Scardinius Hesperidicus
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''Scardinius hesperidicus'', the Italian rudd is a
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
of freshwater
ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fish or actinopterygians, is a class of bony fish that comprise over 50% of living vertebrate species. They are so called because of their lightly built fins made of webbings of sk ...
belonging to the
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Leuciscidae Leuciscidae is a family of freshwater ray-finned fishes, formerly classified as a subfamily of the Cyprinidae, which contains the true minnows. Members of the Old World (OW) clade of minnows within this subfamily are known as European minnow ...
, which includes the daces, Eurasian minnows and related fishes. This species is found in the Po and
Adriatic The Adriatic Sea () is a body of water separating the Italian Peninsula from the Balkans, Balkan Peninsula. The Adriatic is the northernmost arm of the Mediterranean Sea, extending from the Strait of Otranto (where it connects to the Ionian Se ...
drainages east of the Po in Italy, San Marino and Switzerland, and has been introduced into other area watersheds, especially in Italy.


Taxonomy

''Scardinius hesperidicus'' was first formally described in 1845 by the French
art collector A private collection is a privately owned collection of works (usually artworks) or valuable items. In a museum or art gallery context, the term signifies that a certain work is not owned by that institution, but is on loan from an individual ...
and
biologist A biologist is a scientist who conducts research in biology. Biologists are interested in studying life on Earth, whether it is an individual Cell (biology), cell, a multicellular organism, or a Community (ecology), community of Biological inter ...
Charles Lucien Bonaparte Charles Lucien Jules Laurent Bonaparte, 2nd Prince of Canino and Musignano (24 May 1803 – 29 July 1857) was a French naturalist and ornithology, ornithologist, and a nephew of Napoleon. Lucien and his wife had twelve children, including Cardinal ...
with its type locality given as the lakes of
Piedmont Piedmont ( ; ; ) is one of the 20 regions of Italy, located in the northwest Italy, Northwest of the country. It borders the Liguria region to the south, the Lombardy and Emilia-Romagna regions to the east, and the Aosta Valley region to the ...
in Italy. This species was formerly considered to be a
synonym A synonym is a word, morpheme, or phrase that means precisely or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are a ...
of the
common rudd The common rudd (''Scardinius erythrophthalmus'') is a benthos, bentho-pelagic freshwater Actinopterygii, ray-finned fish belonging to the Family (biology), family Leuciscidae. This species is widely spread in Europe and central Asia, around the ...
(''S. erythrophthalmus''). The Italian rudd belongs to the genus '' Scardinius'', commonly referred to as rudds, which belongs to the
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end botanical subfamily names with "-oideae", and zo ...
Leuciscinae Leuciscinae is a subfamily of freshwater Actinopterygii, ray-finned fishes belonging to the Family (biology), family Leuciscidae, which includes the fishes known as daces, chubs, shiners and minnows. The fishes in this subfamily are mainly found i ...
of the family Leuciscidae. The Italian rudd is very similar to the Tiber rudd ('' S. scardafa'') of southern and Central Italy and have been treated as conspecific. These two rudds do show meristic and molecular differences and are currently treated as separate valid, albeit closely related species. The
sister taxon In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and ...
to these two is the Neretva rudd ('' S. plotizza'').


Etymology

''Scardinius hesperidicus'' belongs to the genus ''Scardinius'' and this name is thought to be a latinisation of ''scardafa'', a vernacular name in Italy, Rome in particular, for the Tiber rudd ('' Scardinius scardafa''). The
Specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
, ''hesperidicus'', means "belonging to the
Hesperides In Greek mythology, the Hesperides (; , ) are the nymphs of evening and golden light of sunsets, who were the "Daughters of the Evening" or "Nymphs of the West". They were also called the Atlantides () from their reputed father, Atlas (mytholog ...
, these were nymphs in Greek mythology who were said to tend a paradisical garden in a far western corner of the world, this may be an allusion to Piedmont being in the far west of Italy.


Description

''Scardinius hesperidicus'' may be told apart from other Mediterranean rudd species by having between 36 and 39 scales along the
lateral line The lateral line, also called the lateral line organ (LLO), is a system of sensory organs found in fish, used to detect movement, vibration, and pressure gradients in the surrounding water. The sensory ability is achieved via modified epithelia ...
; the
anal fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported o ...
has between 10 and branched fin rays; 10-13 gill rakers; the dorsal profile of the head is straight or a litte convex with the snout pointing forward, tip at or slightly above the level of the middle of the eye; there is no hump behind the eye; the eye is not close to the dorsal profile of the head in lateral view; the articulation of lower jaw is to the front of the of eye; the head length is equivalent to 24-272% of the
standard length Fish measurement is the measuring of individual fish and various parts of fish anatomy, their anatomies, for data used in many areas of ichthyology, including Taxonomy (biology), taxonomy and fishery biology. Overall length Standard length (SL) is ...
while the depth of the body is equivalent to 35-39% of the standard length; and all fins dark grey in adults with a standard length greater than . This species has a maximum standard length of .


Distribution and habitat

''Scardinius hesperidicus'' is endemic to the northern drainage basin of the Adriatic Sea where its range goes from the Isonzo in
Friuli-Venezia Giulia Friuli-Venezia Giulia () is one of the 20 regions of Italy and one of five autonomous regions with special statute. The regional capital is Trieste on the Gulf of Trieste, a bay of the Adriatic Sea. Friuli-Venezia Giulia has an area of and a ...
south to the Vomano in
Abruzzo Abruzzo (, ; ; , ''Abbrìzze'' or ''Abbrèzze'' ; ), historically also known as Abruzzi, is a Regions of Italy, region of Southern Italy with an area of 10,763 square km (4,156 sq mi) and a population of 1.3 million. It is divided into four ...
. It is also found in the rivers draining into the Ligurian and
Tyrrhenian Sea The Tyrrhenian Sea (, ; or ) , , , , is part of the Mediterranean Sea off the western coast of Italy. It is named for the Tyrrhenians, Tyrrhenian people identified with the Etruscans of Italy. Geography The sea is bounded by the islands of C ...
s in the drainages of the
Arno The Arno is a river in the Tuscany region of Italy. It is the most important river of central Italy after the Tiber. Source and route The river originates on Monte Falterona in the Casentino area of the Apennines, and initially takes a sou ...
,
Ombrone The Ombrone (Latin: ''Umbro'') is a river situated in the provinces of Grosseto and Siena, central Italy. The Ombrone's source is at San Gusmè, near Castelnuovo Berardenga, on the south-eastern side of the Monti del Chianti. After a twistin ...
,
Tiber The Tiber ( ; ; ) is the List of rivers of Italy, third-longest river in Italy and the longest in Central Italy, rising in the Apennine Mountains in Emilia-Romagna and flowing through Tuscany, Umbria, and Lazio, where it is joined by the R ...
and
Volturno The Volturno (ancient Latin name Volturnus, from ''volvere'', to roll) is a river in south-central Italy. Geography It rises in the Abruzzese central Apennines of Samnium near Castel San Vincenzo (province of Isernia, Molise) and flows southe ...
but there is a possibility that the Italian rudd is an introduced species in these systems. It has been introduced to some subalpine lakes in France and Switzerland. This is a species of slow or still waters in rivers but it readily colonises drainage channels, canals and reservoirs.


Biology

''Scardinius hesperidicus'' is a
demersal fish Demersal fish, also known as groundfish, live and feed on or near the bottom of seas or lakes (the demersal zone).Walrond Carl . "Coastal fish - Fish of the open sea floor"Te Ara - the Encyclopedia of New Zealand. Updated 2 March 2009 They oc ...
schooling fish, although it can be pelagic in smaller lakes. The shoals ten to congregate in the littoral and benthic zones of lakes, down to , frequently close to reedbeds or other macrophyte beds. There are omnivorous and feed on both plankton and benthic invertebrates, as well as plant matter and insects taken from the surface. These fishes can live up to 15 years and attain sexual maturirty at 2 or 3 years old. The
spawning Spawn is the Egg cell, eggs and Spermatozoa, sperm released or deposited into water by aquatic animals. As a verb, ''to spawn'' refers to the process of freely releasing eggs and sperm into a body of water (fresh or marine); the physical act is ...
season runs from March to July. Females spawn several times in a season and the eggs are laid among submerged vegetation.


References

{{Taxonbar, from=Q2892057 hesperidicus Fish described in 1845 Taxa named by Charles Lucien Bonaparte