Schleitheimia
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''Schleitheimia'' (named after the type locality of Schleitheim), is an extinct genus of
sauropodiform Sauropodomorpha ( ; from Greek, meaning "lizard-footed forms") is an extinct clade of long-necked, herbivorous, saurischian dinosaurs that includes the sauropods and their ancestral relatives. Sauropods generally grew to very large sizes, had lon ...
sauropodomorph Sauropodomorpha ( ; from Greek, meaning "lizard-footed forms") is an extinct clade of long-necked, herbivorous, saurischian dinosaurs that includes the sauropods and their ancestral relatives. Sauropods generally grew to very large sizes, had lo ...
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...
, from the Gruhalde Member of
Klettgau Formation The Klettgau Formation is a geological formation in Switzerland. It is Late Triassic in age, covering most of the mid to late Norian, the Carnian, and into the Rhaetian, spanning a period of 26-30 million years.Switzerland Switzerland, officially the Swiss Confederation, is a landlocked country located in west-central Europe. It is bordered by Italy to the south, France to the west, Germany to the north, and Austria and Liechtenstein to the east. Switzerland ...
. The
type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
, ''Schleitheimia schutzi'' was formally described in 2020.


Discovery and naming

The type material was collected between 1952 and 1954 by Emil Schutz. They were donated to the
University of Zürich The University of Zurich (UZH, ) is a public university, public research university in Zurich, Switzerland. It is the largest university in Switzerland, with its 28,000 enrolled students. It was founded in 1833 from the existing colleges of the ...
in 1955. In 1986, most of the material was described by
Peter Galton Peter Malcolm Galton (born 14 March 1942 in London) is a British vertebrate paleontologist who has to date written or co-written about 190 papers in scientific journals or chapters in paleontology textbooks, especially on ornithischian and prosau ...
, who referred them all to the species ''
Plateosaurus engelhardti ''Plateosaurus'' (probably meaning "broad lizard", often mistranslated as "flat lizard") is a genus of plateosaurid dinosaur that lived during the Late Triassic period, around 214 to 204 million years ago, in what is now Central and Northern Eu ...
''. In 2020, Oliver Rauhut, Femke Holwerda and Heinz Furrer redescribed most of Schutz's remains, as well as some remains in the collections of the
Museum zu Allerheiligen A museum is an institution dedicated to displaying or preserving culturally or scientifically significant objects. Many museums have exhibitions of these objects on public display, and some have private collections that are used by researchers ...
in Schaffhausen and new remains from an excavation in 2016 led by Holwerda. They found that the earlier found remains represented a new genus and species, which they named ''Schleitheimia schultzi'', honoring the type locality and the discoverer of the type remains. The 2016 finds were not referred. Finds made by Schultz near Hallau-Schwärzibuck were considered to represent a different previously unknown taxon and remained unnamed. The
holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
, PIMUZ A/III 550, was found in a layer of the Klettgau Formation dating from the
Norian The Norian is a division of the Triassic geological period, Period. It has the rank of an age (geology), age (geochronology) or stage (stratigraphy), stage (chronostratigraphy). It lasted from ~227.3 to Mya (unit), million years ago. It was prec ...
. It consists of a partial right ilium. Several other bones from the general locality were designated as
paratype In zoology and botany, a paratype is a specimen of an organism that helps define what the scientific name of a species and other taxon actually represents, but it is not the holotype (and in botany is also neither an isotype (biology), isotype ...
s. Because they are consistent in morphology, the describers referred all of them to the same taxon, if not the same individual.


Description

The holotype of ''Schleitheimia'' shows a number of
autapomorphies In phylogenetics, an autapomorphy is a distinctive feature, known as a derived trait, that is unique to a given taxon. That is, it is found only in one taxon, but not found in any others or outgroup taxa, not even those most closely related to ...
, unique derived traits. There is a broad, rounded ridge on the iliac blade that ends in a large, rounded expansion on the ilium. The fourth tronchanter of the ilium is very robust. The crista tibiofibularis of the femur is broad and there is no posteriorly facing shelf is present lateral to the crista.


Classification

A
phylogenetic analysis In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical data ...
conducted by Rauhut and colleagues found ''Schleitheimia'' to be a derived sauropodiform closely related to
Sauropoda Sauropoda (), whose members are known as sauropods (; from ''wikt:sauro-, sauro-'' + ''wikt:-pod, -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative t ...
, or a basal member of that clade, depending on the definition used (node or stem based). A
cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ...
from that analysis is shown below:


References

{{Taxonbar, from=Q96777241 Massopoda Dinosaur genera Norian dinosaurs Fossil taxa described in 2020 Dinosaurs of Europe