The Erycinae, also known as the Old World sand boas, are a

subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end botanical subfamily names with "-oideae", and zo ...

nonvenomous Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a sti ...

snakes Snakes are elongated Limbless vertebrate, limbless reptiles of the suborder Serpentes (). Cladistically Squamata, squamates, snakes are ectothermic, amniote vertebrates covered in overlapping Scale (zoology), scales much like other members of ...

in the

family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...

Boidae The Boidae, commonly known as boas or boids, are a family of nonvenomous snakes primarily found in the Americas, as well as Africa, Europe, Asia, and some Pacific islands. Boas include some of the world's largest snakes, with the green anaconda ...

Species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...

of the subfamily Erycinae are found in Europe,

Asia Minor Anatolia (), also known as Asia Minor, is a peninsula in West Asia that makes up the majority of the land area of Turkey. It is the westernmost protrusion of Asia and is geographically bounded by the Mediterranean Sea to the south, the Aegean ...

, Africa,

Arabia The Arabian Peninsula (, , or , , ) or Arabia, is a peninsula in West Asia, situated north-east of Africa on the Arabian plate. At , comparable in size to India, the Arabian Peninsula is the largest peninsula in the world. Geographically, the ...

, central and southwestern Asia, India, Sri Lanka, and western North America. Four

genera Genus (; : genera ) is a taxonomic rank above species and below family as used in the biological classification of living and fossil organisms as well as viruses. In binomial nomenclature, the genus name forms the first part of the binomial s ...

comprising 18

species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...

are currently recognized as being valid.

Description

Erycinae is a subfamily of stout-bodied snakes, all of which are competent burrowers. The largest, '' E. johnii'', rarely exceeds in total length (including tail). Most grow to around in total length. They have small eyes and hard, small scales to protect their skin from the grit of sand. A great deal of

sexual dimorphism Sexual dimorphism is the condition where sexes of the same species exhibit different Morphology (biology), morphological characteristics, including characteristics not directly involved in reproduction. The condition occurs in most dioecy, di ...

exists, with females generally becoming much larger than males. Erycines have skeletal adaptations to burrowing. The skull is more compact than in the subfamily Boinae. Also, the vertebrae of the tail are increased in size but reduced in number.

Distribution and habitat

Erycines are found in Southeastern Europe, Asia Minor, North, Central, West and East Africa, Arabia, Central and Southwest Asia, India, Pakistan, Sri Lanka, southwestern Canada, the western United States, and northwestern Mexico. McDiarmid, R.W.; Campbell, J.A., Touré, T. (1999). ''Snake Species of the World: A Taxonomic and Geographic Reference Vol. 1''. Washington, District of Columbia: Herpetologists' League. 511 pp. (series). (volume). Fossil erycines have been found in rock strata over 50 million years old, and were once widespread in

North America North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...

. Now, only four

remain in North America, as well as the Old World sand boas proper in Africa, Asia, and Southeastern

Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...

. The oldest known erycines are from the

Eocene The Eocene ( ) is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (Ma). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes ...

-aged

Messel Pit The Messel Formation is a Formation (geology), geologic formation in Hesse, central Germany, dating back to the Eocene Epoch (geology), epoch (about 47 Ma). Its geographic range is restricted to the Messel pit. There it unconformably overlie ...

in Germany, named ''

Rageryx schmidi ''Rageryx'' is an extinct genus of erycine boid that lived during the Eocene epoch. Distribution ''Rageryx schmidi'' is known from the Messel Formation of Germany Germany, officially the Federal Republic of Germany, is a country in ...

''.

Behavior

The majority of sand boas spend much of their time basking below the surface of the sand, with only their eyes or head exposed. When potential prey approaches, they erupt out of the sand, bite, and employ

constriction Constriction is a method used by several snake species to kill or subdue their prey. Although some species of venomous and mildly venomous snakes do use constriction to subdue their prey, most snakes which use constriction lack venom. The snake ...

to subdue it.

Feeding

The erycines' primary diet is rodents, but they have also been known to prey on lizards and birds.

Reproduction

Otherwise far removed from their boine relatives, erycines are generally

ovoviviparous Ovoviviparity, ovovivipary, ovivipary, or aplacental viviparity is a "bridging" form of reproduction between egg-laying oviparity, oviparous and live-bearing viviparity, viviparous reproduction. Ovoviviparous animals possess embryos that develo ...

, i.e., giving birth to live young. At least two species lay eggs, however: the Arabian sand boa, ''

Eryx jayakari ''Eryx jayakari'', known commonly as the Arabian sand boa or Jayakar's sand boa, is a species of snake in the family Boidae. The species is endemic to the Arabian Peninsula and Iran where it spends the day buried in the sand. Etymology Both th ...

'', and the West African sand boa, ''

Eryx muelleri ''Eryx muelleri'', known commonly as Müller's sand boa or the Saharan sand boa, is a species of snake in the family Boidae. The species is endemic to Africa. They are kept fairly regularly in the pet industry due to their docile nature and easy ...

''.

Smuggling and poaching in India

Poaching Poaching is the illegal hunting or capturing of wild animals, usually associated with land use rights. Poaching was once performed by impoverished peasants for subsistence purposes and to supplement meager diets. It was set against the huntin ...

and smuggling of sand boas is often reported in India. Most of the smuggled snakes go to the United States. There is a misconception about their medicinal and aphrodisiacal properties, as well as the belief that keeping this snake as a pet brings wealth and prosperity.

Captivity

'' Eryx colubrinus'', '' E. conicus'' and '' E. johnii'' are frequently available in the exotic pet trade and are often captive-bred. They breed readily, their small size making them an attractive option. They are usually not aggressive species, though they sometimes have a tendency to bite, and also spend the vast majority of their time hiding. Other species are not commonly available, but are occasionally imported. Sand Boa

Genera

* Not including the

nominate subspecies In biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. ...

.
^T

Type genus In biological taxonomy, the type genus (''genus typica'') is the genus which defines a biological family and the root of the family name. Zoological nomenclature According to the International Code of Zoological Nomenclature, "The name-bearin ...

References

External links

The Sand Boa Page
a
Kingsnake.com
Accessed 16 July 2008. {{Taxonbar, from=Q2220578 Snake subfamilies Taxa named by Charles Lucien Bonaparte