''Scaniornis'' is a prehistoric

bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...

. The only known

species In biology, a species is the basic unit of Taxonomy (biology), classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of ...

, ''Scaniornis lundgreni'', lived in the MP 1–5 (

Early Paleocene The Danian is the oldest age or lowest stage of the Paleocene Epoch or Series, of the Paleogene Period or System, and of the Cenozoic Era or Erathem. The beginning of the Danian (and the end of the preceding Maastrichtian) is at the Cret ...

, perhaps

Middle Paleocene The Selandian is a stage in the Paleocene. It spans the time between . It is preceded by the Danian and followed by the Thanetian. Sometimes the Paleocene is subdivided in subepochs, in which the Selandian forms the "middle Paleocene". Stratig ...

: c. 65–59 million years ago). It is known from a partial

fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...

skeleton of a right wing, namely the

coracoid A coracoid (from Greek κόραξ, ''koraks'', raven) is a paired bone which is part of the shoulder assembly in all vertebrates except therian mammals (marsupials and placentals). In therian mammals (including humans), a coracoid process is pre ...

scapula The scapula (plural scapulae or scapulas), also known as the shoulder blade, is the bone that connects the humerus (upper arm bone) with the clavicle (collar bone). Like their connected bones, the scapulae are paired, with each scapula on eithe ...

and

humerus The humerus (; ) is a long bone in the arm that runs from the shoulder to the elbow. It connects the scapula and the two bones of the lower arm, the radius and ulna, and consists of three sections. The humeral upper extremity consists of a roun ...

found at

Limhamn Limhamn () is, in an administrative sense, the southern district of Malmö Municipality in Sweden. Before 1915, Limhamn was (briefly) a town of its own. The population of Limhamn-Bunkeflo (including suburbs) is 31,000, of which 7,000 live in t ...

( Sweden) and other bones found at Selk, Germany. Thus, it would seem to have been a native of the prehistoric

North Sea The North Sea lies between Great Britain, Norway, Denmark, Germany, the Netherlands and Belgium. An epeiric sea on the European continental shelf, it connects to the Atlantic Ocean through the English Channel in the south and the Norwegian S ...

, which at that time covered part of today's Germany and

France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of overseas regions and territories in the Americas and the Atlantic, Pacific and Indian Oceans. Its metropolitan ar ...

, and sometimes was cut off from the Tethys and

Atlantic Ocean The Atlantic Ocean is the second-largest of the world's five oceans, with an area of about . It covers approximately 20% of Earth#Surface, Earth's surface and about 29% of its water surface area. It is known to separate the "Old World" of Afr ...

s, sometimes connected to them, and sometimes even to the

Turgai Sea The Turgai Sea, also known as the Turgay Sea, Turgai Strait, Obik Sea, Ural Sea or West Siberian Sea, was a large shallow body of salt water (an epicontinental or epeiric sea) during the Mesozoic through Cenozoic Eras. It extended north of the pr ...

. Situated a bit southwestwards — between 44° and 54° North — of its present location due to

plate tectonic Plate tectonics (from the la, label=Late Latin, tectonicus, from the grc, τεκτονικός, lit=pertaining to building) is the generally accepted scientific theory that considers the Earth's lithosphere to comprise a number of large t ...

s, in a fairly wet and warm

epoch In chronology and periodization, an epoch or reference epoch is an instant in time chosen as the origin of a particular calendar era. The "epoch" serves as a reference point from which time is measured. The moment of epoch is usually decided by ...

, the region had probably a warm-temperate to subtropical and fairly

humid Humidity is the concentration of water vapor present in the air. Water vapor, the gaseous state of water, is generally invisible to the human eye. Humidity indicates the likelihood for precipitation, dew, or fog to be present. Humidity dep ...

climate Climate is the long-term weather pattern in an area, typically averaged over 30 years. More rigorously, it is the mean and variability of meteorological variables over a time spanning from months to millions of years. Some of the meteorologica ...

, altogether not too dissimilar from today's

Black Sea The Black Sea is a marginal mediterranean sea of the Atlantic Ocean lying between Europe and Asia, east of the Balkans, south of the East European Plain, west of the Caucasus, and north of Anatolia. It is bounded by Bulgaria, Georgia, ...

region or French

Mediterranean The Mediterranean Sea is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean Basin and almost completely enclosed by land: on the north by Western and Southern Europe and Anatolia, on the south by North Africa, and on th ...

. It appears to be somewhat similar to

flamingo Flamingos or flamingoes are a type of wading bird in the family Phoenicopteridae, which is the only extant family in the order Phoenicopteriformes. There are four flamingo species distributed throughout the Americas (including the Caribbean) ...

s and was long placed with these, and thus would strongly suggest that the Phoenicopteriformes evolved in the

Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', ...

of immediately thereafter. As flamingos are now thought to be related to

grebe Grebes () are aquatic diving birds in the order Podicipediformes . Grebes are widely distributed freshwater birds, with some species also found in marine habitats during migration and winter. Some flightless species exist as well, most notably ...

s, the placement of ''Scaniornis'' requires reanalysis (Mlíkovský 2002). It was also united with other wading or presumed

shorebird 245px, A flock of Red_knot.html" ;"title="Dunlins and Red knot">Dunlins and Red knots Waders or shorebirds are birds of the order Charadriiformes commonly found wikt:wade#Etymology 1, wading along shorelines and mudflats in order to foraging, ...

s into the " Graculavidae", a

form taxon Form classification is the classification of organisms based on their morphology, which does not necessarily reflect their biological relationships. Form classification, generally restricted to palaeontology, reflects uncertainty; the goal of sc ...

of "transitional shorebirds". This group is now known to be

paraphyletic In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be pa ...

and has no standing in

systematics Biological systematics is the study of the diversification of living forms, both past and present, and the relationships among living things through time. Relationships are visualized as evolutionary trees (synonyms: cladograms, phylogenetic t ...

anymore. The presumed relative ''Parascaniornis'' is now known to be a

hesperornithine Hesperornithes is an extinct and highly specialized group of aquatic avialans closely related to the ancestors of modern birds. They inhabited both marine and freshwater habitats in the Northern Hemisphere, and include genera such as ''Hesperorn ...

of the genus ''

Baptornis ''Baptornis'' ("diving bird") is a genus of flightless, aquatic birds from the Late Cretaceous, some 87-80 million years ago (roughly mid-Coniacian to mid-Campanian faunal stages). The fossils of ''Baptornis advenus'', the type species, were dis ...

''. The Hesperornithes which became extinct by the end of the Cretaceous and ''Scaniornis'' which appears clearly a neornithine are not closely related at all. ''Scaniornis'' was sometimes united with the

Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of ...

Gallornis ''Gallornis'' is a genus of prehistoric birds from the Cretaceous. The single known species ''Gallornis straeleni'' lived near today's Auxerre in Yonne ''département'' ( France); it has been dated very tentatively to the Berriasian- Haute ...

'' in the

family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...

Scaniornithidae. ''Gallornis'', however, is of even more unclear relationships; it might be an early member of the Galloanserae. In any case it was subsequently not considered close to ''Scaniornis'' anymore but rather united with the supposed "Cretaceous proto-flamingos" ''"Parascaniornis"'' and ''

Torotix ''Torotix'' is a Late Cretaceous genus of aquatic birds. They lived along the shores of the Western Interior Seaway, but it is not clear whether they were seabirds or freshwater birds, as the genus is only known from a humerus. Consequently, th ...

'', none of which seems even reasonably close to flamingos today.See also the discussion in Sibley ''et al.'' (1969).

Footnotes

References

* * {{Taxonbar, from=Q7430037 Prehistoric bird genera Paleocene birds Basal Neoaves Prehistoric birds of Europe Fossil taxa described in 1890