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''Scirpus'' is a genus of grass-like species in the sedge family
Cyperaceae The Cyperaceae are a family of graminoid (grass-like), monocotyledonous flowering plants known as sedges. The family is large, with some 5,500 known species described in about 90 genera, the largest being the "true sedges" genus '' Carex'' ...
many with the common names club-rush, wood club-rush or bulrush (see also bulrush for other plant genera so-named). They mostly inhabit wetlands and damp locations.


Taxonomy

The taxonomy of the genus is complex, and under review by botanists. Recent studies by taxonomists of the Cyperaceae have resulted in the creation of several new genera, including the genera ''
Schoenoplectus ''Schoenoplectus'' (club-rush ld World species bulrush or tule ew World species is a genus of plants in the sedges with a cosmopolitan distribution. Note that the name bulrush is also applied to species in the unrelated genus ''Typha'' as we ...
'' and ''
Bolboschoenus ''Bolboschoenus'' is a genus of plants in the sedge family, of nearly cosmopolitan distribution. Epipaleolithic and Neolithic peoples used ground root tubers of these plants to make the first breads. ; Accepted species * ''Bolboschoenus caldw ...
''; others (including ''
Blysmus ''Blysmus'' is a genus of sedges of the family Cyperaceae, found in temperate regions across the Northern Hemisphere. Species Species currently accepted by The Plant List are as follows: *''Blysmus compressus'' (L.) Panz. ex Link *''Blysmus mon ...
'', ''
Isolepis ''Isolepis'' is a cosmopolitan genus of sedge containing around 70 species. ''Isolepis'' is found in cool tropical and temperate climates often in Africa and Australasia. ''Isolepis'' was first described by prolific botanist Robert Brown in ...
'', '' Nomochloa'', and ''
Scirpoides ''Scirpoides'' is a genus of sedges (Cyperaceae The Cyperaceae are a family of graminoid (grass-like), monocotyledonous flowering plants known as sedges. The family is large, with some 5,500 known species described in about 90 genera, t ...
'') have also been used. At one point this genus held almost 300 species, but many of the species once assigned to it have now been reassigned, and it now holds an estimated 120 species.


Description

''Scirpus'' are rhizomatous perennial herbs, with 3-angled stems and flat grass-like leaves. The flowers are in clusters of small spikelets, often brown or greenish brown. Some species (e.g. ''S. lacustris'') can reach a height of 3 m, while ''S. sylvaticus'' is about 1.2 m and others, such as ''S. supinus'', are much smaller, only reaching 20–30 cm tall.


Distribution

The genus has a nearly cosmopolitan distribution, found on every continent except
Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
and
Antarctica Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean, it contains the geographic South Pole. Antarctica is the fifth-largest co ...
.


Ecology

Many species are common in
wetlands A wetland is a distinct ecosystem that is flooded or saturated by water, either permanently (for years or decades) or seasonally (for weeks or months). Flooding results in oxygen-free ( anoxic) processes prevailing, especially in the soils. The ...
and can produce dense stands of vegetation, along rivers, in coastal deltas and in ponds and potholes. Although flooding is the most important factor affecting its distribution, drought, ice scour, grazing, fire and salinity also affect its abundance. It can survive unfavourable conditions like prolonged flooding, or drought, as buried seeds ''Scirpus'' species are used as
food plants A crop is a plant that can be grown and harvested extensively for profit or subsistence. When the plants of the same kind are cultivated at one place on a large scale, it is called a crop. Most crops are cultivated in agriculture or hydroponics ...
by the
larva A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. Th ...
e of some Lepidoptera species, including '' Chedra microstigma'' and '' Scirpophaga nivella''.


Selected species

:''(This list is incomplete, and may include some species now assigned to other genera.)'' *'' Scirpus ancistrochaetus'' northeastern bulrush *'' Scirpus atrocinctus'' black-girdle bulrush *'' Scirpus atrovirens'' woolgrass bulrush *'' Scirpus bicolor'' *'' Scirpus campestris'' salt marsh bulrush *'' Scirpus cespitosus'' deergrass, synonymous with Trichophorum cespitosum *'' Scirpus congdonii'' Congdon's bulrush *'' Scirpus cyperinus'' - woolgrass/cottongrass bulrush *'' Scirpus diffusus'' *'' Scirpus divaricatus'' spreading bulrush *'' Scirpus expansus'' Woodland beakrush *'' Scirpus flaccidifolius'' reclining bulrush *'' Scirpus fluitans'' floating club-rush *'' Scirpus fluviatilis'' - river bulrush *'' Scirpus georgianus'' Georgia bulrush *'' Scirpus grossus'' Greater club-rush, Giant bulrush *'' Scirpus hattorianus'' - mosquito bulrush *'' Scirpus lineatus'' drooping bulrush *'' Scirpus longii'' Long's bulrush *'' Scirpus mariqueter'' *'' Scirpus microcarpus'' small-fruit bulrush *'' Scirpus mucronatus'' *''
Scirpus nevadensis ''Amphiscirpus'' is a monotypic genus of flowering plants in the sedge family containing the single species ''Amphiscirpus nevadensis'', which is known by the common name Nevada bulrush. This plant was formerly included in genus '' Scirpus''.Goe ...
'' Nevada bulrush *'' Scirpus olneyi'' Olney bulrush, synonymous with '' Schoenoplectus americanus'' *'' Scirpus pacificus'' Pacific Coast bulrush *'' Scirpus pallidus'' pale bulrush *'' Scirpus paludosus'' salt marsh bulrush *'' Scirpus pedicellatus'' stalked bulrush *'' Scirpus pendulus'' pendulous bulrush *'' Scirpus polyphyllus'' leafy bulrush *'' Scirpus pumilus'' dwarf deergrass *''
Scirpus pungens ''Schoenoplectus pungens'' is a species of flowering plant in the sedge family known as common threesquare, common three-square bulrush and sharp club-rush. It is a herbaceous emergent plant that is widespread across much of North and South Am ...
'' sharp club-rush *'' Scirpus radicans'' *'' Scirpus robustus'' salt marsh bulrush *'' Scirpus supinus'' dwarf club-rush *'' Scirpus sylvaticus'' wood club-rush *'' Scirpus triqueter'' triangular club-rush ;Selected species in a broader view of the genus *''
Bolboschoenus maritimus ''Bolboschoenus maritimus'' is a species of flowering plant from family Cyperaceae. Common names for this species include sea clubrush, cosmopolitan bulrush, alkali bulrush, saltmarsh bulrush, and bayonet grass. It is found in seaside wetland ha ...
'' sea club-rush *''
Isolepis cernua ''Isolepis cernua'' (basionym ''Scirpus cernuus'') is a species of flowering plant in the sedge family known by the common names low bulrush, slender club-rush, tufted clubrush, and fiberoptic grass. It is widespread, being native to many regions ...
'' slender club-rush *'' Isolepis setacea'' bristle club-rush *'' Schoenoplectus acutus'' tule *'' Schoenoplectus hudsonianus'' alpine deergrass *''
Schoenoplectus lacustris ''Schoenoplectus lacustris'', the lakeshore bulrush or common club-rush, is a species of club-rush (genus '' Schoenoplectus'') that grows in fresh water across Europe and some neighbouring areas. Description ''Schoenoplectus lacustris'' grows up ...
'' common club-rush *'' Schoenoplectus tabernaemontani'' *'' Scirpoides holoschoenus'' round-headed club-rush


Fossil record

Several hundred
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
fruits of ''Scirpus ragozinii'' have been described from middle Miocene
strata In geology and related fields, a stratum ( : strata) is a layer of rock or sediment characterized by certain lithologic properties or attributes that distinguish it from adjacent layers from which it is separated by visible surfaces known as e ...
of the Fasterholt area near
Silkeborg Silkeborg () is a Danish town with a population of 49,747 (1 January 2022).Jutland Jutland ( da, Jylland ; german: Jütland ; ang, Ēota land ), known anciently as the Cimbric or Cimbrian Peninsula ( la, Cimbricus Chersonesus; da, den Kimbriske Halvø, links=no or ; german: Kimbrische Halbinsel, links=no), is a peninsula of ...
,
Denmark ) , song = ( en, "King Christian stood by the lofty mast") , song_type = National and royal anthem , image_map = EU-Denmark.svg , map_caption = , subdivision_type = Sovereign state , subdivision_name = Kingdom of Denmark , establishe ...
. 35
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
fruits of the extant ''Scirpus sylvaticus'' have been extracted from
borehole A borehole is a narrow shaft bored in the ground, either vertically or horizontally. A borehole may be constructed for many different purposes, including the extraction of water ( drilled water well and tube well), other liquids (such as petro ...
samples of the Middle Miocene fresh water deposits in Nowy Sacz Basin, West Carpathians,
Poland Poland, officially the Republic of Poland, , is a country in Central Europe. Poland is divided into Voivodeships of Poland, sixteen voivodeships and is the fifth most populous member state of the European Union (EU), with over 38 mill ...
.Łańcucka-Środoniowa M.: Macroscopic plant remains from the freshwater Miocene of the Nowy Sącz Basin (West Carpathians, Poland) zczątki makroskopowe roślin z miocenu słodkowodnego Kotliny Sądeckiej (Karpaty Zachodnie, Polska) Acta Palaeobotanica 1979 20 (1): 3-117.


Uses

''Scirpus'' species are often planted to inhibit
soil erosion Soil erosion is the denudation or wearing away of the upper layer of soil. It is a form of soil degradation. This natural process is caused by the dynamic activity of erosive agents, that is, water, ice (glaciers), snow, air (wind), plants, a ...
and provide habitat for other wildlife. They are also used in some herbal remedies; the plant's
rhizomes In botany and dendrology, a rhizome (; , ) is a modified subterranean plant stem that sends out roots and shoots from its nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from axillary buds and grow hori ...
are collected in the autumn and winter and dried in the sun before use.


References


External links


Genus ''Scirpus''






* ttp://www.northernbushcraft.com/plants/bulrush/notes.htm Edibility of Scirpus Edible parts and identification of ''Scirpus''


Sources

* Muntz, Philip A. ''A California Flora''. Berkeley, CA: University of California Press, 1973, copyright 1959 * Muntz, Philip A. ''A California Flora: Supplement’’. Berkeley, CA: University of California Press, 1976 (Scirpus lacutris, validus, glaucus, p. 183)) {{Taxonbar, from=Q159220 Cyperaceae genera