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The sauger (''Sander canadensis'') is a
freshwater Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. The term excludes seawater and brackish water, but it does include non-salty mi ...
perciform fish of the family Percidae that resembles its close relative, the
walleye The walleye (''Sander vitreus'', Synonym (taxonomy), synonym ''Stizostedion vitreum''), also called the walleyed pike, yellow pike, yellow pikeperch or yellow pickerel, is a freshwater perciform fish native to most of Canada and to the Northern ...
. The species is a member of the largest vertebrate order, the
Perciformes Perciformes (), also called the Acanthopteri, is an order or superorder of ray-finned fish in the clade Percomorpha. ''Perciformes'' means " perch-like". Among the well-known members of this group are perches and darters ( Percidae), and als ...
.Jaeger, Matthew. 2004. Montana's Fish Species of Special Concern: Sauger. Montana Cooperative Fisheries Research Unit. http://www.fisheriessociety.org/AFSmontana/SSCpages/Sauger%20Status.htm. (Accessed May 2011). It is the most migratory percid species in North America.Kuhn et al. 2008. Habitat Use and Movement Patterns by Adult Saugers from fall to Summer in an Unimpounded Small-River System. ''North American Journal of Fisheries Management''. 28: 360–367. American Fisheries Society. 2008. Saugers have two dorsal fins; the first is spiny and the posterior dorsal fin is soft-rayed. Their paired fins are in the thoracic position and their caudal fin is truncated, which means squared off at the corners, a characteristic of the family Percidae. Another physical characteristic of saugers is their ctenoid scales, which are common in advanced fishes. Saugers have a fusiform body structure, and as a result are well adapted predatory fishes and are capable of swimming into fast currents with minimal drag on their bodies. They may be distinguished from walleyes by the distinctly spotted
dorsal fin A dorsal fin is a fin on the back of most marine and freshwater vertebrates. Dorsal fins have evolved independently several times through convergent evolution adapting to marine environments, so the fins are not all homologous. They are found ...
, by the lack of a white splotch on the
caudal fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported only ...
, by the rough skin over their gills, and by their generally more brassy color, or darker (almost black) color in some regions. The typical sauger is in weight.


Distribution

Saugers are widely distributed; their historical range consisted of the eastern U.S. west of the
Appalachian Mountains The Appalachian Mountains, often called the Appalachians, are a mountain range in eastern to northeastern North America. The term "Appalachian" refers to several different regions associated with the mountain range, and its surrounding terrain ...
, mostly southern, central, and western U.S., and north into southern Canada. Sauger distribution and range has decreased from historical ranges because of degraded and fragmented habitat conditions. Sauger distribution within their home range varies by time of year because they are migratory. Saugers are more typical in rivers, whereas walleyes are more common in lakes and reservoirs. In many parts of their range, saugers are sympatric with walleyes. Hybridization between saugers and walleyes is not unknown; the hybrids, referred to as saugeyes, exhibit traits of both species. Being intermediate in appearance between the two species, saugeyes are sometimes difficult to differentiate, but they generally carry the dark blotches characteristic of the sauger. Saugers, however, are usually smaller and better tolerate waters of higher turbidity than walleyes. Saugers require warmer summer water temperatures of . Their need for warm water is thought to affect the northern and western boundaries of their range.


Reproduction and lifecycle

Saugers generally move upstream to spawn during March to May, depending on where they are. They move downstream to their home locations from April–July after their spawning period is over. Saugers have been known to travel between 10 and 600 km from their home to spawning locations downstream. Habitats at spawning sites are less complex and diverse than home locations. Females prefer rocky substrate and pools to deposit their eggs. As females increase in length, egg quality and fecundity increase, but egg production is thought to decline after age 6.Graeb, Brian D.S., Mark A. Kaemingk, David W. Willis. 2007. Early Life History of Sauger in Missouri River Reservoirs. Department of Wildlife and Fisheries Sciences. Brookings, SD. Sexual maturity is reached between 2 and 5 years old. Other measures of sexual maturity are related to size. A sauger is considered to be an adult when it reaches . Upon hatching, larval saugers drift downstream before developing feeding tendencies and horizontal maneuverability. Juvenile saugers tend to develop in diversion canals and backwaters until autumn, when they migrate upstream to their winter habitat. Residing in diversion canals is a large cause of mortality for juvenile saugers.


Diet

Saugers feed on a variety of invertebrates and small fishes depending on the time of year and size of the sauger.
Channel catfish The channel catfish (''Ictalurus punctatus''), known informally as the "channel cat", is a species of catfish native to North America. They are North America's most abundant catfish species, and the official state fish of Kansas, Missouri, Nebra ...
''Ictalurus punctatus'' and freshwater drum ''Aplodinotus grunniens'' are midsize (300–379 mm) and large (>379 mm) saugers' main food source during spring. The diet of a small sauger (200–299 mm) is slightly different from a larger sauger's. Smaller saugers generally feed on benthic invertebrates, mayfly larvae, and catfish during spring and summer. Midsized and large saugers feed mainly on fish from spring to autumn, but their diets alter during summer. Midsized and large saugers feed predominantly on mayfly larvae but only during summer. Freshwater drum ''Aplodinotus grunniens'' and gizzard shad ''Dorosoma cepedianum'' are predominant food sources for saugers of all sizes during autumn. Fish accounted for over 99% of a sauger's diet during autumn. Saugers also prey on shiners during spring and summer, but they do not account for a significant part of their diet. Shiners are absent from a sauger's diet in autumn probably due to their availability.Wickstrom, Gerald A. 2006. Seasonal Distribution, Movement, and Food Habits of Walleye and Sauger in Lewis and Clark Lake. South Dakota Department of Game, Fish & Parks. Chamberlain, South Dakota.


Habitat

Saugers are more likely to be found in large rivers with deep pools (depths greater than 0.6 m). They encounter a variety of habitats because of their migratory tendencies. They are usually found in natural rivers because they have more abundant pools and their flows have not been altered by dams or diversions. They are still common in impounded river systems. Diversions and dams affect habitat and spawning areas of saugers. They are usually found in areas with high turbidity, low channel slope, low stream velocity, and deep water. Saugers tend to select pools with sand and silt substrates, and habitat features that provide cover from the river current. They tend to avoid runs and riffles. They are most commonly found in pools that are at least 1.5 m deep. They can also be found in shallower pools, but in lesser numbers. No differences in habitat preference between males and females has been observed. The number of saugers observed increases with mean summer water temperature, maximum water depth, and alkalinity.


Taxonomy

Alongside the walleye, the sauger forms the North American
clade In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ...
within the genus ''
Sander A sander is a power tool used to smooth surfaces by abrasion with sandpaper. Sanders have a means to attach the sandpaper and a mechanism to move it rapidly contained within a housing with means to handhold it or fix it to a workbench. Woodw ...
'', and the sauger is thought to have first speciated into its modern form about 7.3 million years ago. The sauger was first formally described as ''Lucioperca canadensis'' in 1834 by
British British may refer to: Peoples, culture, and language * British people, nationals or natives of the United Kingdom, British Overseas Territories and Crown Dependencies. * British national identity, the characteristics of British people and culture ...
naturalist Natural history is a domain of inquiry involving organisms, including animals, fungi, and plants, in their natural environment, leaning more towards observational than experimental methods of study. A person who studies natural history is cal ...
s and explorers Edward Griffith (1790–1858) and Charles Hamilton Smith (1776–1859) based on the work of French naturalist and
anatomist Anatomy () is the branch of morphology concerned with the study of the internal structure of organisms and their parts. Anatomy is a branch of natural science that deals with the structural organization of living things. It is an old scien ...
Georges Cuvier Jean Léopold Nicolas Frédéric, baron Cuvier (23 August 1769 – 13 May 1832), known as Georges Cuvier (; ), was a French natural history, naturalist and zoology, zoologist, sometimes referred to as the "founding father of paleontology". Cuv ...
(1769–1832) which Griffith translated from the
French language French ( or ) is a Romance languages, Romance language of the Indo-European languages, Indo-European family. Like all other Romance languages, it descended from the Vulgar Latin of the Roman Empire. French evolved from Northern Old Gallo-R ...
.


Conservation

Saugers face many conservation issues because of migratory barriers,
habitat loss Habitat destruction (also termed habitat loss or habitat reduction) occurs when a natural habitat is no longer able to support its native species. The organisms once living there have either moved elsewhere, or are dead, leading to a decrease ...
, entrainment in
irrigation Irrigation (also referred to as watering of plants) is the practice of applying controlled amounts of water to land to help grow crops, landscape plants, and lawns. Irrigation has been a key aspect of agriculture for over 5,000 years and has bee ...
canals, and
overexploitation Overexploitation, also called overharvesting or ecological overshoot, refers to harvesting a renewable resource to the point of diminishing returns. Continued overexploitation can lead to the destruction of the resource, as it will be unable to ...
. Dams and diversion canals prevent
spawning Spawn is the Egg cell, eggs and Spermatozoa, sperm released or deposited into water by aquatic animals. As a verb, ''to spawn'' refers to the process of freely releasing eggs and sperm into a body of water (fresh or marine); the physical act is ...
in upstream habitats. Altering flows in rivers affect
turbidity Turbidity is the cloudiness or haziness of a fluid caused by large numbers of individual particles that are generally invisible to the naked eye, similar to smoke in air. The measurement of turbidity is a key test of both water clarity and wa ...
, formation of pools, and temperature, all of which are important for the timing and success of spawning saugers. Long migrations are the main reason why saugers struggle in dammed or diverted river systems. Low water levels in periods of drought are the most detrimental to sauger populations because it strands eggs during spawning and prevents larval saugers from reaching their downstream locations. High death rates that occur during spawning are related to degraded and fragmented river systems. Mortality rates in autumn are related to exploitation by fishermen. Sauger are critically imperiled in the state of
New York New York most commonly refers to: * New York (state), a state in the northeastern United States * New York City, the most populous city in the United States, located in the state of New York New York may also refer to: Places United Kingdom * ...
, with a small population still remaining in
Lake Champlain Lake Champlain ( ; , ) is a natural freshwater lake in North America. It mostly lies between the U.S. states of New York (state), New York and Vermont, but also extends north into the Canadian province of Quebec. The cities of Burlington, Ve ...
. Sauger were
extirpated Local extinction, also extirpation, is the termination of a species (or other taxon) in a chosen geographic area of study, though it still exists elsewhere. Local extinctions are contrasted with global extinctions. Local extinctions mark a chan ...
from
Lake Erie Lake Erie ( ) is the fourth-largest lake by surface area of the five Great Lakes in North America and the eleventh-largest globally. It is the southernmost, shallowest, and smallest by volume of the Great Lakes and also has the shortest avera ...
at some point after a population crash in the 1950s, as well as from
Lake Ontario Lake Ontario is one of the five Great Lakes of North America. It is bounded on the north, west, and southwest by the Canadian province of Ontario, and on the south and east by the U.S. state of New York (state), New York. The Canada–United Sta ...
. Populations in the
Allegheny River The Allegheny River ( ; ; ) is a tributary of the Ohio River that is located in western Pennsylvania and New York (state), New York in the United States. It runs from its headwaters just below the middle of Pennsylvania's northern border, nor ...
are prevented from reaching New York by the Kinzua Dam, which has led to the initiation of a sauger stocking program in the upper Allegheny River. As of 2022, fishing for sauger in New York is totally prohibited statewide.


References

*


External links


Fishbase information on Sauger
*Kansas Department of Wildlife and Park

*Montana Fish, Wildlife and Park

*Lake-Link.co

*Sauger Fishing in the Tennessee Valle
Saugeye
{{Authority control Sander (fish) Freshwater fish of the United States Fish of the Eastern United States Fish of the Western United States Fish of the Great Lakes Fish described in 1834 Taxa named by Edward Griffith (zoologist)