Schoeniophylax
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The chotoy spinetail (''Schoeniophylax phryganophilus'') is a species of
bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
in the Furnariinae subfamily of the ovenbird
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Furnariidae. It is found in
Argentina Argentina, officially the Argentine Republic, is a country in the southern half of South America. It covers an area of , making it the List of South American countries by area, second-largest country in South America after Brazil, the fourt ...
,
Bolivia Bolivia, officially the Plurinational State of Bolivia, is a landlocked country located in central South America. The country features diverse geography, including vast Amazonian plains, tropical lowlands, mountains, the Gran Chaco Province, w ...
,
Brazil Brazil, officially the Federative Republic of Brazil, is the largest country in South America. It is the world's List of countries and dependencies by area, fifth-largest country by area and the List of countries and dependencies by population ...
,
Paraguay Paraguay, officially the Republic of Paraguay, is a landlocked country in South America. It is bordered by Argentina to the Argentina–Paraguay border, south and southwest, Brazil to the Brazil–Paraguay border, east and northeast, and Boli ...
, and
Uruguay Uruguay, officially the Oriental Republic of Uruguay, is a country in South America. It shares borders with Argentina to its west and southwest and Brazil to its north and northeast, while bordering the Río de la Plata to the south and the A ...
.


Taxonomy and systematics

The chotoy spinetail was originally described in genus '' Sylvia''. Later authors placed it in ''
Synallaxis ''Synallaxis'' is a genus of birds in the Ovenbird (family), ovenbird family, Furnariidae. It is one of the most diverse genera in the family and is composed of small birds that inhabit dense undergrowth across tropical and subtropical habitats ...
'' but by the early 2010s it was recognized in its own genus ''Schoeniophylax''. It and the white-bellied spinetail (''Mazaria propinqua'') are
sister species In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and ...
.Remsen, Jr., J. V. (2020). Chotoy Spinetail (''Schoeniophylax phryganophilus''), version 1.0. In Birds of the World (J. del Hoyo, A. Elliott, J. Sargatal, D. A. Christie, and E. de Juana, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bow.chospi2.01 retrieved November 20, 2023Derryberry, E. P., S. Claramunt, G. Derryberry, R. T. Chesser, J. Cracraft, A. Aleixo, J. Pérez-Emán, J. V. Remsen, Jr., and R. T. Brumfield. (2011). Lineage diversification and morphological evolution in a large-scale continental radiation: the Neotropical ovenbirds and woodcreepers (Aves: Furnariidae). Evolution 65(10):2973–2986. https://doi.org/10.1111/j.1558-5646.2011.01374.x A high resolution version of the phylogenetic tree in Figure 1 is available from the first author's websit
here
The chotoy spinetail has two subspecies, the
nominate Nomination is part of the process of selecting a candidate for either election to a public office, or the bestowing of an honor or award. A collection of nominees narrowed from the full list of candidates is a short list. Political office In th ...
''S. p. phryganophilus'' (
Vieillot Louis Pierre Vieillot (10 May 1748, Yvetot – 24 August 1830, Sotteville-lès-Rouen) was a French ornithologist. Vieillot is the author of the first scientific descriptions and Linnaean names of a number of birds, including species he collecte ...
, 1817) and ''S. p. petersi'' (
Pinto Pinto is a Portuguese, Spanish, Jewish (Sephardic), and Italian surname. It is a high-frequency surname in all Portuguese-speaking countries and is also widely present in Spanish-speaking countries, Italy, India (especially in Mangalore, Karnata ...
, 1949).


Description

The chotoy spinetail is long and weighs . It is a large spinetail with a unique throat pattern. The sexes have the same plumage. Adults of the nominate subspecies have a white
supercilium The supercilium is a plumage feature found on the heads of some bird species. It is a stripe which runs from the base of the bird's beak above its eye, finishing somewhere towards the rear of the bird's head.Dunn and Alderfer (2006), p. 10 Also k ...
and light brown ear
coverts A covert feather or tectrix on a bird is one of a set of feathers, called coverts (or ''tectrices''), which cover other feathers. The coverts help to smooth airflow over the wings and tail. Ear coverts The ear coverts are small feathers behind t ...
. Their forehead is dark brown, their crown chestnut with faint brown streaks, their back sandy brown with obvious dark brown streaks, and their lower back, rump, and uppertail
coverts A covert feather or tectrix on a bird is one of a set of feathers, called coverts (or ''tectrices''), which cover other feathers. The coverts help to smooth airflow over the wings and tail. Ear coverts The ear coverts are small feathers behind t ...
light brown. Their wings are mostly light brown with a chestnut shoulder. Their tail is long and light brown with blackish feather shafts. Their chin and upper throat are bright yellow; the lower throat is a crisp black square surrounded by white. Their upper breast is apricot, their lower breast buffy white, their belly whitish, their flanks brownish buff, and their undertail coverts rufescent buff. Their iris is red to brown, their
maxilla In vertebrates, the maxilla (: maxillae ) is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The two maxil ...
blackish to dark gray, their
mandible In jawed vertebrates, the mandible (from the Latin ''mandibula'', 'for chewing'), lower jaw, or jawbone is a bone that makes up the lowerand typically more mobilecomponent of the mouth (the upper jaw being known as the maxilla). The jawbone i ...
gray to bluish (sometimes with a dark tip), and their legs and feet light gray to brownish. Juveniles have less sharp streaking on their uppersides than adults, a brown crown, a whitish chin, and an indistinct black throat patch. Subspecies ''S. p. petersi'' has plumage like the nominate's but is smaller.


Distribution and habitat

The nominate subspecies of the chotoy spinetail is by far the more widespread of the two. It is found in eastern Bolivia,
Mato Grosso do Sul Mato Grosso do Sul ( ) is one of Federative units of Brazil, Brazil's 27 federal units, located in the southern part of the Central-West Region, Brazil, Central-West Region, bordering five Brazilian states: Mato Grosso (to the north), Goiás and ...
and
Rio Grande do Sul Rio Grande do Sul (, ; ; "Great River of the South") is a Federative units of Brazil, state in the South Region, Brazil, southern region of Brazil. It is the Federative units of Brazil#List, fifth-most populous state and the List of Brazilian s ...
in Brazil, most of Paraguay, northeastern Argentina as far south as northern
Buenos Aires Province Buenos Aires, officially the Buenos Aires Province, is the largest and most populous Provinces of Argentina, Argentine province. It takes its name from the city of Buenos Aires, the capital of the country, which used to be part of the province an ...
, and essentially all of Uruguay. Subspecies ''S. p. petersi'' is found disjunctly in northeastern Brazil's states of
Minas Gerais Minas Gerais () is one of the 27 federative units of Brazil, being the fourth largest state by area and the second largest in number of inhabitants with a population of 20,539,989 according to the 2022 Brazilian census, 2022 census. Located in ...
,
Bahia Bahia () is one of the 26 Federative units of Brazil, states of Brazil, located in the Northeast Region, Brazil, Northeast Region of the country. It is the fourth-largest Brazilian state by population (after São Paulo (state), São Paulo, Mina ...
, and far separated in
Piauí Piauí ( ) is one of the states of Brazil, located in the country's Northeast Region, Brazil, Northeast Region. The state has 1.6% of the Brazilian population and produces 0.7% of the Brazilian GDP. Piauí has the shortest coastline of any coas ...
. The species inhabits a variety of open to semi-open landscapes including treed savanna;
gallery forest A gallery forest is one formed as a corridor along rivers or wetlands, projecting into landscapes that are otherwise only sparsely treed such as savannas, grasslands, or deserts. The gallery forest maintains a more temperate microclimate above th ...
, thickets, and scrub along watercourses; monte woodlands; and the edges of marshes. In elevation it ranges from near sea level to .


Behavior


Movement

The chotoy spinetail is a year-round resident throughout its range.


Feeding

The chotoy spinetail feeds on
arthropod Arthropods ( ) are invertebrates in the phylum Arthropoda. They possess an arthropod exoskeleton, exoskeleton with a cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate, a body with differentiated (Metam ...
s. It typically forages singly or in pairs, gleaning prey from small branches and foliage within about of the ground.


Breeding

The chotoy spinetail breeds in the austral spring and summer, roughly late September to January. It is thought to be monogamous. Its nest is a large ball of thorny sticks with an entrance tube on the side; the tube and inner chamber are lined with softer plant material. It is typically placed in a bush or small tree up to about above the ground but occasionally much higher. The clutch size is three to six eggs but usually four or five. The incubation period is 15 to 16 days and fledging occurs 13 to 14 days after hatch.


Vocalization

The chotoy spinetail's song is "a distinctive series of low-pitched “cho” notes" that sound like they come from a larger bird. It has been rendered as "rrrrtooo chicheetitichichicheecheychachochoochew". Its call is a "hurried 'sreepsreeptrrit' ".


Status

The
IUCN The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the status ...
has assessed the chotoy spinetail as being of Least Concern. It has a large range and an unknown population size that is believed to be stable. No immediate threats have been identified. It is considered fairly common to common and appears "fairly adaptable, occurring in secondary habitats and scrub".


References


External links


Chotoy spinetail photo gallery
VIREO
Photo-Medium Res
chandra.as.utexas.edu–''"Birds of Brazil"''
Photo-High Res
Tropical Birding {{Taxonbar, from=Q428748 Furnariidae Birds of Uruguay Birds of Paraguay Birds of the Pantanal Birds of Bolivia Birds of Argentina Birds of Brazil Birds described in 1817 Taxa named by Louis Pierre Vieillot Taxonomy articles created by Polbot