Anarhichadidae, the wolffishes, sea wolves or wolf eels, is a family of marine ray finned fishes belonging to the

order Order, ORDER or Orders may refer to: * A socio-political or established or existing order, e.g. World order, Ancien Regime, Pax Britannica * Categorization, the process in which ideas and objects are recognized, differentiated, and understood ...

Perciformes Perciformes (), also called the Acanthopteri, is an order or superorder of ray-finned fish in the clade Percomorpha. ''Perciformes'' means " perch-like". Among the well-known members of this group are perches and darters ( Percidae), and als ...

. These are predatory, eel shaped fishes which are native to the cold waters of the Arctic, North Pacific and North Atlantic Oceans.

Taxonomy

Anarhichadidae was first proposed as a family in 1832 by the French

zoologist Zoology ( , ) is the scientific study of animals. Its studies include the structure, embryology, classification, habits, and distribution of all animals, both living and extinct, and how they interact with their ecosystems. Zoology is one ...

Charles Lucien Bonaparte Charles Lucien Jules Laurent Bonaparte, 2nd Prince of Canino and Musignano (24 May 1803 – 29 July 1857) was a French naturalist and ornithology, ornithologist, and a nephew of Napoleon. Lucien and his wife had twelve children, including Cardinal ...

. The 5th edition of ''

Fishes of the World ''Fishes of the World'' is a standard reference for the systematics of fishes. It was first written in 1976 by the American ichthyologist Joseph S. Nelson (1937–2011). Now in its fifth edition (2016), the work is a comprehensive overview of t ...

'' classifies this family within the

suborder Order () is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between family and class. In biological classification, the order is a taxonomic rank used in the classification of organisms and recognized ...

Zoarcoidei Zoarcoidei is a suborder of marine ray-finned fishes belonging to the Order (biology), order Perciformes. The suborder includes the wolffishes, Pholidae, gunnels and eelpouts. The suborder includes about 400 species. These fishes are predominantl ...

, within the order Scorpaeniformes. Other authorities classify this family in the

infraorder Order () is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between Family_(biology), family and Class_(biology), class. In biological classification, the order is a taxonomic rank used in the classific ...

Zoarcales within the suborder Cottoidei of the

because removing the Scorpaeniformes from the Perciformes renders that taxon non

monophyletic In biological cladistics for the classification of organisms, monophyly is the condition of a taxonomic grouping being a clade – that is, a grouping of organisms which meets these criteria: # the grouping contains its own most recent co ...

Etymology

Anarhichadidae is derived from the name of its

type genus In biological taxonomy, the type genus (''genus typica'') is the genus which defines a biological family and the root of the family name. Zoological nomenclature According to the International Code of Zoological Nomenclature, "The name-bearin ...

''Anarhichas'' which is an

Ancient Greek Ancient Greek (, ; ) includes the forms of the Greek language used in ancient Greece and the classical antiquity, ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Greek ...

name for the Atlantic wolffish (''A. lupus'') and means "the climber", in turn derived from the Greek ''anarrhichesis'' which means "to climb or scramble up". This may be an allusion to the ancient belief that wolffishes left the water and climbed up on the rocks.

Genera and species

Anarhichadidae contains two genera and five species:

Timeline of genera

ImageSize = width:1000px height:auto barincrement:15px PlotArea = left:10px bottom:50px top:10px right:10px Period = from:-65.5 till:10 TimeAxis = orientation:horizontal ScaleMajor = unit:year increment:5 start:-65.5 ScaleMinor = unit:year increment:1 start:-65.5 TimeAxis = orientation:hor AlignBars = justify Colors = #legends id:CAR value:claret id:ANK value:rgb(0.4,0.3,0.196) id:HER value:teal id:HAD value:green id:OMN value:blue id:black value:black id:white value:white id:cenozoic value:rgb(0.54,0.54,0.258) id:paleogene value:rgb(0.99,0.6,0.32) id:paleocene value:rgb(0.99,0.65,0.37) id:eocene value:rgb(0.99,0.71,0.42) id:oligocene value:rgb(0.99,0.75,0.48) id:neogene value:rgb(0.999999,0.9,0.1) id:miocene value:rgb(0.999999,0.999999,0) id:pliocene value:rgb(0.97,0.98,0.68) id:quaternary value:rgb(0.98,0.98,0.5) id:pleistocene value:rgb(0.999999,0.95,0.68) id:holocene value:rgb(0.999,0.95,0.88) BarData= bar:eratop bar:space bar:periodtop bar:space bar:NAM1 bar:NAM2 bar:space bar:period bar:space bar:era PlotData= align:center textcolor:black fontsize:M mark:(line,black) width:25 shift:(7,-4) bar:periodtop from: -65.5 till: -55.8 color:paleocene text:

Paleocene The Paleocene ( ), or Palaeocene, is a geological epoch (geology), epoch that lasted from about 66 to 56 mya (unit), million years ago (mya). It is the first epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), ...

from: -55.8 till: -33.9 color:eocene text:

Eocene The Eocene ( ) is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (Ma). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes ...

from: -33.9 till: -23.03 color:oligocene text:

Oligocene The Oligocene ( ) is a geologic epoch (geology), epoch of the Paleogene Geologic time scale, Period that extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that defin ...

from: -23.03 till: -5.332 color:miocene text:

Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...

from: -5.332 till: -2.588 color:pliocene text: Plio. from: -2.588 till: -0.0117 color:pleistocene text: Pleist. from: -0.0117 till: 0 color:holocene text: H. bar:eratop from: -65.5 till: -23.03 color:paleogene text:

Paleogene The Paleogene Period ( ; also spelled Palaeogene or Palæogene) is a geologic period and system that spans 43 million years from the end of the Cretaceous Period Ma (million years ago) to the beginning of the Neogene Period Ma. It is the fir ...

from: -23.03 till: -2.588 color:neogene text:

Neogene The Neogene ( ,) is a geologic period and system that spans 20.45 million years from the end of the Paleogene Period million years ago ( Mya) to the beginning of the present Quaternary Period million years ago. It is the second period of th ...

from: -2.588 till: 0 color:quaternary text: Q. PlotData= align:left fontsize:M mark:(line,white) width:5 anchor:till align:left color:pliocene bar:NAM1 from: -5.332 till: 0 text:

Anarhichas ''Anarhichas'' is a genus of marine ray-finned fishes belonging to the Family (biology), family Anarhichadidae, the wolffishes or sea wolves. These fishes are found in the northern North Atlantic and North Pacific oceans. Taxonomy ''Anarhichas ...

color:pleistocene bar:NAM2 from: -2.588 till: 0 text: Anarrhichthys PlotData= align:center textcolor:black fontsize:M mark:(line,black) width:25 bar:period from: -65.5 till: -55.8 color:paleocene text:

from: -55.8 till: -33.9 color:eocene text:

from: -33.9 till: -23.03 color:oligocene text:

from: -23.03 till: -5.332 color:miocene text:

from: -5.332 till: -2.588 color:pliocene text: Plio. from: -2.588 till: -0.0117 color:pleistocene text: Pleist. from: -0.0117 till: 0 color:holocene text: H. bar:era from: -65.5 till: -23.03 color:paleogene text:

from: -23.03 till: -2.588 color:neogene text:

from: -2.588 till: 0 color:quaternary text: Q.

Characteristics

Anarhichadidae wolfishes have a largely compressed and, in the genus ''Anarhichas'', a moderately elongate body. ''Anarrhichthys'' has an extremely elongate body, and this has given rise to its common name of wolf-eel. The long

dorsal fin A dorsal fin is a fin on the back of most marine and freshwater vertebrates. Dorsal fins have evolved independently several times through convergent evolution adapting to marine environments, so the fins are not all homologous. They are found ...

starts at the head and has many flexible spines and soft rays. The

anal fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported o ...

may have a single spine in ''Anarrhichthys'', and again there is a large number of soft rays. The

caudal fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported only ...

is separate from the other median fins in ''Anarhichas'' but they are all three joined in ''Anarrhichthys''. There is a single pair of nostrils. The scales, if present, are cycloid, tiny and do not overlap. There are well developed. movement sensitive sensory canals on the head and as the fish ages the pores grow very large. There are 1 or 2

lateral line The lateral line, also called the lateral line organ (LLO), is a system of sensory organs found in fish, used to detect movement, vibration, and pressure gradients in the surrounding water. The sensory ability is achieved via modified epithelia ...

s made up of superficial neuromasts. There are robust conical teeth in the front of the jaws and large molar like teeth to the rear of those. The gill membranes are joined to the isthmus and the gill openings are set widely apart. There is no

swim bladder The swim bladder, gas bladder, fish maw, or air bladder is an internal gas-filled organ (anatomy), organ in bony fish that functions to modulate buoyancy, and thus allowing the fish to stay at desired water depth without having to maintain lift ...

. The counts of vertebrae are 72–89 in ''Anarhichas'' and 221–251 in ''Anarrhichthys''. The longest published

total length Fish measurement is the measuring of individual fish and various parts of their anatomies, for data used in many areas of ichthyology, including taxonomy and fishery biology. Overall length Standard length (SL) is the length of a fish measured f ...

is for ''Anarrhichthys ocellatus'' and is .

Distribution and habitat

Anarhichadidae wolfishes prefer cooler waters and are found in the northern parts of the North Pacific and North Atlantic Oceans as well as in the Arctic Ocean. They are

demersal fish Demersal fish, also known as groundfish, live and feed on or near the bottom of seas or lakes (the demersal zone).Walrond Carl . "Coastal fish - Fish of the open sea floor"Te Ara - the Encyclopedia of New Zealand. Updated 2 March 2009 They oc ...

es occurring in shallow to moderately deep and cold seas.

Biology

Anarhichadidae wolffishes use their large teeth to feed on a diet of shelled invertebrates such as crabs, starfishes and sea urchins, as well as other prey, The peak mating season for wolffish is September to October. The male wolffish will guard the eggs 3–9 months until they hatch.

Fisheries

Anarhichadidae wolffishes, in particular two Atlantic species, the spotted wolffish and the Atlantic wolffish, are targeted by commercial fisheries. The flesh is used for food and the skin to make leather.

Trivia

The collegiate sports teams of the University of Alaska Anchorage have adopted the Seawolf, based on

Tlingit The Tlingit or Lingít ( ) are Indigenous peoples of the Pacific Northwest Coast of North America. , they constitute two of the 231 federally recognized List of Alaska Native tribal entities, Tribes of Alaska. Most Tlingit are Alaska Natives; ...

and Haida legends, as their mascot.

References

* * *“The Wolffishes (Family : Anarhichadidae).” ''Finfish Aquaculture Diversification'', by R. Le François Nathalie, CABI, 2010, pp. 417–418. {{Taxonbar, from=Q604663 Perciformes families Articles which contain graphical timelines Taxa named by Charles Lucien Bonaparte