The brown meagre or corb (''Sciaena umbra'') is a species of marine

ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fish or actinopterygians, is a class of bony fish that comprise over 50% of living vertebrate species. They are so called because of their lightly built fins made of webbings of sk ...

belonging to the

family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...

Sciaenidae Sciaenidae is a family (biology), family of ray-finned fishes belonging to the Order (biology), order Acanthuriformes. They are commonly called drums or croakers in reference to the repetitive throbbing or drumming sounds they make. The family co ...

, the drums and croakers. This species found in, the eastern Atlantic,

Mediterranean Sea The Mediterranean Sea ( ) is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean basin and almost completely enclosed by land: on the east by the Levant in West Asia, on the north by Anatolia in West Asia and Southern Eur ...

and

Black Sea The Black Sea is a marginal sea, marginal Mediterranean sea (oceanography), mediterranean sea lying between Europe and Asia, east of the Balkans, south of the East European Plain, west of the Caucasus, and north of Anatolia. It is bound ...

occurring in shallow waters and sandy bottoms. It is harvested for human consumption, especially in the Mediterranean.

Taxonomy

The brown meagre was first formally described in 1758 by

Carl Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...

in the 10th edition of the ''

Systema Naturae ' (originally in Latin written ' with the Orthographic ligature, ligature æ) is one of the major works of the Sweden, Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the syste ...

''. Linnaeus gave the type locality as

Zadar Zadar ( , ), historically known as Zara (from Venetian and Italian, ; see also other names), is the oldest continuously inhabited city in Croatia. It is situated on the Adriatic Sea, at the northwestern part of Ravni Kotari region. Zadar ...

in Croatia. In 1972 the International Commission for Zoological Nomenclature confirmed that this species was the

type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...

of the

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

Sciaena ''Sciaena'' is a genus of marine ray-finned fishes belonging to the family Sciaenidae, the drums and croakers. These fishes are found in the Eastern Pacific Ocean and the Eastern Atlantic Ocean. Taxonomy ''Sciaena'' was first proposed as a gen ...

''. The genus ''Sciaena'' is placed in the

subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end botanical subfamily names with "-oideae", and zo ...

Sciaeninae by some authors but the 5th edition of ''

Fishes of the World ''Fishes of the World'' is a standard reference for the systematics of fishes. It was first written in 1976 by the American ichthyologist Joseph S. Nelson (1937–2011). Now in its fifth edition (2016), the work is a comprehensive overview of t ...

'' does not recognise subfamilies within the Sciaenidae which it places in the

order Order, ORDER or Orders may refer to: * A socio-political or established or existing order, e.g. World order, Ancien Regime, Pax Britannica * Categorization, the process in which ideas and objects are recognized, differentiated, and understood ...

Acanthuriformes Acanthuriformes is a large, diverse order of mostly marine ray-finned fishes, part of the Percomorpha clade. In the past, members of this clade were placed in the suborders Acanthuroidea and Percoidea of the order Perciformes, but this treatment ...

Etymology

The specific name ''umbra'' is derived from the

Latin Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...

for a shadow or phantom while the generic name is derived from the

Greek Greek may refer to: Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group *Greek language, a branch of the Indo-European language family **Proto-Greek language, the assumed last common ancestor of all kno ...

''skiaina'' or ''skion'' meaning a fish, or more specifically a

red mullet The red mullets or surmullets are two species of goatfish, '' Mullus barbatus'' and ''Mullus surmuletus'', found in the Mediterranean Sea, east North Atlantic Ocean, and the Black Sea. Both "red mullet" and "surmullet" can also refer to the Mull ...

Distribution

The brown meagre is found in the eastern Atlantic Ocean from the southern

English Channel The English Channel, also known as the Channel, is an arm of the Atlantic Ocean that separates Southern England from northern France. It links to the southern part of the North Sea by the Strait of Dover at its northeastern end. It is the busi ...

south to

Senegal Senegal, officially the Republic of Senegal, is the westernmost country in West Africa, situated on the Atlantic Ocean coastline. It borders Mauritania to Mauritania–Senegal border, the north, Mali to Mali–Senegal border, the east, Guinea t ...

and

Cape Verde Cape Verde or Cabo Verde, officially the Republic of Cabo Verde, is an island country and archipelagic state of West Africa in the central Atlantic Ocean, consisting of ten volcanic islands with a combined land area of about . These islands ...

, including the

Canary Islands The Canary Islands (; ) or Canaries are an archipelago in the Atlantic Ocean and the southernmost Autonomous communities of Spain, Autonomous Community of Spain. They are located in the northwest of Africa, with the closest point to the cont ...

, records from West Africa south of Senegal are questionable. Also in the Mediterranean Sea, (where IUCN classifies it as '' Vulnerable'') the

and

Sea of Azov The Sea of Azov is an inland Continental shelf#Shelf seas, shelf sea in Eastern Europe connected to the Black Sea by the narrow (about ) Strait of Kerch, and sometimes regarded as a northern extension of the Black Sea. The sea is bounded by Ru ...

Habitat

The brown meagre is found at depths between , mainly over rocky and sandy substrates and the young enter estuarine environments.

Description

The brown meagre is between in length but can grow to . It has a flat belly and its strongly arched back which give it an easily recognisable shape, the body is laterally compressed and the large, horizontal mouth reaches the level of the eye and contains villiform teeth. The anal and pelvic fins are black with an anterior white border. Both dorsal fins and the truncate caudal fin, are yellow with a black border. The body is grey with flashes of gold and silver. The scales are

ctenoid A fish scale is a small rigid plate that grows out of the skin of a fish. The skin of most jawed fishes is covered with these protective scales, which can also provide effective camouflage through the use of reflection and colouration, as w ...

on the nape and the body while the head scales are

cycloid In geometry, a cycloid is the curve traced by a point on a circle as it Rolling, rolls along a Line (geometry), straight line without slipping. A cycloid is a specific form of trochoid and is an example of a roulette (curve), roulette, a curve g ...

Biology and behaviour

It is a rather nocturnal fish but it can occasionally be found during the day among beds of sea grass and on rocky bottoms in the vicinity of caves or large crevices where it can shelter. This species is social and lives in small groups. It feeds off the small fishes and crustaceans. They are capable of creating sound using some muscles under their well developed swim bladder. This is their way of communication since they have a very good hearing ability. They can manage their buoyancy perfectly. The spawning period is from March to August in the Mediterranean.

Fisheries

The brown meagre is a commercial species throughout the Mediterranean basin and has been over exploited by fishing practices. It is fished mainly by spear fishing, trammel nets, and gill nets and is heavily exposed to fisheries during spawning aggregations at the mouths of estuaries. Sold fresh or frozen across fish markets in Turkey. The

otolith An otolith (, ' ear + , ', a stone), also called otoconium, statolith, or statoconium, is a calcium carbonate structure in the saccule or utricle (ear), utricle of the inner ear, specifically in the vestibular system of vertebrates. The saccule ...

s of the brown meagre are ground and used for urinary infections by local people in Turkey. The sport and

commercial fishing Commercial fishing is the activity of catching fish and other seafood for Commerce, commercial Profit (economics), profit, mostly from wild fisheries. It provides a large quantity of food to many countries around the world, but those who practice ...

of this species was banned in Turkey until 2003 and 2006 to help conserve the stock.

References

External links

* {{Taxonbar, from=Q1383737 Fish of the Black Sea Fish of the Mediterranean Sea Fish described in 1758 Taxa named by Carl Linnaeus Fish of West Asia Fish of Europe Sciaena