aerobic respiration, namely combining the carbohydrates, fats, and protein ...
, and is a rich source (20% or more of the
Daily Value
In the U.S. and Canada, the Reference Daily Intake (RDI) is used in nutrition labeling on food and dietary supplement products to indicate the daily intake level of a nutrient that is considered to be sufficient to meet the requirements of 97� ...
, DV) of
essential nutrients, including
dietary fiber
Dietary fiber (fibre in English in the Commonwealth of Nations, Commonwealth English) or roughage is the portion of plant-derived food that cannot be completely broken down by human digestive enzymes. Dietary fibers are diverse in chemical co ...
,
B vitamins
B vitamins are a class of water-soluble vitamins that play important roles in Cell (biology), cell metabolism and synthesis of red blood cells. They are a chemically diverse class of compounds.
Dietary supplements containing all eight are referr ...
, such as
thiamine
Thiamine, also known as thiamin and vitamin B1, is a vitamin – an Nutrient#Micronutrients, essential micronutrient for humans and animals. It is found in food and commercially synthesized to be a dietary supplement or medication. Phosp ...
and
niacin (each at 25% DV), and several
dietary minerals
In the context of nutrition, a mineral is a chemical element. Some "minerals" are essential for life, but most are not. ''Minerals'' are one of the four groups of essential nutrients; the others are vitamins, essential fatty acids, and essen ...
. Highest micronutrient contents are for
manganese
Manganese is a chemical element; it has Symbol (chemistry), symbol Mn and atomic number 25. It is a hard, brittle, silvery metal, often found in minerals in combination with iron. Manganese was first isolated in the 1770s. It is a transition m ...
(130% DV) and
phosphorus
Phosphorus is a chemical element; it has Chemical symbol, symbol P and atomic number 15. All elemental forms of phosphorus are highly Reactivity (chemistry), reactive and are therefore never found in nature. They can nevertheless be prepared ar ...
(27% DV) (table).
Health effects
According to
Health Canada
Health Canada (HC; )Health Canada is the applied title under the Federal Identity Program; the legal title is Department of Health (). is the Structure of the Canadian federal government#Departments, with subsidiary units, department of the Gove ...
and the U.S.
Food and Drug Administration
The United States Food and Drug Administration (FDA or US FDA) is a List of United States federal agencies, federal agency of the United States Department of Health and Human Services, Department of Health and Human Services. The FDA is respo ...
, consuming at least per day of rye
beta-glucan or per serving of
soluble fiber can lower levels of
blood cholesterol, a risk factor for
cardiovascular diseases
Cardiovascular disease (CVD) is any disease involving the heart or blood vessels. CVDs constitute a class of diseases that includes: coronary artery diseases (e.g. angina, heart attack), heart failure, hypertensive heart disease, rheuma ...
.
Eating whole-grain rye, as well as other high-fibre grains, improves regulation of
blood sugar
The blood sugar level, blood sugar concentration, blood glucose level, or glycemia is the measure of glucose concentrated in the blood. The body tightly regulates blood glucose levels as a part of metabolic homeostasis.
For a 70 kg (1 ...
(i.e., reduces blood glucose response to a meal).
Consuming
breakfast cereal
Breakfast cereal is a category of food, including food products, made from food processing, processed cereal, cereal grains, that are eaten as part of breakfast or as a snack food, primarily in Western societies.
Although warm, cooked cereals li ...
s containing rye over weeks to months also improved cholesterol levels and glucose regulation.
Health concerns
Like wheat, barley, and their hybrids and derivatives, rye contains
gluten
Gluten is a structural protein naturally found in certain Cereal, cereal grains. The term ''gluten'' usually refers to the elastic network of a wheat grain's proteins, gliadin and glutenin primarily, that forms readily with the addition of water ...
s and related prolamines, which makes it an unsuitable grain for consumption by people with
gluten-related disorders
Gluten-related disorders is the term for the diseases triggered by gluten, including celiac disease (CD), non-celiac gluten sensitivity (NCGS), gluten ataxia, dermatitis herpetiformis (DH) and wheat allergy. The umbrella category has also been r ...
, such as
celiac disease
Coeliac disease (British English) or celiac disease (American English) is a long-term autoimmune disorder, primarily affecting the small intestine. Patients develop intolerance to gluten, which is present in foods such as wheat, rye, spel ...
,
non-celiac gluten sensitivity
Non-celiac gluten sensitivity (NCGS) or gluten sensitivity is a controversial disorder which can cause both gastrointestinal and other problems.
NCGS is included in the spectrum of gluten-related disorders. The definition and diagnostic criteria ...
, and
wheat allergy, among others.
Nevertheless, some wheat allergy patients can tolerate rye or barley.
Uses
Food and drink
Rye grain is refined into a
flour
Flour is a powder made by Mill (grinding), grinding raw grains, List of root vegetables, roots, beans, Nut (fruit), nuts, or seeds. Flours are used to make many different foods. Cereal flour, particularly wheat flour, is the main ingredie ...
high in
gliadin
Gliadin (a type of prolamin) is a class of proteins present in wheat and several other cereals within the grass genus ''Triticum''. Gliadins, which are a component of gluten, are essential for giving bread the ability to rise properly during ba ...
but low in
glutenin
Glutenin (a type of glutelin) is a major protein within wheat flour, making up 47% of the total protein content. The glutenins are protein aggregates of high- molecular-mass (HMW) and low-molecular-mass (LMW) subunits with molar masses from ab ...
and rich in soluble fiber.
Alkylresorcinols are phenolic lipids present in high amounts in the bran layer (e.g.
pericarp
Fruits are the mature ovary or ovaries of one or more flowers. They are found in three main anatomical categories: aggregate fruits, multiple fruits, and simple fruits.
Fruitlike structures may develop directly from the seed itself rather th ...
,
testa and
aleurone layers) of wheat and rye (0.1–0.3% of dry weight).
Rye bread, including
pumpernickel, is made using rye flour and is a widely eaten food in Northern and Eastern Europe.
In Scandinavia, rye is widely used to make
crispbread (); in the Middle Ages it was a
staple food
A staple food, food staple, or simply staple, is a food that is eaten often and in such quantities that it constitutes a dominant portion of a standard diet for an individual or a population group, supplying a large fraction of energy needs an ...
in the region, and it remains popular in the 21st century.
Rye grain is used to make alcoholic drinks, such as
rye whiskey and
rye beer.
The traditional cloudy and sweet-sour low-alcohol beverage
kvass
Kvass is a fermented, cereal-based, low-alcoholic beverage of cloudy appearance and sweet-sour taste.
Kvass originates from northeastern Europe, where grain production was considered insufficient for beer to become a daily drink. The first wr ...
is fermented from rye bread or rye flour and malt.
Oval rye pumpkin bread 2024-03-10 08.jpg, Rye bread
Naturaplan Bio Pumpernickel.png, Pumpernickel
IKEA knäckebröd.JPG, Swedish rye crispbread ()
File:Sultsina.png, Sultsina, a traditional Karelia
Karelia (; Karelian language, Karelian and ; , historically Коре́ла, ''Korela'' []; ) is an area in Northern Europe of historical significance for Russia (including the Soviet Union, Soviet era), Finland, and Sweden. It is currentl ...
n dish made of unleavened rye dough and a farina filling
Old Overholt Rye Whiskey bottle and tumbler.jpg, Rye whiskey
Terrapin Brewing Co. Rye Squared Imperial Pale Ale.jpg, Rye beer
Other uses
Rye is a useful
forage
Forage is a plant material (mainly plant leaves and stems) eaten by grazing livestock. Historically, the term ''forage'' has meant only plants eaten by the animals directly as pasture, crop residue, or immature cereal crops, but it is also used m ...
crop in cool climates; it grows vigorously and provides plentiful
fodder
Fodder (), also called provender (), is any agriculture, agricultural foodstuff used specifically to feed domesticated livestock, such as cattle, domestic rabbit, rabbits, sheep, horses, chickens and pigs. "Fodder" refers particularly to food ...
for grazing animals, or
green manure
In agriculture, a green manure is a crop specifically cultivated to be incorporated into the soil while still green. Typically, the green manure's Biomass (ecology), biomass is incorporated with a plow or disk, as is often done with (brown) man ...
to improve the soil. It forms a good
cover crop in winter with its rapid growth and deep roots.
Rye straw is used as
livestock bedding Bedding, in ethology and animal husbandry, is material, usually organic, used by animals to support their bodies when resting or otherwise stationary. It reduces pressure on skin, heat loss, and contamination by waste produced by an animal or tho ...
, despite the risk of ergot poisoning. It is used on a small scale to make
craft
A craft or trade is a pastime or an occupation that requires particular skills and knowledge of skilled work. In a historical sense, particularly the Middle Ages and earlier, the term is usually applied to people occupied in small scale pr ...
s such as
corn dollies. More recently it has found uses as a raw material for bioconversion to products such as the sweetener
xylitol
Xylitol is a chemical compound with the formula , or HO(CH2)(CHOH)3(CH2)OH; specifically, one particular Stereoisomerism, stereoisomer with that structural formula. It is a colorless or white crystalline solid. It is classified as a polyalcoho ...
.
Rye flour is boiled with
red iron oxide pigments and some additives to make traditional
Falun red paint, widely used as a house paint in Sweden and other Scandinavian countries.
Production of hybrids
Plant breeders, starting in the 19th century in Germany and Scotland, but mainly from the 1950s, worked to develop a hybrid cereal with the best qualities of wheat and rye, now called
triticale. Modern triticales are
hexaploid with six sets of chromosomes; they are used to produce millions of tons of cereal annually.
Varieties of rye hold much genetic diversity,
[ which cites ] which can be used to improve other crops such as wheat. For example, the pollination abilities of wheat can be improved by the addition of the rye chromosome 4R; this increases the size of the wheat
anther
The stamen (: stamina or stamens) is a part consisting of the male reproductive organs of a flower. Collectively, the stamens form the androecium., p. 10
Morphology and terminology
A stamen typically consists of a stalk called the filament ...
and the amount of pollen. The chromosome is the source of many
crop disease resistance genes.
Varieties such as Petkus, Insave, Amigo, and Imperial have donated 1R-originating resistance to wheat.
AC Hazlet rye is a medium-sized winter rye with resistance to both
lodging
Lodging refers to the use of a short-term dwelling, usually by renting the living space or sometimes through some other arrangement. People who travel and stay away from home for more than a day need lodging for sleep, rest, food, safety, shel ...
and
shattering. Rye was the
gene donor of ''
Sr31'' – a
stem rust resistance gene – introgressed into wheat.
The characteristics of ''S. cereale'' have been combined with another perennial rye, ''
S. montanum'', to produce ''
S. cereanum'', which has the beneficial characteristics of each. The hybrid rye can be grown in harsh environments and on poor soil. It provides improved forage with digestible fiber and protein.
In human culture

In European folklore, the ''Roggenwolf'' ("rye wolf") is a carnivorous corn demon or ''
Feldgeist'', a field spirit shaped like a
wolf
The wolf (''Canis lupus''; : wolves), also known as the grey wolf or gray wolf, is a Canis, canine native to Eurasia and North America. More than thirty subspecies of Canis lupus, subspecies of ''Canis lupus'' have been recognized, includin ...
.
The ''Roggenwolf'' steals children and feeds on them. The last grain heads are often left at their place as a sacrifice for the agricultural spirits.
In contrast, the ''Roggenmuhme'' or ''Roggenmutter'' ("rye aunt" or "rye mother") is an anthropomorphic female corn demon with fiery fingers. Her bosoms are filled with tar and may end in tips of iron. Her bosoms are also long, and as such must be thrown over her shoulders when she runs. The ''Roggenmuhme'' is completely black or white, and in her hand she has a birch or whip from which lightning sparks. She can change herself into different animals, such as snakes, turtles, and frogs.
The classical scholar
Carl A. P. Ruck
Carl Anton Paul Ruck (born December 8, 1935, Bridgeport, Connecticut) is a professor in the Classical Studies department at Boston University. He received his B.A. at Yale University, his M.A. at the University of Michigan, and a Ph.D. at Harvar ...
writes that the ''Roggenmutter'' was believed to go through the fields, rustling like the wind, with a pack of rye wolves running after her. They spread ergot through the sheaves of harvested rye. According to Ruck, they then lured children into the fields to nurse on the infected grains "like the iron teats of the ''Roggenmutter''".
The enlarged reddish ergot-infected grains were known as ''Wulfzähne'' (wolf teeth).
References
External links
{{Authority control
Rye
Rye (''Secale cereale'') is a grass grown extensively as a grain, a cover crop and a forage crop. It is grown principally in an area from Eastern and Northern Europe into Russia. It is much more tolerant of cold weather and poor soil than o ...
Cereals
Plants described in 1753
Pooideae
Staple foods