''Santalum ellipticum'', commonly known as ''Iliahialoe'' ( Hawaiian) or coastal sandalwood, is a species of

flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Ancient Greek, Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed with ...

in the

mistletoe Mistletoe is the common name for obligate parasite, obligate parasitic plant, hemiparasitic plants in the Order (biology), order Santalales. They are attached to their host tree or shrub by a structure called the haustorium, through which they ...

family,

Santalaceae The Santalaceae, sandalwoods, are a widely distributed family of flowering plants (including small trees, shrubs, perennial plants, perennial herbs, and epiphyte, epiphytic climbersHewson & George t al.'Santalaceae'' taxonomy, 1984, pp. 191-1 ...

, that is

endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...

to the

Hawaiian Islands The Hawaiian Islands () are an archipelago of eight major volcanic islands, several atolls, and numerous smaller islets in the Pacific Ocean, North Pacific Ocean, extending some from the Hawaii (island), island of Hawaii in the south to nort ...

. It is a sprawling

shrub A shrub or bush is a small to medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees by their multiple ...

to small

tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, e.g., including only woody plants with secondary growth, only ...

, typically reaching a height of and a canopy spread of , but is extremely variable in size and shape. Like other members of the genus, ''S. ellipticum'' is a hemi-parasite, deriving some of its

nutrients A nutrient is a substance used by an organism to survive, grow and reproduce. The requirement for dietary nutrient intake applies to animals, plants, fungi and protists. Nutrients can be incorporated into cells for metabolic purposes or excret ...

from the host plant by attaching to its roots.

Habitat and range

''Iliahialoe'' inhabits

dry forests The tropical and subtropical dry broadleaf forest is a habitat type defined by the World Wide Fund for Nature and is located at tropical and subtropical latitudes. Though these forests occur in climates that are warm year-round, and may receive ...

, low shrublands, and lava plains throughout the archipelago, including the

Northwestern Hawaiian Islands The Northwestern Hawaiian Islands also known as the Leeward Hawaiian Islands, are a series of islands and atolls located northwest of Kauai and Niihau, Niihau in the Hawaiian Islands, Hawaiian island chain. Politically, these islands are part of ...

, but has been

extirpated Local extinction, also extirpation, is the termination of a species (or other taxon) in a chosen geographic area of study, though it still exists elsewhere. Local extinctions are contrasted with global extinctions. Local extinctions mark a chan ...

from Laysan and Kahoolawe. Although never recorded on

Niihau Niihau (Hawaiian language, Hawaiian: ), anglicized as Niihau ( ), is the seventh largest island in Hawaii and the westernmost of the main islands. It is southwest of Kauai, Kauai across the Channels of the Hawaiian Islands#Kaulakahi Channel, Ka ...

, its historic presence on the island is almost certain. ''S. ellipticum'' is generally found at elevations from sea level to , but populations can occur as high as . An isolated individual was observed growing at on the island of Hawaii.

Uses

Non-medicinal

The ''laau ala'' (

heartwood Wood is a structural tissue/material found as xylem in the stems and roots of trees and other woody plants. It is an organic materiala natural composite of cellulosic fibers that are strong in tension and embedded in a matrix of lignin t ...

) of ''iliahialoe'' contains valuable, aromatic

essential oils An essential oil is a concentrated hydrophobic liquid containing volatile (easily evaporated at normal temperatures) chemical compounds from plants. Essential oils are also known as volatile oils, ethereal oils, aetheroleum, or simply as the o ...

. Trees were harvested for export to

China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...

between 1791 and 1840, where the hard, yellowish-brown

wood Wood is a structural tissue/material found as xylem in the stems and roots of trees and other woody plants. It is an organic materiala natural composite of cellulosic fibers that are strong in tension and embedded in a matrix of lignin t ...

was made into carved objects, chests, and

incense Incense is an aromatic biotic material that releases fragrant smoke when burnt. The term is used for either the material or the aroma. Incense is used for aesthetic reasons, religious worship, aromatherapy, meditation, and ceremonial reasons. It ...

. The ''iliahialoe'' trade peaked from 1815 to 1826.

Native Hawaiian Native Hawaiians (also known as Indigenous Hawaiians, Kānaka Maoli, Aboriginal Hawaiians, or simply Hawaiians; , , , and ) are the Indigenous peoples of Oceania, Indigenous Polynesians, Polynesian people of the Hawaiian Islands. Hawaiʻi was set ...

s used the wood to make ''pola'', the deck on a ''waa kaulua'' ( double-hulled canoe). Powdered ''laau ala'' was used as a

perfume Perfume (, ) is a mixture of fragrance, fragrant essential oils or aroma compounds (fragrances), Fixative (perfumery), fixatives and solvents, usually in liquid form, used to give the human body, animals, food, objects, and living-spaces an agre ...

and added to '' kapa'' cloth.

Medicinal

Native Hawaiians combined leaves and bark of the ''iliahialoe'' with ''naio'' (''

Myoporum sandwicense ''Myoporum sandwicense'', commonly known as ''naio'', bastard sandalwood or false sandalwood is a species of flowering plant in the figwort family, Scrophulariaceae. It is a tree or shrub highly variable in its form, the size and shape of its le ...

'') ashes to treat ''kepia o ke poo'' (

dandruff Dandruff is a skin condition of the scalp. Symptoms include flaking and sometimes mild itchiness. It can result in social or self-esteem problems. A more severe form of the condition, which includes inflammation of the skin, is known as s ...

) and ''liha o ka lauoho'' (

head lice The head louse (''Pediculus humanus capitis'') is an obligate ectoparasite of humans. Head lice are wingless insects that spend their entire lives on the human scalp and feed exclusively on human blood. Humans are the only known hosts of this s ...

). ''Iliahialoe'' shavings mixed with ''awa'' ('' Piper methysticum''), ''nioi'' ('' Eugenia reinwardtiana''), ''ahakea'' ('' Bobea'' spp.), and ''kauila'' ('' Alphitonia ponderosa'') was used to treat

sexually transmitted disease A sexually transmitted infection (STI), also referred to as a sexually transmitted disease (STD) and the older term venereal disease (VD), is an infection that is spread by sexual activity, especially vaginal intercourse, anal sex, or ...

References

ellipticum Plants described in 1829 Trees of Hawaii Endemic flora of Hawaii Flora without expected TNC conservation status {{Santalales-stub