
Sand sharks, also known as sand tiger sharks, gray nurse sharks or ragged tooth sharks, are
mackerel shark
The Lamniformes (, from Greek ''lamna'' "fish of prey") are an order of sharks commonly known as mackerel sharks (which may also refer specifically to the family Lamnidae). It includes some of the most familiar species of sharks, such as the gr ...
s of the
family
Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Odontaspididae. They are found worldwide in temperate and tropical waters. The three species are in two genera.
Description

The body tends to be brown with dark markings in the upper half. These markings disappear as they mature. Their needle-like teeth are highly adapted for impaling fish, their main prey. Their teeth are long, narrow, and very sharp with smooth edges, with one and on occasion two smaller cusplets on either side. Sand sharks have a large second
dorsal fin
A dorsal fin is a fin located on the back of most marine and freshwater vertebrates within various taxa of the animal kingdom. Many species of animals possessing dorsal fins are not particularly closely related to each other, though through c ...
.
The sand shark can grow up to long, and most adults can weigh around . The average lifespan of both sexes is only about 7 years, though they may live longer in captivity.
Location and origins
The name sand shark comes from their tendency to migrate towards shoreline habitats, and they are often seen swimming around the ocean floor in the surf zone; at times, they come very close to shore. They are often found in warm or temperate waters throughout the world's oceans, except the eastern
Pacific
The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the continen ...
. They also frequent the
Mediterranean
The Mediterranean Sea is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean Basin and almost completely enclosed by land: on the north by Western and Southern Europe and Anatolia, on the south by North Africa, and on th ...
and
Adriatic Sea
The Adriatic Sea () is a body of water separating the Italian Peninsula from the Balkans, Balkan Peninsula. The Adriatic is the northernmost arm of the Mediterranean Sea, extending from the Strait of Otranto (where it connects to the Ionian Sea) ...
s at depths from and sometimes more.
Behavior
The sand shark has a unique hunting strategy. It is able to gulp air from above the surface and collect the air in its
stomach
The stomach is a muscular, hollow organ in the gastrointestinal tract of humans and many other animals, including several invertebrates. The stomach has a dilated structure and functions as a vital organ in the digestive system. The stomach i ...
. This enables it to become
buoyant
Buoyancy (), or upthrust, is an upward force exerted by a fluid that opposes the weight of a partially or fully immersed object. In a column of fluid, pressure increases with depth as a result of the weight of the overlying fluid. Thus the pr ...
and approach its
prey
Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill the ...
virtually motionless. During the day, the sand shark stays mostly inactive, but at night, it becomes active and resumes hunting activities. Its staple is small fish, but it eats crustaceans and
squid
True squid are molluscs with an elongated soft body, large eyes, eight arms, and two tentacles in the superorder Decapodiformes, though many other molluscs within the broader Neocoleoidea are also called squid despite not strictly fitting ...
, as well. It occasionally hunts in shivers (groups), and has even been known to attack full
fishing nets
A fishing net is a Net (device), net used for fishing. Nets are devices made from fibers woven in a grid-like structure. Some fishing nets are also called fish traps, for example #Fyke nets, fyke nets. Fishing nets are usually meshes formed by ...
.
Reproduction
Sand sharks only develop two
embryo
An embryo is an initial stage of development of a multicellular organism. In organisms that reproduce sexually, embryonic development is the part of the life cycle that begins just after fertilization of the female egg cell by the male sperm ...
s, one in each uterus. The largest and strongest embryos consume their siblings in the womb (
intrauterine cannibalism
Cannibalism is the act of consuming another individual of the same species as food. Cannibalism is a common ecological interaction in the animal kingdom and has been recorded in more than 1,500 species. Human cannibalism is well documented, bo ...
) before each surviving pup is born. It has one of the lowest reproduction rates of all sharks and is susceptible to even minimal
population pressure
Population pressure, a term summarizing the stress brought about by an excessive population density and its consequences, is used both in conjunction with human overpopulation and with other animal populations that suffer from too many individuals ...
, so it is listed as
vulnerable
Vulnerable may refer to:
General
* Vulnerability
* Vulnerability (computing)
* Vulnerable adult
* Vulnerable species
Music
Albums
* ''Vulnerable'' (Marvin Gaye album), 1997
* ''Vulnerable'' (Tricky album), 2003
* ''Vulnerable'' (The Used album) ...
and is protected in much of its range.
Attacks on people
Sand sharks are not known to attack humans. If a person were to provoke a sand shark, it may retaliate defensively. Sand sharks are generally not aggressive, but harass divers who are spearfishing. In North America, wreck divers regularly visit the
World War II
World War II or the Second World War, often abbreviated as WWII or WW2, was a world war that lasted from 1939 to 1945. It involved the World War II by country, vast majority of the world's countries—including all of the great power ...
shipwrecks
A shipwreck is the wreckage of a ship that is located either beached on land or sunken to the bottom of a body of water. Shipwrecking may be intentional or unintentional. Angela Croome reported in January 1999 that there were approximately ...
to dive with the sharks that make the wrecks their home.
Conservation
A recent report from the PEW Charitable Trusts suggests a new management approach used for large mammals that have suffered population declines could hold promise for sharks. Because of the life-history characteristics of sharks, conventional fisheries management approaches, such as reaching maximum
sustainable yield
The sustainable yield of natural capital is the ecological yield that can be extracted without reducing the base of capital itself, i.e. the surplus required to maintain ecosystem services at the same or increasing level over time. The term onl ...
, may not be sufficient to rebuild depleted shark populations. Some of the more stringent approaches used to reverse declines in large mammals may be appropriate for sharks, including prohibitions on the retention of the most vulnerable species and regulation of international trade.
Species

The family contains three extant species, in two genera, as well as many extinct species in several genera. Recent
mitochondrial DNA analysis of extant members has found the two extant members do not actually form a
monophyletic
In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic ...
clade. This family is therefore
polyphyletic
A polyphyletic group is an assemblage of organisms or other evolving elements that is of mixed evolutionary origin. The term is often applied to groups that share similar features known as homoplasies, which are explained as a result of conver ...
and in need of revision.
* Genus ''
Carcharias
''Carcharias'' is a genus of sand tiger sharks belonging to the family Odontaspididae. Once bearing many prehistoric species, all have gone extinct with the exception of the critically endangered sand tiger shark.
Description
''Carcharias'' ar ...
''
Rafinesque
Constantine Samuel Rafinesque-Schmaltz (; October 22, 1783September 18, 1840) was a French 19th-century polymath born near Constantinople in the Ottoman Empire and self-educated in France. He traveled as a young man in the United States, ultimat ...
, 1810
** ''
Carcharias taurus
The sand tiger shark (''Carcharias taurus''), gray nurse shark, spotted ragged-tooth shark or blue-nurse sand tiger, is a species of shark that inhabits subtropical and temperate waters worldwide. It inhabits the continental shelf, from sandy sho ...
''
Rafinesque, 1810 (sand tiger shark)
*Genus ''
Odontaspis
''Odontaspis'' (from el, ὀδούς 'tooth') and el, ἀσπίς 'shield') is a genus of sand shark with two extant species.
Description
Bigeye sand tigers can reach a length of about and smalltooth sand tigers of about 4.1 m.
They ...
''
Agassiz 1838
** ''
Odontaspis ferox''
A. Risso, 1810 (smalltooth sand tiger)
** ''
Odontaspis noronhai''
Maul
A maul may refer to any number of large hammers, including:
* War hammer, a medieval weapon
* Post maul, a type of sledgehammer
* Spike maul, railroad hand tool
* Splitting maul, heavy wood-splitting tool resembling both axe and hammer
People
* A ...
, 1955 (bigeye sand tiger)
* Subfamily
Odontaspinae †
Herman, 1975
**Genus ''
Striatolamia
''Striatolamia'' is an extinct genus of sharks belonging to the family Odontaspididae. These extinct sharks lived from the Early Paleocene to Late Miocene (61.7 to 10.3 Ma).
Etymology
The Latin genus name ''Striatolamia'' refers to the striati ...
'' †
Glikman, 1964
**Genus ''
Carcharoides
''Carcharoides'' is an extinct genus of mackerel shark which lived during the Oligocene and Miocene epochs. It is a widespread genus, known from specimens in North and South America, Europe, Africa, and Australia. It is most common in the Europe ...
'' †
Ameghino, 1901
**Genus ''
Parodontaspis'' †
White, 1931
**Genus ''
Priodontaspis'' †
Ameghino, 1901
**Genus ''
Pseudoisurus'' †
Glikman, 1957
**Genus ''
Synodontaspis'' †
White, 1931
*Subfamily
Johnlonginae
Johnlonginae is an extinct subfamily of sand shark with two genera: ''Johnlongia'' and ''Pseudomegachasma''. The latter genus is thought to have evolved from former. They are among the older sand sharks, and are only known from the Cretaceous
T ...
†
Shimada, 2015
**Genus ''
Johnlongia
''Johnlongia'' is an extinct genus of sand shark from the Cretaceous period. It contains two described species, ''J. parvidens'' and ''J. allocotodon'', and possibly a third unnamed species from the Niobrara Chalk. It is presumed piscivorous; how ...
'' †
**Genus ''
Pseudomegachasma
''Pseudomegachasma'' ("false megamouth") is an extinct genus of filter-feeding shark that was closely related to the modern sand tiger shark. It is known from Cretaceous strata in Russia and the United States, and is the only known planktivorous ...
'' † (
Shimada, 2015)
References
External links
FishBase Family Odontaspididae - Sand tigers fishbase.org
Sand Tiger Shark SchoolSand Tigers Sharks Fayetteville Observer
{{Taxonbar, from=Q259297
Ovoviviparous fish
Extant Late Cretaceous first appearances
Taxa named by Friedrich Gustav Jakob Henle