Sclerocactus
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''Sclerocactus'' ("hard
cactus A cactus (: cacti, cactuses, or less commonly, cactus) is a member of the plant family Cactaceae (), a family of the order Caryophyllales comprising about 127 genera with some 1,750 known species. The word ''cactus'' derives, through Latin, ...
", from
Greek Greek may refer to: Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group *Greek language, a branch of the Indo-European language family **Proto-Greek language, the assumed last common ancestor of all kno ...
; refers to the hard, dry fruit) is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
cacti A cactus (: cacti, cactuses, or less commonly, cactus) is a member of the plant family Cactaceae (), a family of the order Caryophyllales comprising about 127 genera with some 1,750 known species. The word ''cactus'' derives, through Latin, ...
. It comprises about 15 species, the exact number depending on the authority. These species are
xerophytic A xerophyte () is a species of plant that has adaptations to survive in an environment with little liquid water. Examples of xerophytes include cactus, cacti, pineapple and some gymnosperm plants. The morphology (biology), morphology and physiology ...
. They are sometimes called 'fishhook cactus' or 'little barrels.'


Description

''Sclerocactus'' range in shape from ovoid to elongate cylindric, have rigid stems with
tubercle In anatomy, a tubercle (literally 'small tuber', Latin for 'lump') is any round nodule, small eminence, or warty outgrowth found on external or internal organs of a plant or an animal. In plants A tubercle is generally a wart-like projectio ...
s that are generally coalesced into ribs, and are covered with spines that come out of the
areole In botany, areoles are small light- to dark-colored bumps on cactus, cacti out of which grow clusters of Thorns, spines, and prickles, spines. Areoles are important diagnostic features of cactus, cacti, and identify them as a family distinct fr ...
s., in Most species have at least one hooked spine at each areole. Less often, species may not have hooks. These plants are found in higher elevation deserts such as on the
Colorado Plateau The Colorado Plateau is a physiographic and desert region of the Intermontane Plateaus, roughly centered on the Four Corners region of the Southwestern United States. This plateau covers an area of 336,700 km2 (130,000 mi2) within w ...
, or in the Mohave Desert or the
Great Basin The Great Basin () is the largest area of contiguous endorheic watersheds, those with no outlets to the ocean, in North America. It spans nearly all of Nevada, much of Utah, and portions of California, Idaho, Oregon, Wyoming, and Baja Californi ...
. They are well suited to extremes due to lack of rainfall, hot summers and below freezing winters. The little barrel cactus typically grows in areas where water flows irregularly or depressions where water can accumulate for short periods of time. They are most often found growing along washes and arroyos where their seeds have been subjected to scarification due to water movement, but they oddly also tend to grow along ridges in spots where depressions have formed and can hold water for some period of time. The sometimes hooked or curved spines and the armored web of spines enclosing the cactus body in species of this genus is an adaptation which allows the plant to move to more favorable locations. This plants seeds germinate in areas where water movement occurs or in areas where standing water accumulates for some period of time, and during flash floods, the hooked spines allow the plants to be caught on water borne debris and be uprooted and carried to areas where water tends to accumulate. A large percentage of plants in habitat show signs of water debris damage along the stems which has healed and are found growing on their sides in places where floodwaters have deposited them. The habitat these plants exist in is very arid, and the plants have adapted to exploit water movement to concentrate their biomass in areas where water is likely to be present.


Synonymy

''Sclerocactus'' was once reduced to
synonymy A synonym is a word, morpheme, or phrase that means precisely or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are a ...
with '' Pediocactus''—but this is not accepted; these genera are not closely related. The following genera have been brought into synonymy with ''Sclerocactus'': *''Ancistrocactus'' Britton &
Rose A rose is either a woody perennial plant, perennial flowering plant of the genus ''Rosa'' (), in the family Rosaceae (), or the flower it bears. There are over three hundred Rose species, species and Garden roses, tens of thousands of cultivar ...
*''Coloradoa'' Boissev. & C.Davidson *''Echinomastus'' Britton &
Rose A rose is either a woody perennial plant, perennial flowering plant of the genus ''Rosa'' (), in the family Rosaceae (), or the flower it bears. There are over three hundred Rose species, species and Garden roses, tens of thousands of cultivar ...
*''Toumeya'' Britton &
Rose A rose is either a woody perennial plant, perennial flowering plant of the genus ''Rosa'' (), in the family Rosaceae (), or the flower it bears. There are over three hundred Rose species, species and Garden roses, tens of thousands of cultivar ...


Species

the following species are recognized in the genus ''Sclerocactus '' by
Plants of the World Online Plants of the World Online (POWO) is an online taxonomic database published by the Royal Botanic Gardens, Kew. History Following the Convention on Biological Diversity, the Royal Botanic Gardens in Kew launched Plants of the World Online i ...
: Many species of ''Sclerocactus'' are protected under the provisions of the
Endangered Species Act The Endangered Species Act of 1973 (ESA; 16 U.S.C. § 1531 et seq.) is the primary law in the United States for protecting and conserving imperiled species. Designed to protect critically imperiled species from extinction as a "consequence of e ...
and all species should be treated as imperiled.


Cultivation and propagation

''Sclerocactus'' are easy to cultivate. They are generally a cold-adapted genus. They grow in very arid areas which are subjected to deep cold in spots where water flows irregularly or depressions where water can accumulate for short period of time. They are most often found growing along washes and arroyos where their seeds have been subjected to
scarification Scarification involves scratching, etching, burning/ branding, or superficially cutting designs, pictures, or words into the skin as a permanent body modification or body art. The body modification can take roughly 6–12 months to heal. In t ...
due to water movement, but they oddly also tend to grow along ridges in spots where depressions have formed and can hold water for some period of time. They undergo most of their growth in spring and benefit from moderate
fertilizer A fertilizer or fertiliser is any material of natural or synthetic origin that is applied to soil or to plant tissues to supply plant nutrients. Fertilizers may be distinct from liming materials or other non-nutrient soil amendments. Man ...
in cultivation. They prefer mineral-rich, sandy soils. They are tolerant of overwatering provided they are grown in small pots that drain and dry out quickly. They do best out of doors do well in full sun. They are not particularly attractive plants and their vicious hooked spines are a hazard, making them poor houseplants. Like '' Pediocactus'', ''Sclerocactus'' "deflate" in late fall in preparation for winter and freezing; they push water out of the plant flesh, through the roots, and into the soil, dramatically decreasing in size. They require deep cold and
dormancy Dormancy is a period in an organism's Biological life cycle, life cycle when growth, development, and (in animals) physical activity are temporarily stopped. This minimizes metabolism, metabolic activity and therefore helps an organism to conserv ...
to bloom well. Some species, such as '' S. parviflorus'', produce large, showy flowers and put a short, but impressive floral display in early spring. ''Sclerocactus'' seeds are difficult to germinate as they contain inhibitors, and require cold stratification, prolonged soaking, or scarification of the seeds to trigger
germination Germination is the process by which an organism grows from a seed or spore. The term is applied to the sprouting of a seedling from a seed of an angiosperm or gymnosperm, the growth of a sporeling from a spore, such as the spores of fungi, ...
. Seeds of members of this genus can take up to three years to germinate in some cases if not scarified or subjected to repeated freeze-thaw cycles in the presence of water. The plants are easy to maintain in cultivation but difficult to start from seed.


References


External links


Southwest Colorado Wildflowers
* {{Authority control Cacti of Mexico Cacti of the United States Flora of the California desert regions Flora of the Great Basin Natural history of the Mojave Desert North American desert flora Cactoideae genera