Scolopendra Michoacana
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''Scolopendra'', from
Ancient Greek Ancient Greek (, ; ) includes the forms of the Greek language used in ancient Greece and the classical antiquity, ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Greek ...
σκόλοψ (''skólops''), meaning "thorn", and ἔντερον (''énteron''), meaning "earthworm", is a species-rich genus of large tropical centipedes of the family Scolopendridae.


Description

The genus ''Scolopendra'' contains many species of centipedes found across the world's tropics and warmer temperate areas. The species vary considerably in coloration and size. ''Scolopendra'' are mostly very large centipedes. The largest species found in tropical climates can exceed and are the largest living centipedes in the world. All ''Scolopendra'' species can deliver a painful bite, injecting
venom Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a sti ...
through their forcipules, which are not fangs or other mouthparts; rather, these are modified legs on the first body segment.


Ecology

''Scolopendra'' species are active predators, feeding primarily on insects and other invertebrates. Larger specimens have been observed preying on frogs, tarantulas, lizards, birds,
snakes Snakes are elongated Limbless vertebrate, limbless reptiles of the suborder Serpentes (). Cladistically Squamata, squamates, snakes are ectothermic, amniote vertebrates covered in overlapping Scale (zoology), scales much like other members of ...
,
rodent Rodents (from Latin , 'to gnaw') are mammals of the Order (biology), order Rodentia ( ), which are characterized by a single pair of continuously growing incisors in each of the upper and Mandible, lower jaws. About 40% of all mammal specie ...
s, and even bats. Two southeast Asian species, '' S. cataracta'' and '' S. paradoxa'', as well as '' S. alcyona'' from the Ryukyu Islands, are amphibious, as these species can travel underwater by swimming or walking.


Venom

The venom of most species is not medically significant; however, bites from several species can cause intense and long-lasting pain and swelling. Large ''Scolopendra'' species from Asian/Pacific regions, such as '' Scolopendra subspinipes'' and '' Scolopendra dehaani'', are particularly potent, and have caused one reported fatality. In 2014, a fatality was reported for a bite from a '' Scolopendra gigantea''. The venom of certain ''Scolopendra'' species were found to contain compounds such as serotonin, haemolytic phospholipase, a cardiotoxic
protein Proteins are large biomolecules and macromolecules that comprise one or more long chains of amino acid residue (biochemistry), residues. Proteins perform a vast array of functions within organisms, including Enzyme catalysis, catalysing metab ...
, and a cytolysin.


Taxonomic history

''Scolopendra'' was one of the genera created by
Carl Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
in his 1758 10th edition of ''Systema Naturae'', the starting point for zoological nomenclature. Only two of the species originally assigned to the genus remain so: '' Scolopendra gigantea'' and '' S. morsitans''; the latter was chosen to be the
type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
by Opinion 454 of the International Commission on Zoological Nomenclature, overruling a previous designation by
Pierre André Latreille Pierre André Latreille (; 29 November 1762 – 6 February 1833) was a French zoology, zoologist, specialising in arthropods. Having trained as a Roman Catholic priest before the French Revolution, Latreille was imprisoned, and only regained hi ...
, in which he chose Linnaeus' ''Scolopendra forficata'' (now '' Lithobius forficatus'') as the type species.


Species

The genus ''Scolopendra'' contains these species: *'' Scolopendra abnormis'' Lewis & Daszak, 1996 *'' Scolopendra afer'' (Meinert, 1886) *'' Scolopendra alcyona'' Tsukamoto & Shimano, 2021 *'' Scolopendra algerina'' Newport, 1845 *'' Scolopendra alternans'' Leach, 1813 *'' Scolopendra andhrensis'' Jangi & Dass, 1984 *'' Scolopendra angulata'' Newport, 1844 *'' Scolopendra angusticollis'' Murray, 1887 *'' Scolopendra anomia'' Newport, 1844 *'' Scolopendra appendiculata'' Daday, 1891 *'' Scolopendra arenicola'' (Lawrence, 1975) *'' Scolopendra arthrorhabdoides'' Ribaut, 1912 *'' Scolopendra attemsi'' Lewis, Minelli & Shelley, 2006 *'' Scolopendra aztecorum'' Verhoeff, 1934 *'' Scolopendra calcarata'' Porat, 1876 *'' Scolopendra canidens'' Newport, 1844 * '' Scolopendra cataracta'' Siriwut, Edgecombe & Panha, 2016 *'' Scolopendra childreni'' Newport, 1844 *'' Scolopendra chlora'' Chamberlin, 1942 *'' Scolopendra chlorotes'' C. L. Koch, 1856 *'' Scolopendra cingulata'' Latreille, 1829 *'' Scolopendra clavipes'' C. L. Koch, 1847 *'' Scolopendra concolor'' Newport, 1845 *'' Scolopendra crassa'' Templeton, 1846 *'' Scolopendra cretica'' Attems, 1902 *'' Scolopendra cribrifera'' Gervais, 1847 *'' Scolopendra crudelis'' C. L. Koch, 1847 *'' Scolopendra dalmatica'' C. L. Koch, 1847 *'' Scolopendra dawydoffi'' Kronmüller, 2012 *'' Scolopendra dehaani'' Brandt, 1840 *'' Scolopendra ellorensis'' Jangi & Dass, 1984 *'' Scolopendra fissispina'' L. Koch, 1865 *'' Scolopendra foveolata'' Verhoeff, 1937 *'' Scolopendra galapagoensis'' Bollman, 1889 *'' Scolopendra gigantea''
Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
, 1758
*'' Scolopendra gracillima'' Attems, 1898 *'' Scolopendra hainanum'' Kronmüller, 2012 *'' Scolopendra hardwickei'' Newport, 1844 *'' Scolopendra hermosa'' Chamberlin, 1941 *'' Scolopendra heros'' Girard, 1853 *'' Scolopendra horrida'' C. L. Koch, 1847 *'' Scolopendra inaequidens'' Gervais, 1847 *'' Scolopendra indiae'' (Chamberlin, 1914) *'' Scolopendra indica'' Meinert, 1886 *'' Scolopendra inermipes'' C. L. Koch, 1847 *'' Scolopendra inermis'' Newport, 1845 *'' Scolopendra jangii'' Khanna & Yadav, 1997 *'' Scolopendra japonica'' Koch, 1878 *'' Scolopendra koreana'' (Verhoeff, 1934) *'' Scolopendra labiata'' C. L. Koch, 1863 *'' Scolopendra laeta'' Haase, 1887 *'' Scolopendra langi'' (Chamberlin, 1927) *'' Scolopendra latro'' Meinert, 1886 *'' Scolopendra leki'' (Waldock & Edgecombe 2012) *'' Scolopendra limicolor'' Wood, 1861 *'' Scolopendra lucasii'' Gervais, 1847 *'' Scolopendra lufengia'' S. Kang ''et al.'', 2017 *'' Scolopendra lutea'' (Attems, 1928) *'' Scolopendra madagascariensis'' Attems, 1910 *'' Scolopendra malkini'' Chamberlin, 1955 *'' Scolopendra mazbii'' Gravely, 1912 *'' Scolopendra media'' (Muralewicz, 1926) *'' Scolopendra melionii'' Lucas, 1853 *'' Scolopendra metuenda'' Pocock, 1895 *'' Scolopendra michoacana'' Chamberlin, 1941 *'' Scolopendra mima'' Chamberlin, 1942 *'' Scolopendra mirabilis'' (Porat, 1876) *'' Scolopendra monticola'' (Lawrence, 1975) *'' Scolopendra morsitans''
Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
, 1758
*'' Scolopendra multidens'' Newport, 1844 *'' Scolopendra negrocapitis'' Zhang & Wang, 1999 *'' Scolopendra nuda'' (Jangi & Dass, 1980) *'' Scolopendra occidentalis'' F. Meinert, 1886 *'' Scolopendra octodentata'' Verhoeff, 1934 *'' Scolopendra oraniensis'' Lucas, 1846 *'' Scolopendra paradoxa'' Doménech, 2018 *'' Scolopendra pachygnatha'' Pocock, 1895 *'' Scolopendra paranuda'' (Khanna & Tripathi, 1987) *'' Scolopendra pentagramma'' Motschoulsky, 1886 *'' Scolopendra pinguis'' Pocock, 1891 *'' Scolopendra polymorpha'' Wood, 1861 *'' Scolopendra pomacea'' C. L. Koch, 1847 *'' Scolopendra puncticeps'' Wood, 1861 *'' Scolopendra punensis'' Jangi & Dass, 1984 *'' Scolopendra robusta'' Kraepelin, 1903 *'' Scolopendra sanatillae'' Bollman, 1893 *'' Scolopendra silhetensis'' Newport, 1845 *'' Scolopendra spinipriva'' Bücherl, 1946 *'' Scolopendra spinosissima'' Kraepelin, 1903 *'' Scolopendra subspinipes'' Leach, 1816 *'' Scolopendra sumichrasti'' Saussure, 1860 *'' Scolopendra tenuitarsis'' Pocock, 1895 *'' Scolopendra teretipes'' (Porat,1893) *'' Scolopendra valida'' Lucas, 1840 *'' Scolopendra violacea'' Fabricius, 1798 *'' Scolopendra viridicornis'' Newport, 1844 *'' Scolopendra viridipes'' Dufour, 1820 *'' Scolopendra viridis'' Say, 1821 *'' Scolopendra zuluana'' (Lawrence, 1958) One
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
species, †'' Scolopendra proavita'', is known from Baltic amber deposits from the
Eocene The Eocene ( ) is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (Ma). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes ...
of
Poland Poland, officially the Republic of Poland, is a country in Central Europe. It extends from the Baltic Sea in the north to the Sudetes and Carpathian Mountains in the south, bordered by Lithuania and Russia to the northeast, Belarus and Ukrai ...
. Fossil remains of a species tentatively assigned to '' S. morsitans'' (as ''S. (cf)'' ''morsitans'') are also known from
Pliocene The Pliocene ( ; also Pleiocene) is the epoch (geology), epoch in the geologic time scale that extends from 5.33 to 2.58Makapansgat of South Africa.


Species gallery


See also

* Gu (poison)


References

{{Authority control Centipede genera