A sea eagle or fish eagle (also called erne or ern, mostly in reference to the white-tailed eagle) is any of the
birds of prey
Birds of prey or predatory birds, also known as (although not the same as) raptors, are hypercarnivorous bird species that actively predation, hunt and feed on other vertebrates (mainly mammals, reptiles and smaller birds). In addition to speed ...
in the
subfamily
In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end botanical subfamily names with "-oideae", and zo ...
Haliaeetinae of the bird of prey family
Accipitridae
The Accipitridae () is one of the four families within the order Accipitriformes, and is a family of small to large birds of prey with strongly hooked bills and variable morphology based on diet. They feed on a range of prey items from insects ...
. Ten extant species exist, currently described with this label.
The subfamily has a significant reach, with a scholarly article in 2005 reporting that they were "found in riverine and coastal habitat
throughout the world". However, ''Haliaeetinae'' inhabited areas have experienced particular threats given the context of
human impacts on the environment
Human impact on the environment (or anthropogenic environmental impact) refers to changes to biophysical environments and to ecosystems, biodiversity, and natural resources caused directly or indirectly by humans. Modifying the environment to ...
.
Taxonomy and evolution
The genus ''Haliaeetus'' was introduced in 1809 by French naturalist
Marie Jules César Savigny in his chapter on birds in the ''
Description de l'Égypte''. The two fish eagles in the genus ''Ichthyophaga'' were found to lie within ''Haliaeetus'' in a genetic study in 2005.
They were then moved accordingly. They are very similar to the tropical ''Haliaeetus'' species. A prehistoric (i.e. extinct before 1500) form from Maui in the Hawaiian Islands may represent a species or subspecies in this genus.
The relationships to other genera in the family are less clear; they have long been considered closer to the genus ''
Milvus'' (kites) than to the true eagles in the genus ''
Aquila'' on the basis of their morphology and display behaviour;
[Brown, L. H, & Amadon, D. (1968). ''Eagles, Hawks and Falcons of the World''. Country Life Books, Feltham.] more recent genetic evidence agrees with this, but points to their being related to the genus ''
Buteo'' (buzzards/hawks), as well, a relationship not previously thought close.
A 2005 molecular study found that the genus is
paraphyletic
Paraphyly is a taxonomic term describing a grouping that consists of the grouping's last common ancestor and some but not all of its descendant lineages. The grouping is said to be paraphyletic ''with respect to'' the excluded subgroups. In co ...
and subsumes ''
Ichthyophaga'', the species diverging into a temperate and tropical group.
Evolution
''Haliaeetus'' is possibly one of the oldest genera of living birds. A
distal
Standard anatomical terms of location are used to describe unambiguously the anatomy of humans and other animals. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position provi ...
left
tarsometatarsus
The tarsometatarsus is a bone that is only found in the lower leg of birds and some non-avian dinosaurs. It is formed from the fusion of several bird bones found in other types of animals, and homologous to the mammalian tarsus (ankle bones) a ...
(DPC 1652) recovered from early
Oligocene
The Oligocene ( ) is a geologic epoch (geology), epoch of the Paleogene Geologic time scale, Period that extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that defin ...
deposits of Fayyum,
Egypt
Egypt ( , ), officially the Arab Republic of Egypt, is a country spanning the Northeast Africa, northeast corner of Africa and Western Asia, southwest corner of Asia via the Sinai Peninsula. It is bordered by the Mediterranean Sea to northe ...
(
Jebel Qatrani Formation
The Jebel Qatrani Formation (also Gebel Qatrani, Gabal Qatrani or Djebel Qatrani) is a geologic formation located in the Faiyum Governorate of central Egypt. It is exposed between the Jebel Qatrani escarpment and the Qasr el Sagha escarpment, nor ...
, about 33 million years ago (Mya)) is similar in general pattern and some details to that of a modern sea eagle.
[Rasmussen, D., Tab, O., Storrs, L., & Simons, E. L. (1987). Fossil Birds from the Oligocene Jebel Qatrani Formation, Fayum Province, Egypt. ''Smithsonian Contributions to Paleobiology'' 62: 1–20]
PDF Fulltext
(file size 8.1 MB) The genus was present in the middle
Miocene
The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...
(12–16 Mya) with certainty.
[ Lambrecht, K. (1933). ''Handbuch der Palaeornithologie''. Gebrüder Bornträger, Berlin.]
The origin of the sea eagles and fishing eagles is probably in the general area of the
Bay of Bengal
The Bay of Bengal is the northeastern part of the Indian Ocean. Geographically it is positioned between the Indian subcontinent and the Mainland Southeast Asia, Indochinese peninsula, located below the Bengal region.
Many South Asian and Southe ...
. During the
Eocene
The Eocene ( ) is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (Ma). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes ...
/
Oligocene
The Oligocene ( ) is a geologic epoch (geology), epoch of the Paleogene Geologic time scale, Period that extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that defin ...
, as the
Indian subcontinent
The Indian subcontinent is a physiographic region of Asia below the Himalayas which projects into the Indian Ocean between the Bay of Bengal to the east and the Arabian Sea to the west. It is now divided between Bangladesh, India, and Pakista ...
slowly collided with
Eurasia
Eurasia ( , ) is a continental area on Earth, comprising all of Europe and Asia. According to some geographers, Physical geography, physiographically, Eurasia is a single supercontinent. The concept of Europe and Asia as distinct continents d ...
, this was a vast expanse of fairly shallow ocean; the initial sea eagle divergence seems to have resulted in the four
tropical
The tropics are the regions of Earth surrounding the equator, where the sun may shine directly overhead. This contrasts with the temperate or polar regions of Earth, where the Sun can never be directly overhead. This is because of Earth's ax ...
(and
Southern Hemisphere subtropical
The subtropical zones or subtropics are geographical zone, geographical and Köppen climate classification, climate zones immediately to the Northern Hemisphere, north and Southern Hemisphere, south of the tropics. Geographically part of the Ge ...
) species living around the
Indian Ocean
The Indian Ocean is the third-largest of the world's five oceanic divisions, covering or approximately 20% of the water area of Earth#Surface, Earth's surface. It is bounded by Asia to the north, Africa to the west and Australia (continent), ...
today. The
Central Asia
Central Asia is a region of Asia consisting of Kazakhstan, Kyrgyzstan, Tajikistan, Turkmenistan, and Uzbekistan. The countries as a group are also colloquially referred to as the "-stans" as all have names ending with the Persian language, Pers ...
n Pallas's sea eagle's relationships to the other
taxa
In biology, a taxon (back-formation from ''taxonomy''; : taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular name and ...
is more obscure; it seems closer to the three
Holarctic
The Holarctic realm is a biogeographic realm that comprises the majority of habitats found throughout the continents in the Northern Hemisphere. It corresponds to the floristic Boreal Kingdom. It includes both the Nearctic zoogeographical reg ...
species which evolved later and may be an early offshoot of this northward expansion; it does not have the hefty yellow bill of the northern forms, retaining a smaller, darker beak like the tropical species.
The rate of
molecular evolution
Molecular evolution describes how Heredity, inherited DNA and/or RNA change over evolutionary time, and the consequences of this for proteins and other components of Cell (biology), cells and organisms. Molecular evolution is the basis of phylogen ...
in ''Haliaeetus'' is fairly slow, as is to be expected in long-lived birds which take years to successfully reproduce. In the
mtDNA
Mitochondrial DNA (mtDNA and mDNA) is the DNA located in the mitochondria organelles in a eukaryotic cell that converts chemical energy from food into adenosine triphosphate (ATP). Mitochondrial DNA is a small portion of the DNA contained in ...
cytochrome ''b'' gene, a
mutation
In biology, a mutation is an alteration in the nucleic acid sequence of the genome of an organism, virus, or extrachromosomal DNA. Viral genomes contain either DNA or RNA. Mutations result from errors during DNA or viral replication, ...
rate of 0.5–0.7% per million years (if assuming an Early Miocene divergence) or maybe as little as 0.25–0.3% per million years (for a Late Eocene divergence) has been shown.
Issues in the modern era
The ''Haliaeetinae'' subfamily is an especially threatened collection of creatures within the broader ''Accipitridae'' species, according to the academic journal ''
Molecular Phylogenetics and Evolution
''Molecular Phylogenetics and Evolution'' is a peer-reviewed scientific journal of evolutionary biology and phylogenetics. The journal is edited by E.A. Zimmer.
Indexing
The journal is indexed in:
* EMBiology
*Journal Citation Reports
*Scopus ...
'', given the "
anthropogenic factors" involved. The publication reported in 2005 that prior trends had meant that sea eagles could be "found in riverine and coastal habitat
throughout the world". In terms of international scientific campaigns, the
Convention on International Trade in Endangered Species
Convention may refer to:
* Convention (norm), a custom or tradition, a standard of presentation or conduct
** Treaty, an agreement in international law
** Convention (political norm), uncodified legal or political tradition
* Convention (meeting ...
(CITES) protects all entities in the broader species, including sea eagles.
Species
Current sea eagles
Description
Sea eagles vary in size, from
Sanford's sea eagle, averaging , to
Steller's sea eagle, weighing up to . At up to , the
white-tailed eagle is the largest eagle in Europe.
Bald eagle
The bald eagle (''Haliaeetus leucocephalus'') is a bird of prey found in North America. A sea eagle, it has two known subspecies and forms a species pair with the white-tailed eagle (''Haliaeetus albicilla''), which occupies the same niche ...
s can weigh up to , making them the largest eagle native to North America. There are exceptional records of even heavier individuals in both the white-tailed and bald eagles, although not surpassing the largest Steller's sea eagles. The
white-bellied sea eagle can weigh up to . They are generally overall brown (from rich brown to dull grey-brown), often with white to the head, tail or underparts. Some of the species have an all-yellow beak as adults, which is unusual among eagles.
Their diets consist mainly of fish, aquatic birds, and small mammals. Nests are typically very large and positioned in a tree, but sometimes on a cliff.
The tail is entirely white in adult ''Haliaeetus'' species except for Sanford's, white-bellied, and Pallas's. Three
species pairs exist: white-tailed and bald eagles, Sanford's and white-bellied sea eagles, and the African and Madagascar fish eagles, each of these consists of a white- and a tan-headed species.
In popular culture

* The
bald eagle
The bald eagle (''Haliaeetus leucocephalus'') is a bird of prey found in North America. A sea eagle, it has two known subspecies and forms a species pair with the white-tailed eagle (''Haliaeetus albicilla''), which occupies the same niche ...
is the
national symbol
A national symbol is a manifestation of a nation or community, serving as a representation of their National identity, identity and values. National symbols may be not only applied to sovereign states but also nations and countries in a state of ...
of the
United States
The United States of America (USA), also known as the United States (U.S.) or America, is a country primarily located in North America. It is a federal republic of 50 U.S. state, states and a federal capital district, Washington, D.C. The 48 ...
.
* The silver eagle on red shield on the arms of Poland has been interpreted as the sea eagle.
*
Namibia
Namibia, officially the Republic of Namibia, is a country on the west coast of Southern Africa. Its borders include the Atlantic Ocean to the west, Angola and Zambia to the north, Botswana to the east and South Africa to the south; in the no ...
,
Zambia
Zambia, officially the Republic of Zambia, is a landlocked country at the crossroads of Central Africa, Central, Southern Africa, Southern and East Africa. It is typically referred to being in South-Central Africa or Southern Africa. It is bor ...
, and
Zimbabwe
file:Zimbabwe, relief map.jpg, upright=1.22, Zimbabwe, relief map
Zimbabwe, officially the Republic of Zimbabwe, is a landlocked country in Southeast Africa, between the Zambezi and Limpopo Rivers, bordered by South Africa to the south, Bots ...
have the
African fish eagle
The African fish eagle (''Icthyophaga vocifer'') or the African sea eagle is a large species of eagle found throughout sub-Saharan Africa wherever large bodies of open water with an abundant food supply occur. It is the national bird of Malawi, ...
as their
national bird.
* The
white-tailed eagle is the
national bird of
Poland
Poland, officially the Republic of Poland, is a country in Central Europe. It extends from the Baltic Sea in the north to the Sudetes and Carpathian Mountains in the south, bordered by Lithuania and Russia to the northeast, Belarus and Ukrai ...
.
* The
Manly Warringah Sea Eagles are an Australian professional
rugby league
Rugby league football, commonly known as rugby league in English-speaking countries and rugby 13/XIII in non-Anglophone Europe, is a contact sport, full-contact sport played by two teams of thirteen players on a rectangular Rugby league playin ...
club that competes in the
National Rugby League
The National Rugby League (also known as the NRL Telstra Premiership for sponsorship reasons) is a professional rugby league competition in Oceania which contains clubs from New South Wales, Queensland, Victoria (state), Victoria, the Austral ...
(NRL).
* Nesting pairs of both the bald eagle and white-bellied sea eagle have been subject to live-streaming webcam footage.
* In
heraldic language, the
osprey
The osprey (; ''Pandion haliaetus''), historically known as sea hawk, river hawk, and fish hawk, is a diurnal, fish-eating bird of prey with a cosmopolitan range. It is a large raptor, reaching more than in length and a wingspan of . It ...
is termed a "sea-eagle", although ospreys come from the taxonomic family ''
Pandionidae'' and are not classified as true sea eagles.
* Novelist
Kurt Vonnegut
Kurt Vonnegut ( ; November 11, 1922 – April 11, 2007) was an American author known for his Satire, satirical and darkly humorous novels. His published work includes fourteen novels, three short-story collections, five plays, and five nonfict ...
invented the extinct “Bermuda ern” as a literary device in his novel
Breakfast of Champions.
See also
*
Brahminy kite, also called "red-backed sea eagle"
*
Osprey
The osprey (; ''Pandion haliaetus''), historically known as sea hawk, river hawk, and fish hawk, is a diurnal, fish-eating bird of prey with a cosmopolitan range. It is a large raptor, reaching more than in length and a wingspan of . It ...
, also called "sea hawk"
References
General sources
*
*
{{Authority control
Eagles