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The Erycinae, known as the Old World sand boas, are a
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...
of
nonvenomous Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a sti ...
snakes Snakes are elongated, limbless, carnivorous reptiles of the suborder Serpentes . Like all other squamates, snakes are ectothermic, amniote vertebrates covered in overlapping scales. Many species of snakes have skulls with several more joi ...
in the family Boidae. Species of the subfamily Erycinae are found in Europe, Asia Minor, Africa, Arabia, central and southwestern Asia, India, Sri Lanka, and western North America. Four
genera Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomenclat ...
comprising 18 species are currently recognized as being valid.


Description

Erycinae is a subfamily of stout-bodied snakes, all of which are competent burrowers. The largest, '' E. johnii'', rarely exceeds in total length (including tail). Most grow to around in total length. They have small eyes and hard, small scales to protect their skin from the grit of sand. A great deal of sexual dimorphism exists, with females generally becoming much larger than males. Erycines have skeletal adaptations to burrowing. The skull is more compact than in the subfamily Boinae. Also, the vertebrae of the tail are increased in size but reduced in number.


Distribution and habitat

Erycines are found in Southeastern Europe, Asia Minor, North, Central, West and East Africa, Arabia, Central and Southwest Asia, India, Sri Lanka, southwestern Canada, the western United States, and northwestern Mexico. McDiarmid RW, Campbell JA, Touré T (1999). ''Snake Species of the World: A Taxonomic and Geographic Reference, Volume 1''. Washington, District of Columbia: Herpetologists' League. 511 pp. (series). (volume). Fossil erycines have been found in rock strata over 50 million years old, and were once widespread in
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
. Now, only four species remain in North America, as well as the sand boas in Africa, Asia, and Southeastern Europe. The oldest known erycines are from the Eocene-aged Messel Pit in Germany.


Behavior

The majority of sand boas spend much of their time basking below the surface of the sand, with only their eyes or head exposed. When potential prey approaches, they erupt out of the sand, bite, and employ constriction to subdue it.


Feeding

The erycines' primary diet is rodents, but they have also been known to prey on lizards and birds.


Reproduction

Otherwise far removed from their boine relatives, erycines are generally ovoviviparous, i.e., giving birth to live young. At least three species lay eggs, however: the Calabar python, ''Calabaria reinhardtii'' (once regarded as a python for this reason), the Arabian sand boa, '' Eryx jayakari'', and the West African sand boa, ''E. muelleri''.


Smuggling and poaching in India

Poaching and smuggling of sand boas is often reported in India. Most of the smuggled snakes go to the United States. There is a misconception about their medicinal and aphrodisiacal properties, as well as the belief that keeping this snake as a pet brings wealth and prosperity.


Captivity

'' Eryx colubrinus'', '' E. conicus'' and '' E. johnii'' are frequently available in the exotic pet trade and are often captive-bred. They breed readily, their small size making them an attractive option. They are usually not aggressive species, though they sometimes have a tendency to bite, and also spend the vast majority of their time hiding. Other species are not commonly available, but are occasionally imported.


Genera

* Not including the nominate subspecies.
T Type genus.


See also

*
List of erycine species and subspecies This is a list of all extant genera, species and subspecies of the snakes of the subfamily Erycinae, otherwise referred to as erycines or Old World sand boas. It follows the taxonomy currently provided by ITIS, which is based on the continui ...


References


Further reading

* Bonaparte, Carlo Luciano (1831). ''Saggio di una distribuzione metodica degli animali vertebrati.'' Rome: Antonio Boulzaler. 144 pp. (Subfamily "Erycina ic, p. 67). (in Italian). * Goin CJ, Goin OB, Zug GR (1978). ''Introduction to Herpetology, Third Edition''. San Francisco: W.H. Freeman and Company. xi + 378 pp. . (Subfamily Erycinae, p. 319).


External links


The Sand Boa Page
a
Kingsnake.com
Accessed 16 July 2008. {{Taxonbar, from=Q2220578 Tetrapod subfamilies Taxa named by Charles Lucien Bonaparte