Scytinium Biatorinum
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''Scytinium'' is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
lichen A lichen ( , ) is a hybrid colony (biology), colony of algae or cyanobacteria living symbiotically among hypha, filaments of multiple fungus species, along with yeasts and bacteria embedded in the cortex or "skin", in a mutualism (biology), m ...
-forming
fungi A fungus (: fungi , , , or ; or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and mold (fungus), molds, as well as the more familiar mushrooms. These organisms are classified as one ...
in the family
Collemataceae The Collemataceae are a family of mostly lichen-forming fungi in the order Peltigerales. The family contains twelve genera and about 325 species. The family has a widespread distribution. Taxonomy The family was circumscribed by Jonathan Carl Z ...
. It has 49 species. These lichens are typically found on basic rocks, soil, and trees, occasionally in association with
moss Mosses are small, non-vascular plant, non-vascular flowerless plants in the taxonomic phylum, division Bryophyta (, ) ''sensu stricto''. Bryophyta (''sensu lato'', Wilhelm Philippe Schimper, Schimp. 1879) may also refer to the parent group bryo ...
es. Despite the morphological and ecological diversity within ''Scytinium'', its species share similar
ascospore In fungi, an ascospore is the sexual spore formed inside an ascus—the sac-like cell that defines the division Ascomycota, the largest and most diverse Division (botany), division of fungi. After two parental cell nucleus, nuclei fuse, the ascu ...
features, such as shape and septation, as well as a small to medium-sized
thallus Thallus (: thalli), from Latinized Greek (), meaning "a green shoot" or "twig", is the vegetative tissue of some organisms in diverse groups such as algae, fungi, some liverworts, lichens, and the Myxogastria. A thallus usually names the entir ...
with at least a partial .


Description

''Scytinium'' encompasses lichen species that exhibit a variety of thallus forms, such as
crustose Crustose is a Habit (biology), habit of some types of algae and lichens in which the organism grows tightly appressed to a substrate, forming a biological layer. ''Crustose'' adheres very closely to the Substrate (biology), substrates at all poin ...
,
squamulose A squamulose lichen is a lichen that is composed of small, often overlapping "scales" called . If they are raised from the substrate and appear leafy, the lichen may appear to be a foliose lichen, but the underside does not have a "skin" (cortex) ...
, , or . These lichens have a gelatinous texture, and their colour ranges from dark brown and bluish-grey to olive-green. The of ''Scytinium'' can be spreading, elongate, or somewhat cylindrical in shape. The upper and lower , when present, is composed of either cuboid cells or flattened, degraded tissue. The medulla contains loosely interwoven or compact
hypha A hypha (; ) is a long, branching, filamentous structure of a fungus, oomycete, or actinobacterium. In most fungi, hyphae are the main mode of vegetative growth, and are collectively called a mycelium. Structure A hypha consists of one o ...
e, along with the photobiont ''
Nostoc ''Nostoc'', also known as star jelly, troll's butter, spit of moon, fallen star, witch's butter (not to be confused with the fungi commonly known as witches' butter), and witch's jelly, is the most common genus of cyanobacteria found in a variety ...
''–a common genus of
cyanobacteria Cyanobacteria ( ) are a group of autotrophic gram-negative bacteria that can obtain biological energy via oxygenic photosynthesis. The name "cyanobacteria" () refers to their bluish green (cyan) color, which forms the basis of cyanobacteri ...
. Both the upper and lower surfaces of ''Scytinium'' lichens can vary from smooth to wrinkled or ridged and typically exhibit a matte appearance. While
isidia An isidium (plural: isidia) is a tiny, wart- or finger-like outgrowth on the thallus surface of certain lichen species. It is one of two principal types of vegetative reproduction, vegetative reproductive structures in lichens, the other being ...
may or may not be present,
soredia Soredia are common reproduction, reproductive structures of lichens. Lichens asexual reproduction, reproduce asexually by employing simple fragmentation and production of soredia and isidia. Soredia are powdery propagules composed of fungus, fung ...
are absent in this genus. The cells are generally arranged in distinct chains. The
ascomata An ascocarp, or ascoma (: ascomata), is the fruiting body (sporocarp (fungi), sporocarp) of an ascomycete phylum fungus. It consists of very tightly interwoven hyphae and millions of embedded ascus, asci, each of which typically contains four to ...
are in the form of
apothecia An ascocarp, or ascoma (: ascomata), is the fruiting body ( sporocarp) of an ascomycete phylum fungus. It consists of very tightly interwoven hyphae and millions of embedded asci, each of which typically contains four to eight ascospores. As ...
with a red-brown , which can be sessile, laminal, or marginal. The is smooth to , to , and often persistent. The is raised, cup-shaped, and predominantly composed of cells, ranging in colour from colourless to reddish-brown. The disc is either concave or flat. The is colourless to reddish-brown and does not react with a solution of
potassium hydroxide Potassium hydroxide is an inorganic compound with the formula K OH, and is commonly called caustic potash. Along with sodium hydroxide (NaOH), KOH is a prototypical strong base. It has many industrial and niche applications, most of which utili ...
or ammonia. The
hymenium The hymenium is the tissue layer on the hymenophore of a fungal fruiting body where the cells develop into basidia or asci, which produce spores. In some species all of the cells of the hymenium develop into basidia or asci, while in oth ...
is colourless, turning blue when exposed to
iodine Iodine is a chemical element; it has symbol I and atomic number 53. The heaviest of the stable halogens, it exists at standard conditions as a semi-lustrous, non-metallic solid that melts to form a deep violet liquid at , and boils to a vi ...
. The is shallow, and its colour varies from colourless to pale yellowish. The consists of numerous, that separate in potassium hydroxide and sometimes have branched apices that are somewhat swollen. The asci contain eight spores and are club-shaped (clavate), with a strongly thickened apex that reacts to potassium hydroxide and iodine, turning blue. are primarily
ellipsoid An ellipsoid is a surface that can be obtained from a sphere by deforming it by means of directional Scaling (geometry), scalings, or more generally, of an affine transformation. An ellipsoid is a quadric surface;  that is, a Surface (mathemat ...
al, , and colourless, lacking any distinct surface ornamentation or . , or , are infrequent in ''Scytinium'' species. No
lichen product Lichen products, also known as lichen substances, are organic compounds produced by a lichen. Specifically, they are secondary metabolites. Lichen products are represented in several different chemical classes, including terpenoids, orcinol deri ...
s have been detected in ''Scytinium'' species using
thin-layer chromatography Thin-layer chromatography (TLC) is a chromatography technique that separates components in non-volatile mixtures. It is performed on a TLC plate made up of a non-reactive solid coated with a thin layer of adsorbent material. This is called the sta ...
.


Species

,
Species Fungorum ''Index Fungorum'' is an international project to index all formal names (Binomial nomenclature, scientific names) in the fungus Kingdom (biology), kingdom. As of 2015, the project is based at the Royal Botanic Gardens, Kew, one of three partn ...
(in the
Catalogue of Life The Catalogue of Life (CoL) is an online database that provides an index of known species of animals, plants, fungi, and microorganisms. It was created in 2001 as a partnership between the global Species 2000 and the American Integrated Taxono ...
) accepts 48 species of ''Scytinium''. *'' Scytinium apalachense'' *''
Scytinium aquale ''Scytinium'' is a genus of lichen-forming fungi in the family Collemataceae. It has 49 species. These lichens are typically found on basic rocks, soil, and trees, occasionally in association with mosses. Despite the morphological and ecologica ...
'' *'' Scytinium aragonii'' *'' Scytinium biatorinum'' *'' Scytinium bosoense'' *'' Scytinium californicum'' *'' Scytinium callopismum'' *'' Scytinium cellulosum'' *'' Scytinium chibaense'' *'' Scytinium contortum'' *'' Scytinium dactylinum'' *'' Scytinium erectum'' *'' Scytinium euthallinum'' *'' Scytinium ferax'' *'' Scytinium fragile'' *'' Scytinium fragrans'' *''
Scytinium gelatinosum ''Scytinium'' is a genus of lichen-forming fungi in the family Collemataceae. It has 49 species. These lichens are typically found on basic rocks, soil, and trees, occasionally in association with mosses. Despite the morphological and ecologica ...
'' *'' Scytinium imbricatum'' *'' Scytinium intermedium'' *'' Scytinium juniperinum'' *'' Scytinium kauaiense'' *'' Scytinium leptogioides'' *'' Scytinium lichenoides'' *'' Scytinium magnussonii'' *'' Scytinium massiliense'' *'' Scytinium nanum'' *'' Scytinium palmatum'' *'' Scytinium palustre'' *'' Scytinium parculum'' *'' Scytinium parvum'' *'' Scytinium platynum'' *'' Scytinium plicatile'' *'' Scytinium polycarpum'' *'' Scytinium pulvinatum'' *'' Scytinium quadrifidum'' *'' Scytinium rivale'' *'' Scytinium rogersii'' *'' Scytinium schraderi'' *'' Scytinium singulare'' *'' Scytinium siskiyouense'' *'' Scytinium subaridum'' *'' Scytinium subfragrans'' *'' Scytinium subtile'' *'' Scytinium subtorulosum'' *'' Scytinium tacomae'' *'' Scytinium tenuilobum'' – Australia *'' Scytinium tenuissimum'' *'' Scytinium tetrasporum'' *'' Scytinium turgidum''


References

{{Taxonbar, from=Q23072885 Peltigerales Taxa described in 1810 Taxa named by Erik Acharius Peltigerales genera Lichen genera