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''Sarracenia rubra'', also known as the sweet or purple pitcher plant, is a
carnivorous plant Carnivorous plants are plants that derive some or most of their nutrients from trapping and consuming animals or protozoans, typically insects and other arthropods, and occasionally small mammals and birds. They have adapted to grow in waterlo ...
in the
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
''
Sarracenia ''Sarracenia'' ( or ) is a genus comprising 8 to 11 species of North American pitcher plants, commonly called trumpet pitchers. The genus belongs to the family Sarraceniaceae, which also contain the closely allied genera '' Darlingtonia'' and '' ...
''. Like all ''Sarracenia'', it is native to the
New World The term "New World" is used to describe the majority of lands of Earth's Western Hemisphere, particularly the Americas, and sometimes Oceania."America." ''The Oxford Companion to the English Language'' (). McArthur, Tom, ed., 1992. New York: ...
. Its range extends from southern
Mississippi Mississippi ( ) is a U.S. state, state in the Southeastern United States, Southeastern and Deep South regions of the United States. It borders Tennessee to the north, Alabama to the east, the Gulf of Mexico to the south, Louisiana to the s ...
, through southern
Alabama Alabama ( ) is a U.S. state, state in the Southeastern United States, Southeastern and Deep South, Deep Southern regions of the United States. It borders Tennessee to the north, Georgia (U.S. state), Georgia to the east, Florida and the Gu ...
, the
Florida panhandle The Florida panhandle (also known as West Florida and Northwest Florida) is the northwestern part of the U.S. state of Florida. It is a Salient (geography), salient roughly long, bordered by Alabama on the west and north, Georgia (U.S. state ...
and
Georgia Georgia most commonly refers to: * Georgia (country), a country in the South Caucasus * Georgia (U.S. state), a state in the southeastern United States Georgia may also refer to: People and fictional characters * Georgia (name), a list of pe ...
, to the coastal plains of
North Carolina North Carolina ( ) is a U.S. state, state in the Southeastern United States, Southeastern region of the United States. It is bordered by Virginia to the north, the Atlantic Ocean to the east, South Carolina to the south, Georgia (U.S. stat ...
and
South Carolina South Carolina ( ) is a U.S. state, state in the Southeastern United States, Southeastern region of the United States. It borders North Carolina to the north and northeast, the Atlantic Ocean to the southeast, and Georgia (U.S. state), Georg ...
.


Morphology and carnivory

Like other members of the genus ''Sarracenia'', the sweet pitcherplant traps insects using a rolled leaf, which in this species is generally smaller and narrower than most species, usually not exceeding 65 cm (26 inches) in height. ''Sarracenia rubra'' is generally clump-forming. The uppermost part of the leaf is flared into a lid (the operculum), which prevents excess rain from entering the pitcher and diluting the digestive secretions within. The upper regions of the pitcher are covered in short, stiff, downwards-pointing
hairs Hair is a protein filament that grows from follicles found in the dermis. Hair is one of the defining characteristics of mammals. The human body, apart from areas of glabrous skin, is covered in follicles which produce thick terminal and fin ...
, which serve to guide
insect Insects (from Latin ') are Hexapoda, hexapod invertebrates of the class (biology), class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (Insect morphology#Head, head, ...
s alighting on the upper portions of the leaf towards the opening of the pitcher tube. The opening of the pitcher tube is retroflexed into a 'nectar roll' or
peristome Peristome (from the Greek language, Greek ''peri'', meaning 'around' or 'about', and ''stoma'', 'mouth') is an anatomical feature that surrounds an opening to an organ or structure. Some plants, fungi, and shelled gastropods have peristomes. In mo ...
, whose surface is studded with
nectar Nectar is a viscous, sugar-rich liquid produced by Plant, plants in glands called nectaries, either within the flowers with which it attracts pollination, pollinating animals, or by extrafloral nectaries, which provide a nutrient source to an ...
-secreting glands. Prey entering the tube find that their footing is made extremely uncertain by the smooth, waxy secretions found on the surfaces of the upper portion of the tube. Insects losing their footing on this surface plummet to the bottom of the tube, where a combination of digestive fluid, wetting agents and inward-pointing hairs prevent their escape. Some large insects (such as
wasp A wasp is any insect of the narrow-waisted suborder Apocrita of the order Hymenoptera which is neither a bee nor an ant; this excludes the broad-waisted sawflies (Symphyta), which look somewhat like wasps, but are in a separate suborder ...
s) have been reported to escape from the pitchers on occasion, by chewing their way out through the wall of the tube.Flora of North America, ''Sarracenia rubra'' Walter, 1788. Sweet or red pitcher plant
/ref>


Flowers

''See also the section on ''Sarracenia'' flowers in the main article''. In spring, the plant produces small, bright red
flower Flowers, also known as blooms and blossoms, are the reproductive structures of flowering plants ( angiosperms). Typically, they are structured in four circular levels, called whorls, around the end of a stalk. These whorls include: calyx, m ...
s with 5-fold
symmetry Symmetry () in everyday life refers to a sense of harmonious and beautiful proportion and balance. In mathematics, the term has a more precise definition and is usually used to refer to an object that is Invariant (mathematics), invariant und ...
. The red
petal Petals are modified leaves that form an inner whorl surrounding the reproductive parts of flowers. They are often brightly coloured or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''corol ...
s are long and strap-like, and dangle over the
umbrella An umbrella or parasol is a folding canopy supported by wooden or metal ribs that is mounted on a wooden, metal, or plastic pole. It is usually designed to protect a person against rain. The term ''umbrella'' is traditionally used when protec ...
-like style of the flower, which is held upside down at the end of a 50 cm long scape. The
stigmata Stigmata (, plural of , 'mark, spot, brand'), in Roman Catholicism, Catholicism, are bodily wounds, scars and pain which appear in locations corresponding to the Crucifixion of Jesus, crucifixion Five Holy Wounds, wounds of Jesus in Christian ...
of the flower are found at the tips of the 'spokes' of this umbrella. Pollinating insects generally enter the flower from above, forcing their way into the cavity between the petals and umbrella, and depositing any pollen they are carrying on the stigmata as they enter. The pollinators generally exit the flower, having been dusted with the plant's own pollen, by lifting a petal. This one-way system helps to ensure cross pollination. ''Sarracenia rubra'' flowers are generally sweet-scented, hence the common name.


Growth cycle

This species is characterised by producing quite floppy pitchers in spring with large wings, perhaps as a method of producing a large surface area of tissue in order to rapidly photosynthesise at the start of the year. Later on in summer and autumn, much more substantial pitchers are produced.


Taxonomy

''Sarracenia rubra'' taxonomy is rather complicated and still much debated. Currently, there are six subspecies: *''S. rubra'' subsp. ''rubra'' *''S. rubra'' subsp. ''jonesii'' ''S. jonesii''*''S. rubra'' subsp. ''gulfensis'' *''S. rubra'' subsp. ''viatorum'' *''S. rubra'' subsp. ''alabamensis'' ''S. alabamensis''*''S. rubra'' subsp. ''wherryi'' ''S. alabamensis'' subsp. ''wherryi'' These placings are somewhat disputed, however, as each could arguably be its own species due to differences in pitcher morphology. The similarity between the flowers of these taxa indicates a close relationship, however.


References

{{Taxonbar, from=Q159647 rubra Carnivorous plants of North America Endemic flora of the United States Flora of the Southeastern United States Plants described in 1788 Flora without expected TNC conservation status