The Sardinian warbler (''Curruca melanocephala'') is a common and widespread

typical warbler The typical warblers are small birds belonging to the genus ''Sylvia'' in the "Old World warbler" (or sylviid warbler) family Sylviidae.Del Hoyo, J.; Elliot, A., & Christie, D. (editors). (2006). ''Handbook of the Birds of the World''. Volume 1 ...

from the

Mediterranean The Mediterranean Sea ( ) is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean basin and almost completely enclosed by land: on the east by the Levant in West Asia, on the north by Anatolia in West Asia and Southern ...

region. Like most ''Curruca''

species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...

, it has distinct male and female

plumage Plumage () is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, there can b ...

s. The adult male has a grey back, whitish underparts, black head, white throat and red eyes. Plumages are somewhat variable even in the same locality, with the intensity of a reddish hue on upper- and/or underside that varies from absent to (in some

subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...

) pronounced. The female is mainly brown above and buff below, with a grey head. The Sardinian warbler's song is fast and rattling, and is very characteristic of the

areas where this bird breeds.

Taxonomy and systematics

The Sardinian warbler was

formally described A species description is a formal scientific description of a newly encountered species, typically articulated through a scientific publication. Its purpose is to provide a clear description of a new species of organism and explain how it differ ...

in 1789 by the German naturalist

Johann Friedrich Gmelin Johann Friedrich Gmelin (8 August 1748 – 1 November 1804) was a German natural history, naturalist, chemist, botanist, entomologist, herpetologist, and malacologist. Education Johann Friedrich Gmelin was born as the eldest son of Philipp F ...

in his revised and expanded edition of

Carl Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...

's ''

Systema Naturae ' (originally in Latin written ' with the Orthographic ligature, ligature æ) is one of the major works of the Sweden, Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the syste ...

''. He placed it with the wagtails in the

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

'' Motacilla'' and coined the

binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin grammatical forms, altho ...

''Motacilla melanocephala''. Gmelin based his entry on a description by the Italian zoologist

Francesco Cetti Francesco Cetti (9 August 1726 – 20 November 1778) was an Italian Jesuit priest, zoologist and mathematician. Biography Cetti was born in Mannheim in Germany, but his parents were natives of Como. He was educated in Lombardy and at the Jesu ...

in his book ''Gli uccelli di Sardegna'' (''The Birds of Sardinia'') that was published in 1776. This was the second volume of his ''Storia naturale di Sardegna''. Cetti did not use scientific names for the species. When a translation of Cetti's three volumes were published in German, the translator, David Piesch, and the editor, Nathanael Gottfried Leske, included an Appendix at the end of the third volume (published in 1784) in which they proposed the binomial name ''Motacilla melanocephala'', the identical name that was later adopted by Gmelin. In spite of this earlier publication, as of 2023 Gmelin is treated as the authority for the species. The species was formerly placed in the genus ''Sylvia'' that was introduced in 1769 by the Italian naturalist

Giovanni Antonio Scopoli Giovanni Antonio Scopoli (sometimes Latinisation of names, Latinized as Johannes Antonius Scopolius) (3 June 1723 – 8 May 1788) was an Italians, Italian physician and natural history, naturalist. His biographer Otto Guglia named him the "first ...

. The genus name is from

Modern Latin Neo-LatinSidwell, Keith ''Classical Latin-Medieval Latin-Neo Latin'' in ; others, throughout. (also known as New Latin and Modern Latin) is the style of written Latin used in original literary, scholarly, and scientific works, first in Italy d ...

''silvia'', a woodland sprite, related to ''silva'', a wood. The specific ''melanocephala'' is from

Ancient Greek Ancient Greek (, ; ) includes the forms of the Greek language used in ancient Greece and the classical antiquity, ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Greek ...

''melas'', "black", and ''kephale'', "head". Currently, the Sardinian warbler is placed in the genus ''

Curruca ''Curruca'' is a genus of Sylviid warblers, best represented in Europe, Africa, and Asia. All of these species were formerly placed in the genus '' Sylvia''. Taxonomy The genus ''Curruca'' was introduced by the German naturalist Johann Matthäus ...

'' by the IOC, along with most of the species formerly classified in the genus ''Sylvia''. Together with Menetries's warbler the Sardinian warbler forms a

superspecies In biology, a species complex is a group of closely related organisms that are so similar in appearance and other features that the boundaries between them are often unclear. The taxa in the complex may be able to hybridize readily with each oth ...

. Both have white malar areas and light throats, and otherwise black heads in adult males, as well as a naked ring around the eye. The

eastern subalpine warbler The eastern subalpine warbler (''Curruca cantillans'') is a small typical warbler which breeds in the southernmost areas of Europe. It was Species description, first described by the German naturalist Peter Simon Pallas in 1764 and given the bino ...

, which seems the superspecies' closest relative, has a dark throat and breast and a dark gray upper head in males, but otherwise shares these characters. These three species are related to a dark-throated superspecies consisting of Rüppell's warbler and the Cyprus warbler, which also share the white malar area with blackish above.Helbig (2001), Jønsson & Fjeldså (2006) This bird may be considered a

, divided into the western ''Curruca melanocephala'' and ''Curruca momus'' from the more arid regions of the

Near East The Near East () is a transcontinental region around the Eastern Mediterranean encompassing the historical Fertile Crescent, the Levant, Anatolia, Egypt, Mesopotamia, and coastal areas of the Arabian Peninsula. The term was invented in the 20th ...

and adjacent Africa.

Subspecies

The geographical variation in the Sardinian warbler conforms to some extent with

Gloger's rule Gloger's rule is an ecogeographical rule which states that within a species of endotherms, more heavily pigmented forms tend to be found in more humid environments, e.g. near the equator. It was named after the zoologist Constantin Wilhelm Lamb ...

, though not as strongly as in some other typical warblers. The validity of ''leucogastra'' and ''norissae'' is not accepted by some authors, and ''valverdei'' has been described very recently. On the other hand, ''leucogastra'' might be more than one subspecies.Cabot & Urdiales (2005) * ''Curruca melanocephala melanocephala'' (Gmelin, 1789) :

Iberia The Iberian Peninsula ( ), also known as Iberia, is a peninsula in south-western Europe. Mostly separated from the rest of the European landmass by the Pyrenees, it includes the territories of peninsular Spain and Continental Portugal, compri ...

across the northern

to western

Turkey Turkey, officially the Republic of Türkiye, is a country mainly located in Anatolia in West Asia, with a relatively small part called East Thrace in Southeast Europe. It borders the Black Sea to the north; Georgia (country), Georgia, Armen ...

. Extends into the

Maghreb The Maghreb (; ), also known as the Arab Maghreb () and Northwest Africa, is the western part of the Arab world. The region comprises western and central North Africa, including Algeria, Libya, Mauritania, Morocco, and Tunisia. The Maghreb al ...

from Iberia, and into

Libya Libya, officially the State of Libya, is a country in the Maghreb region of North Africa. It borders the Mediterranean Sea to the north, Egypt to Egypt–Libya border, the east, Sudan to Libya–Sudan border, the southeast, Chad to Chad–L ...

from

Italy Italy, officially the Italian Republic, is a country in Southern Europe, Southern and Western Europe, Western Europe. It consists of Italian Peninsula, a peninsula that extends into the Mediterranean Sea, with the Alps on its northern land b ...

via

Sicily Sicily (Italian language, Italian and ), officially the Sicilian Region (), is an island in the central Mediterranean Sea, south of the Italian Peninsula in continental Europe and is one of the 20 regions of Italy, regions of Italy. With 4. ...

. Migrates to the

Sahel The Sahel region (; ), or Sahelian acacia savanna, is a Biogeography, biogeographical region in Africa. It is the Ecotone, transition zone between the more humid Sudanian savannas to its south and the drier Sahara to the north. The Sahel has a ...

and

oases In ecology, an oasis (; : oases ) is a fertile area of a desert or semi-desert environment phylogenetic In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical dat ...

status requires review :

Canary Islands The Canary Islands (; ) or Canaries are an archipelago in the Atlantic Ocean and the southernmost Autonomous communities of Spain, Autonomous Community of Spain. They are located in the northwest of Africa, with the closest point to the cont ...

, resident, probably some vagrancy between eastern islands and Maghreb. :Medium size, short-winged and large-billed.

Tenerife Tenerife ( ; ; formerly spelled ''Teneriffe'') is the largest and most populous island of the Canary Islands, an Autonomous communities of Spain, autonomous community of Spain. With a land area of and a population of 965,575 inhabitants as of A ...

and

La Palma La Palma (, ), also known as ''La isla bonita'' () and historically San Miguel de La Palma, is the most northwesterly island of the Canary Islands, a Spanish autonomous community and archipelago in Macaronesia in the North Atlantic Ocean. La Pa ...

(western) birds are most distinct, being dark above with some rusty/beige hue on the underside in males. Eastern birds (

Fuerteventura Fuerteventura () is one of the Canary Islands, in the Atlantic Ocean, geographically part of Macaronesia, and politically part of Spain. It is located away from the coast of North Africa. The island was declared a biosphere reserve by UNESCO i ...

Lanzarote Lanzarote (, , ) is a Spanish island, the easternmost of the Canary Islands, off the north coast of Africa and from the Iberian Peninsula. Covering , Lanzarote is the fourth-largest of the islands in the archipelago. With 163,230 inhabi ...

and

Gran Canaria Gran Canaria (, ; ), also Grand Canary Island, is the third-largest and second-most-populous island of the Canary Islands, a Spain, Spanish archipelago off the Atlantic coast of Northwest Africa. the island had a population of that constitut ...

) are more like ''melanocephala'' and ''momus'' but differ in measurements. * ''Curruca melanocephala momus'' (Hemprich & Ehrenberg, 1833) :

. Resident, some local movements. :Smallish, short-winged. Varies between brownish grey and rusty above; underside almost always has reddish hue. Females rusty to rusty olive. * ''Curruca melanocephala norissae'' Nicoll, 1917: Fayyum warbler - probably only a local morph of ''momus'' :

Nile Delta The Nile Delta (, or simply , ) is the River delta, delta formed in Lower Egypt where the Nile River spreads out and drains into the Mediterranean Sea. It is one of the world's larger deltas—from Alexandria in the west to Port Said in the eas ...

region.

Extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...

since around 1940. :Like ''momus'', but tend to be very reddish. * ''Curruca melanocephala valverdei'' Cabot & Urdiales, 2005 - recently split from ''melanocephala'' :From

Tiznit Tiznit or Tiznet (, ) is a town in the west coast of the Moroccan region of Souss-Massa, founded in 1881 by Alawi Sultan Hassan I. It is the capital of Tiznit Province and recorded a population of 74,699 in the 2014 Moroccan census. Internati ...

(

Morocco Morocco, officially the Kingdom of Morocco, is a country in the Maghreb region of North Africa. It has coastlines on the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to Algeria–Morocc ...

) south to the

Tropic of Cancer The Tropic of Cancer, also known as the Northern Tropic, is the Earth's northernmost circle of latitude where the Sun can be seen directly overhead. This occurs on the June solstice, when the Northern Hemisphere is tilted toward the Sun ...

, inland to the edge of the

Sahara The Sahara (, ) is a desert spanning across North Africa. With an area of , it is the largest hot desert in the world and the list of deserts by area, third-largest desert overall, smaller only than the deserts of Antarctica and the northern Ar ...

. Resident, but some seasonal movements. :Medium-sized, tail tip quite square. A very pure-colored form, the palest subspecies. Undersides clean white. Matte black cap in males. Juveniles decidedly sandy.

Distribution and habitat

It breeds in the southernmost areas of

Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...

and just into

Asia Asia ( , ) is the largest continent in the world by both land area and population. It covers an area of more than 44 million square kilometres, about 30% of Earth's total land area and 8% of Earth's total surface area. The continent, which ...

in Turkey and the eastern end of the Mediterranean. This small

passerine A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped') which includes more than half of all bird species. Sometimes known as perching birds, passerines generally have an anisodactyl arrangement of their ...

bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...

, unlike most "

warbler Various Passeriformes (perching birds) are commonly referred to as warblers. They are not necessarily closely related to one another, but share some characteristics, such as being fairly small, vocal, and insectivorous. Sylvioid warblers T ...

s", is not particularly migratory, but some birds winter in

North Africa North Africa (sometimes Northern Africa) is a region encompassing the northern portion of the African continent. There is no singularly accepted scope for the region. However, it is sometimes defined as stretching from the Atlantic shores of t ...

, and it occurs as a vagrant well away from the breeding range, as far as

Great Britain Great Britain is an island in the North Atlantic Ocean off the north-west coast of continental Europe, consisting of the countries England, Scotland, and Wales. With an area of , it is the largest of the British Isles, the List of European ...

Behaviour and ecology

This is a

of open country and cultivation, with bushes for nesting. The nest is built in low shrub or brambles, and 3-6 eggs are laid. Like most "

s", it is

insectivorous A robber fly eating a hoverfly An insectivore is a carnivorous animal or plant which eats insects. An alternative term is entomophage, which can also refer to the human practice of eating insects. The first vertebrate insectivores we ...

, but will also take berries and other soft

fruit In botany, a fruit is the seed-bearing structure in flowering plants (angiosperms) that is formed from the ovary after flowering. Fruits are the means by which angiosperms disseminate their seeds. Edible fruits in particular have long propaga ...

References

* * Helbig, A.J. (2001): Phylogeny and biogeography of the genus ''Sylvia''. ''In:'' Shirihai, Hadoram: ''Sylvia warblers'': 24–29. Princeton University Press, Princeton, N.J.. * Jønsson, Knud A. & Fjeldså, Jon (2006): A phylogenetic supertree of oscine passerine birds (Aves: Passeri). '' Zool. Scripta'' 35(2): 149–186. (HTML abstract)

External links

Xeno-canto: audio recordings of the Sardinian warblerAgeing and sexing (PDF; 4.2 MB) by Javier Blasco-Zumeta & Gerd-Michael Heinze

Avibase
{{Taxonbar, from=Q110257507, from2=Q669647

Sardinian warbler The Sardinian warbler (''Curruca melanocephala'') is a common and widespread typical warbler from the Mediterranean region. Like most ''Curruca'' species, it has distinct male and female plumages. The adult male has a grey back, whitish underpart ...

Birds of Southern Europe Birds of West Asia Birds of North Africa Birds of the Canary Islands Fauna of Sardinia