Scarce Copper
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The scarce copper (''Lycaena virgaureae'') is a
butterfly Butterflies are winged insects from the lepidopteran superfamily Papilionoidea, characterized by large, often brightly coloured wings that often fold together when at rest, and a conspicuous, fluttering flight. The oldest butterfly fossi ...
of the family
Lycaenidae Lycaenidae is the second-largest family (biology), family of butterflies (behind Nymphalidae, brush-footed butterflies), with over 6,000 species worldwide, whose members are also called gossamer-winged butterflies. They constitute about 30% of ...
(copper or gossamer-winged butterflies).


Appearance

The lower surfaces of the back wings are yellowish and have only a few black dots; there are characteristic white marks in the immediate vicinity of these. The species exhibits one kind of
sexual dimorphism Sexual dimorphism is the condition where sexes of the same species exhibit different Morphology (biology), morphological characteristics, including characteristics not directly involved in reproduction. The condition occurs in most dioecy, di ...
: male butterflies are colored bright gold-red on the upper side of wing, while the females have broader orange wings with a dark design. (MHNT) Lycaena virgaureae - Radawa Poland - male dorsal.jpg , ''Lycaena virgaureae'' ♂ (MHNT) Lycaena virgaureae - Radawa Poland - male ventral.jpg , ''Lycaena virgaureae'' ♂ △


Life cycle

A generation appears from mid-July to mid-September. Eggs are laid on dried-out plant parts, for example on dry
sorrel Sorrel (''Rumex acetosa''), also called common sorrel or garden sorrel, is a perennial herbaceous plant in the family Polygonaceae. Other names for sorrel include spinach dock and narrow-leaved dock ("dock" being a common name for the genus ''Ru ...
stems. The eggs are white in color and somewhat larger than those of other ''Lycaena'' species. The caterpillars are green and nocturnal and eat sorrel. ''Lycaena virgaureae'' is the only species of this genus whose eggs last over the winter. The butterflies feed on blossoms of such plants as the
ground-elder ''Aegopodium podagraria'', commonly called ground elder, is a species of flowering plant in the carrot family Apiaceae that grows in shady places. The name "ground elder" comes from the superficial similarity of its leaves and flowers to those o ...
, ''
Eupatorium ''Eupatorium'' is a genus of flowering plants in the family Asteraceae, containing from 36 to 60 species depending on the classification system. Most are Herbaceous plant, herbaceous perennial plant, perennials growing to tall. A few are shrubs ...
'', ''
Valeriana ''Valeriana'' is a genus of flowering plants in the family Caprifoliaceae, members of which may be commonly known as valerians. It contains many species, including the garden valerian, ''Valeriana officinalis''. ''Valeriana'' has centers of div ...
'', and
burnet saxifrage ''Pimpinella saxifraga'', known as burnet-saxifrage, solidstem burnet saxifrage, lesser burnet is a plant species in the family Apiaceae, a native of the British Isles and temperate Europe and Western Asia. It is neither a burnet, which its l ...
.


Habitat

''Lycaena virgaureae'' lives preferably on flower-rich, dry and damp meadows in
Central Europe Central Europe is a geographical region of Europe between Eastern Europe, Eastern, Southern Europe, Southern, Western Europe, Western and Northern Europe, Northern Europe. Central Europe is known for its cultural diversity; however, countries in ...
. In
Spain Spain, or the Kingdom of Spain, is a country in Southern Europe, Southern and Western Europe with territories in North Africa. Featuring the Punta de Tarifa, southernmost point of continental Europe, it is the largest country in Southern Eur ...
it is found in the
Pyrenees The Pyrenees are a mountain range straddling the border of France and Spain. They extend nearly from their union with the Cantabrian Mountains to Cap de Creus on the Mediterranean coast, reaching a maximum elevation of at the peak of Aneto. ...
and in the
Cantabrian Mountains The Cantabrian Mountains or Cantabrian Range () are one of the main systems of mountain ranges in Spain. They stretch for over 300 km (180 miles) across northern Spain, from the western limit of the Pyrenees to the Galician Massif ...
. In
southern France Southern France, also known as the south of France or colloquially in French as , is a geographical area consisting of the regions of France that border the Atlantic Ocean south of the Marais Poitevin,Louis Papy, ''Le midi atlantique'', Atlas e ...
it is found in the Massif Central. Its area of distribution reaches the
Arctic Circle The Arctic Circle is one of the two polar circles, and the northernmost of the five major circle of latitude, circles of latitude as shown on maps of Earth at about 66° 34' N. Its southern counterpart is the Antarctic Circle. The Arctic Circl ...
in
Fennoscandia __NOTOC__ Fennoscandia (Finnish language, Finnish, Swedish language, Swedish and ; ), or the Fennoscandian Peninsula, is a peninsula in Europe which includes the Scandinavian Peninsula, Scandinavian and Kola Peninsula, Kola peninsulas, mainland ...
in the north; in the south it reaches north and northwest
Greece Greece, officially the Hellenic Republic, is a country in Southeast Europe. Located on the southern tip of the Balkan peninsula, it shares land borders with Albania to the northwest, North Macedonia and Bulgaria to the north, and Turkey to th ...
. In
Belgium Belgium, officially the Kingdom of Belgium, is a country in Northwestern Europe. Situated in a coastal lowland region known as the Low Countries, it is bordered by the Netherlands to the north, Germany to the east, Luxembourg to the southeas ...
there are a few threatened populations in the south of the country. It is not found, however, on the
British Isles The British Isles are an archipelago in the Atlantic Ocean, North Atlantic Ocean off the north-western coast of continental Europe, consisting of the islands of Great Britain, Ireland, the Isle of Man, the Inner Hebrides, Inner and Outer Hebr ...
or the
Netherlands , Terminology of the Low Countries, informally Holland, is a country in Northwestern Europe, with Caribbean Netherlands, overseas territories in the Caribbean. It is the largest of the four constituent countries of the Kingdom of the Nether ...
. It can be found as far east as
Mongolia Mongolia is a landlocked country in East Asia, bordered by Russia to the north and China to the south and southeast. It covers an area of , with a population of 3.5 million, making it the world's List of countries and dependencies by po ...
. The scarce copper is found at elevations of 1,000 to 2,000 meters, although in Armenia it inhabits meadows up to 2,700 meters above sea level. There have been several scientific debates about the Scarce Copper presence in Britain during the 19th century.


Subspecies

* ''Lycaena virgaureae montanus'' Meyer-Dür, 1851 Found in the
Alps The Alps () are some of the highest and most extensive mountain ranges in Europe, stretching approximately across eight Alpine countries (from west to east): Monaco, France, Switzerland, Italy, Liechtenstein, Germany, Austria and Slovenia. ...
in
France France, officially the French Republic, is a country located primarily in Western Europe. Overseas France, Its overseas regions and territories include French Guiana in South America, Saint Pierre and Miquelon in the Atlantic Ocean#North Atlan ...
,
Switzerland Switzerland, officially the Swiss Confederation, is a landlocked country located in west-central Europe. It is bordered by Italy to the south, France to the west, Germany to the north, and Austria and Liechtenstein to the east. Switzerland ...
,
Italy Italy, officially the Italian Republic, is a country in Southern Europe, Southern and Western Europe, Western Europe. It consists of Italian Peninsula, a peninsula that extends into the Mediterranean Sea, with the Alps on its northern land b ...
,
Germany Germany, officially the Federal Republic of Germany, is a country in Central Europe. It lies between the Baltic Sea and the North Sea to the north and the Alps to the south. Its sixteen States of Germany, constituent states have a total popu ...
, and
Austria Austria, formally the Republic of Austria, is a landlocked country in Central Europe, lying in the Eastern Alps. It is a federation of nine Federal states of Austria, states, of which the capital Vienna is the List of largest cities in Aust ...
at elevations of . * ''Lycaena virgaureae miegii'' Vogel, 1857 North and central Spain at elevations of . * ''Lycaena virgaureae armeniaca'' (A. Bang-Haas, 1906) Endemic of Armenia, occupies elevation range .


Description from Seitz

Male above red-golden, with a narrow black margin, at the proximal side of which there are dark dots on the hindwing: female cinnabar-red, spotted with black, the hindwing partly dusted with black. Underside leather-yellow, more sparsely spotted with black, before the outer third of the hind- wing pale dots, which are occasionally united in a white chain. The anal area dusted with red. The species occurs from the Atlantic coasts throughout Europe to East- Siberia and from the coast of the North Sea to the Mediterranean, but is absent from Great Britain and Japan. — In Lapland flies a small form, ''oranula'' Frr. (76 b), which is otherwise very similar to the name-typical form. — ''estonica'' Huene (76 b), from the Baltic provinces and eastern Russia, exactly resembles the preceding in size and shape, but has a broader black margin. —'' virgaureola'' Stgr. (76 b) is in size like ''virgaureae'', but the upperside of the male is as in ''estonica''; beneath the more reddish disc of the forewing contrasts with the more yellow hindwing and the white discal stripe of the latter is absent or reduced; from northern Central Asia, Dauria, Mongolia, and as similar aberration in the Swiss Alps. — males from the Apennines, in which the upperside is deeper red-golden and the base of the hindwing more densely dusted with dark, are ''apennina'' Calb. The females the contrary have the ground-colour paler and are less dusted with dark than many Central European specimens, the underside being hghter. According to Fallou the males from the Pyrenees are also deeper red. - On the other hand, the specimens from Mersina and the neighbouring Taurus Mts. have a bright light golden-red upperside in both sexes, the black margin being narrowed in the male. This is aureomicans Heyne. — Specimens with the underside of the hindwing strongly dusted with grey, the upperside of the female more-over having a brown-grey tint on account of the dark dusting on the golden ground, occur in many alpine districts with the ordinary form, being especially plentiful and well marked in the Alps of Valais; all such individuals are united as ab. ''zermattensis'' Fall. (76 a). It is hardly possible to decide from Fallou's very long description of this form which are the real characteristics of his aberration, so that strictly speaking not all the specimens standing in collections under this name belong here, particularly not all the specimens from the Valais; in fact only those individuals are true ab. zermattensis which, like ''phlaeas caeruleopunctata'', have a chain of white dots before the outer third of the hindwing, as shown in the figure given in ''Ann. Soc. Ent. Fr.'' vol. V. pl. 2. Besides the modification of the colour of the upperside into sepia-brown, both sexes differ on the underside in the forewing bearing a broad dark margin and the hindwing being washed with fuscous, ab. ''seriata'' Fruhst. are ''zermattensis'' females which have a discal row of white dots on the hindwing proximally to the row of black spots, ab. ''fredegunda'' Fruhst. bears, besides those dots, a row of submarginal spots on the forewing beneath. ore vars describedhe egg is semiglobular, grey-green, with a network of polygons, laid singly or several together on the stalk of the foot-plant. Larva green with a yellow stripe on back and at the sides, brownish head and brownish legs; at first glassy, transparent, later yellow on the back. Emerges in April and feeds until June on ''Rumex'' and ''Solidago''. Pupa rounded, smooth, similar to a small bean, brownish, with dark markings; on the thorax a dark dorsal longitudinal stripe, which is continued on the abdomen by a row of impressed dots. The butterflies are on the wing from the end of June into August on meadows, clearings in woods, mountain-sides and flowery slopes; they are abundant almost everywhere in the area aru:! ascend in the high mountains above 10,000 ft.Seitz, A. ed. Band 1: Abt. 1, ''Die Großschmetterlinge des palaearktischen Faunengebietes, Die palaearktischen Tagfalter'', 1909, 379 Seiten, mit 89 kolorierten Tafeln (3470 Figuren)


Gallery

File:Lycaenidae - Lycaena virgaureae-female.JPG, ''Lycaena virgaureae'', female - lateral view File:Lycaena virgaureae female.JPG, ''Lycaena virgaureae'', female - dorsal view File:Lycaenidae - Lycaena virgaureae - male-2.JPG, ''Lycaena virgaureae'', male - lateral view File:Lycaenidae - Lycaena virgaureae - male.JPG, ''Lycaena virgaureae'', male - dorsal view File:The Macrolepidoptera of the world (Taf. 76) (8145299572).jpg , Seitz


References


External links


www.eurobutterflies.com


* ttp://www.lepiforum.de/cgi-bin/lepiwiki.pl?Lycaena_Virgaureae www.lepiforum.de: Fotos und Taxonomie
www.schmetterling-raupe.de




{{Taxonbar, from=Q520934 Lycaena Butterflies of Asia Butterflies of Europe Insects of Mongolia Butterflies described in 1758 Taxa named by Carl Linnaeus