Sambirano
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The Sambirano region, also known as the Sambirano domain, is a
biogeographic Biogeography is the study of the distribution of species and ecosystems in geographic space and through geological time. Organisms and biological communities often vary in a regular fashion along geographic gradients of latitude, elevation, ...
region of northwestern
Madagascar Madagascar, officially the Republic of Madagascar, is an island country that includes the island of Madagascar and numerous smaller peripheral islands. Lying off the southeastern coast of Africa, it is the world's List of islands by area, f ...
. It has a distinctive lowland climate with year-round rain, differing from the rest of seasonally-dry western and northern Madagascar. The climate supports humid lowland rainforests, with greater similarity to those of eastern Madagascar than the dry deciduous forests characteristic of the Madagascar's western lowlands.


Geography

The region takes its name from the
Sambirano River The Sambirano is a river of northwestern Madagascar in the region of Diana. It has its sources at the Maromokotra peak and flows through the Tsaratanana Reserve to the Indian Ocean. Its delta occupies 250 km. The ecosystem of the river, mai ...
, which runs through the center of the region. The Sambirano region extends from the Ifasy River in the north to the Ampasindava Peninsula in the south, in the southwestern portion of
Diana Region Diana is a region in Madagascar at the northern part of the island. It borders the regions of Sava to the southeast and Sofia to the southwest. It covers an area of 19,266 km2, and had a population of 889,736 in 2018. The regional capital is ...
. It extends from the coast to the slopes of the
Tsaratanana Massif Madagascar is a large island in the Indian Ocean located off the eastern coast of Southern Africa, east of Mozambique. It has a total area of with of land and of water. Madagascar is the list of islands by area, fourth-largest island in the ...
, which bounds the region on the east. It also includes the islands of
Nosy Be Nosy Be (; formerly Nossi-bé and Nosse Be, ) is a volcanic island off the northwest coast of Madagascar. Nosy Be is Madagascar's largest and busiest tourist destination. It has an area of , and its population was 109,465 according to the provisio ...
and
Nosy Komba Nosy Komba (; ), also known as Nosy Ambariovato, is a small island in Madagascar, situated between the island of Nosy Be and the northwestern coast of the main island of Madagascar. Roughly circular, it rises sharply towards a plateau and the su ...
.


Climate

The region receives summer monsoon rains characteristic of the west coast, but also rain during the winter months from the easterly trade winds. The trade winds bring winter rain to eastern Madagascar but across most of the west are intercepted by Madagascar's central highlands. The convergence of easterly wind currents in the lee of the Tsaratanana Massif is responsible for the Sambirano region's higher rainfall. Average annual rainfall in
Ambanja Ambanja is a city and commune in northern Madagascar. According to 2018 census the population of Ambanja was 60,321. Geography Ambanja is located on the northern berth of the Sambirano River and is crossed by the Route Nationale 6 (Antsiranan ...
is 2171 mm. January is rainiest month, averaging 538.4 mm. May to October are drier months, with the lowest rainfall (24.6 mm) in July. Mean annual temperature averages 25.7 °C at Ambanja, with little seasonal variation and humidity throughout most of the year. As one ascends the slopes of Tsaratanana, rainfall and humidity increase, and temperature and seasonal rainfall variation decrease.


Flora

The predominant natural vegetation in the region is lowland rainforest, although little remains. Mature forests have a canopy of trees 25 to 30 meters high, festooned with abundant
liana A liana is a long-Plant stem, stemmed Woody plant, woody vine that is rooted in the soil at ground level and uses trees, as well as other means of vertical support, to climb up to the Canopy (biology), canopy in search of direct sunlight. T ...
s. There is a middle stratum of smaller, shade-tolerant trees, and a sparse understory of ferns, dwarf palms, shrubs, and herbaceous plants. Trees of the families
Sarcolaenaceae The Sarcolaenaceae are a family of flowering plants endemic to Madagascar. The family includes 79 species of mostly evergreen trees and shrubs in ten genera. Recent DNA studies indicate that the Sarcolaenaceae are a sibling taxon to the family ...
and
Myristicaceae The Myristicaceae are a family of flowering plants native to Africa, Asia, Pacific islands, and the Americas and has been recognized by most taxonomists. It is sometimes called the "nutmeg family", after its most famous member, '' Myristica fragr ...
, and genus ''
Anthostema ''Anthostema'' is a flowering plant genus in the Family Euphorbiaceae (spurge family) first described as a genus in 1824. It is native to Africa and Madagascar Madagascar, officially the Republic of Madagascar, is an island country that ...
'' are characteristic canopy trees, along with trees of families
Fabaceae Fabaceae () or Leguminosae,International Code of Nomen ...
(''
Cynometra ''Cynometra'' (from Greek language, Greek calque of ''puki anjing'', local Malay name for ''Cynometra cauliflora, C. cauliflora'' according to Georg Eberhard Rumphius, Rumphius) is genus of tropical forest trees with a pantropical distribution. ...
''),
Lauraceae Lauraceae, or the laurels, is a plant Family (biology), family that includes the bay laurel, true laurel and its closest relatives. This family comprises about 2850 known species in about 45 genus (biology), genera worldwide. They are dicotyled ...
(''
Ocotea ''Ocotea'' is a genus of flowering plants belonging to the family Lauraceae. Many are evergreen trees with lauroid leaves. There are over 520 species currently accepted within the genus, distributed mostly in tropical and subtropical areas of ...
'' and ''
Cryptocarya agathophylla ''Cryptocarya agathophylla'' (known sometimes as clove nutmeg) is a member of the laurel family, Lauraceae, and originates in Madagascar. Malagasy names include ', ', ' and '. The former genus name ''Ravensara'' is a latinization of the Malaga ...
''),
Ebenaceae The Ebenaceae are a family of flowering plants belonging to order Ericales. The family includes ebony and persimmon among about 768 species of trees and shrubs. It is distributed across the tropical and warmer temperate regions of the world. It i ...
(''
Diospyros ''Diospyros'' is a genus of over 700 species of deciduous and evergreen trees and shrubs. The majority are native to the tropics, with only a few species extending into temperate regions. Individual species valued for their hard, heavy, dark timb ...
''),
Euphorbiaceae Euphorbiaceae (), the spurge family, is a large family of flowering plants. In English, they are also commonly called euphorbias, which is also the name of Euphorbia, the type genus of the family. Most spurges, such as ''Euphorbia paralias'', ar ...
,
Arecaceae The Arecaceae () is a family (biology), family of perennial plant, perennial, flowering plants in the Monocotyledon, monocot order Arecales. Their growth form can be climbing palm, climbers, shrubs, tree-like and stemless plants, all commonly k ...
,
Rubiaceae Rubiaceae () is a family (biology), family of flowering plants, commonly known as the coffee, madder, or bedstraw family. It consists of terrestrial trees, shrubs, lianas, or herbs that are recognizable by simple, opposite leaves with Petiole ( ...
,
Sapindaceae The Sapindaceae are a family (biology), family of flowering plants in the order Sapindales known as the soapberry family. It contains 138 genera and 1,858 accepted species. Examples include Aesculus, horse chestnut, maples, ackee and lychee. The ...
, and
Anacardiaceae The Anacardiaceae, commonly known as the cashew family or sumac family, are a family of flowering plants, including about 83 genera with about 860 known species. Members of the Anacardiaceae bear fruits that are drupes and in some cases produce ...
. Human disturbance, has reduced the rainforest to scattered patches. Grasslands are maintained through regular burning, and are used for livestock grazing. Secondary forest, known as ''savoka'', grows in areas which were cleared for
shifting cultivation Shifting cultivation is an agricultural system in which plots of land are cultivated temporarily, then abandoned while post-disturbance fallow vegetation is allowed to freely grow while the cultivator moves on to another plot. The period of cul ...
and then left fallow. The palm-like native plant ''
Ravenala madagascariensis ''Ravenala madagascariensis'', commonly known as the traveller's tree, traveller's palm or East-West palm, is a species of monocotyledonous flowering plant found in Madagascar. It is not a true palm but a member of the family Strelitziaceae. N ...
'' is common in disturbed areas. Introduced species are increasingly common, including
citrus ''Citrus'' is a genus of flowering trees and shrubs in the family Rutaceae. Plants in the genus produce citrus fruits, including important crops such as oranges, mandarins, lemons, grapefruits, pomelos, and limes. ''Citrus'' is nativ ...
, ''
Psidium ''Psidium'' is a genus of trees and shrubs in the family Myrtaceae. It is native to warmer parts of the Western Hemisphere (Mexico, Central and South America, the West Indies the Galápagos islands). Many of the species bear edible fruits, and ...
'' (guava), and the bamboo ''
Cathariostachys capitata ''Cathariostachys'' is a genus of Madagascan bamboo in the grass family. ;Species # '' Cathariostachys capitata'' - eastern Madagascar # ''Cathariostachys madagascariensis'' (A.Camus) S.Dransf. - central Madagascar Madagascar, offici ...
''.


Fauna

Several species of
lemur Lemurs ( ; from Latin ) are Strepsirrhini, wet-nosed primates of the Superfamily (biology), superfamily Lemuroidea ( ), divided into 8 Family (biology), families and consisting of 15 genera and around 100 existing species. They are Endemism, ...
are endemic or mostly endemic to the Sambirano region, including the
black lemur The black lemur (''Eulemur macaco'') is a species of lemur from the family Lemuridae. Like all lemurs, it is endemic to Madagascar. Originally, the species was thought to have two subspecies, ''Eulemur macaco macaco'' and ''Eulemur macaco flavi ...
(''Eulemur macaco''), Pariente's fork-marked lemur (''Phaner parienti''), Nosy Be mouse lemur (''Microcebus mamiratra''), and Nosy Be sportive lemur (''Lepilemur tymerlachsoni'').


Conservation

The largest remaining area of lowland rainforest is in Lokobe National Park on Nosy Be.
Manongarivo Special Reserve Manongarivo Reserve is a wildlife reserve in the North-West of Madagascar in the region of Diana Diana most commonly refers to: * Diana (name), given name (including a list of people with the name) * Diana (mythology), ancient Roman goddess of ...
preserves some enclaves of humid forest along with transitional lowland dry forest.


References

{{reflist Madagascar subhumid forests Diana Region