Saltasaurinae is a
subfamily of
titanosaur
Titanosaurs (or titanosaurians; members of the group Titanosauria) were a diverse group of sauropod dinosaurs, including genera from all seven continents. The titanosaurs were the last surviving group of long-necked sauropods, with taxa still th ...
ian
sauropods known from the late
Cretaceous
The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of ...
period of
South America
South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the souther ...
,
India
India, officially the Republic of India ( Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the ...
and
Madagascar
Madagascar (; mg, Madagasikara, ), officially the Republic of Madagascar ( mg, Repoblikan'i Madagasikara, links=no, ; french: République de Madagascar), is an island country in the Indian Ocean, approximately off the coast of East Africa ...
.
Description

Saltasaurines are relatively small sauropods with the general body shape of a small head, long neck, four limbs, and a long tail. They range from the small ''
Ibirania
''Ibirania'' (meaning "Ibirá wanderer" or "tree wanderer") is a genus of dwarf saltasaurine titanosaur dinosaur from the Late Cretaceous (Santonian to Campanian) São José do Rio Preto Formation ( Bauru Basin) of south-east Brazil. The type spe ...
'' at around , to the larger ''
Neuquensaurus'' at .
A currently unnamed fragmentary sauropod from
Madagascar
Madagascar (; mg, Madagasikara, ), officially the Republic of Madagascar ( mg, Repoblikan'i Madagasikara, links=no, ; french: République de Madagascar), is an island country in the Indian Ocean, approximately off the coast of East Africa ...
may turn out to be a saltasaurine longer than ''Neuquensaurus''. The weight of saltasaurines is very light compared to that of some of the largest dinosaurs.
Thomas R. Holtz Jr.
Thomas Richard Holtz Jr. (born September 13, 1965) is an American vertebrate palaeontologist, author, and principal lecturer at the University of Maryland's Department of Geology. He has published extensively on the phylogeny, morphology, ecomorp ...
found the genera range from around , with ''Saltasaurus'' and an unnamed genus on both extremes, respectively.
[Holtz, Thomas R. Jr. (2011) ''Dinosaurs: The Most Complete, Up-to-date Encyclopedia for Dinosaur Lovers of All Ages,']
Winter 2010 Appendix.
/ref>
Saltasaurinae is the only known group of sauropods found with armour from almost every species
In biology, a species is the basic unit of Taxonomy (biology), classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of ...
. The most probable reason for the bony studs and plates is that it evolved for defence against theropods like ''Abelisaurus
''Abelisaurus'' (; "Abel's lizard") is a genus of predatory abelisaurid theropod dinosaur alive during the Late Cretaceous Period (Campanian) of what is now South America. It was a bipedal carnivore that probably reached about in length, althou ...
'' and ''Carnotaurus
''Carnotaurus'' (; ) is a genus of theropod dinosaur that lived in South America during the Late Cretaceous period, probably sometime between 71 and 69 million years ago. The only species is ''Carnotaurus sastrei''. Known from a single well-p ...
''. Saltasaurine armour has led to controversies; in 1929, the paleontologist Friedrich von Huene
Friedrich von Huene, born Friedrich Richard von Hoinigen, (March 22, 1875 – April 4, 1969) was a German paleontologist who renamed more dinosaurs in the early 20th century than anyone else in Europe. He also made key contributions about v ...
named the genus '' Loricosaurus'' for armour he thought to be from ankylosauria
Ankylosauria is a group of herbivorous dinosaurs of the order Ornithischia. It includes the great majority of dinosaurs with armor in the form of bony osteoderms, similar to turtles. Ankylosaurs were bulky quadrupeds, with short, powerful limbs ...
ns. These bones were found to have similarities to those later discovered on sauropods like ''Saltasaurus'' and ''Neuquensaurus'', and as such, ''Loricosaurus'' may be the same as one of the other genera.
Age and distribution
Saltasaurines lived in the late Cretaceous
The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of ...
, from the early Campanian
The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campani ...
to the Maastrichtian
The Maastrichtian () is, in the ICS geologic timescale, the latest age (uppermost stage) of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spanned the interv ...
(about 80–66 million years ago) when they went extinct along with all other non-avian dinosaurs. ''Saltasaurus'' is the only named Saltasaurine that lived later in the Maastrichtian
The Maastrichtian () is, in the ICS geologic timescale, the latest age (uppermost stage) of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spanned the interv ...
than 68 million years ago. ''Loricosaurus'' and ''Neuquensaurus'' lived around 71 million years ago and the later surviving ''Jainosaurus'' lived around 68 million years ago. An unnamed Saltasaurine from Madagascar
Madagascar (; mg, Madagasikara, ), officially the Republic of Madagascar ( mg, Repoblikan'i Madagasikara, links=no, ; french: République de Madagascar), is an island country in the Indian Ocean, approximately off the coast of East Africa ...
would have probably survived later, until the Cretaceous-Paleogene extinction event, around 66 million years ago.
The subfamily Saltasaurinae is known almost completely from the Southern Hemisphere with South American forms. '' Jainosaurus'' and ''Abditosaurus
''Abditosaurus'' (meaning "forgotten lizard") is an extinct genus of titanosaurian sauropod dinosaur from the Late Cretaceous (Maastrichtian) Tremp Group of Catalonia, Spain. The type and only species is ''Abditosaurus kuehnei''. Phylogenetic ana ...
'' are two of the only definite saltasaurines from outside the Southern Hemisphere and is two of the only ones from outside of South America.
Classification
In a 1992 study on '' Saltasaurus'', Jaime Powell
Jaime Eduardo Powell (January 13, 1953 – February 1, 2016) was an Argentine paleontologist who described the titanosaur sauropod dinosaur taxa Aeolosaurus and found evidence that titanosaurs have osteoderms.
Research
Powell described the first ...
named Saltasaurinae, a new subfamily within Titanosauridae
Lithostrotia is a clade of derived titanosaur sauropods that lived during the Early Cretaceous and Late Cretaceous. The group was defined by Upchurch ''et al.'' in 2004 as the most recent common ancestor of '' Malawisaurus'' and '' Saltasaurus ...
(a family now considered invalid). He found many features uniting the group, consisting of the type genus and '' Neuquensaurus''. This group was later supported and defined by Salgado ''et al.'' (1997). They defined the subfamily as "the clade including the most recent common ancestor of ''Neuquensaurus australis'', ''Saltasaurus loricatus'', and all of its descendants". They conducted a phylogeny and found that the subfamily was sister to ''Alamosaurus
''Alamosaurus'' (; meaning "Ojo Alamo lizard") is a genus of opisthocoelicaudiine titanosaurian sauropod dinosaurs, containing a single known species, ''Alamosaurus sanjuanensis'', from the late Cretaceous Period (geology), Period of what is now ...
'' and only included ''Neuquensaurus'' and ''Saltasaurus''. Paul Sereno
Paul Callistus Sereno (born October 11, 1957) is a professor of paleontology at the University of Chicago and a National Geographic "explorer-in-residence" who has discovered several new dinosaur species on several continents, including at sites ...
defined it in 1998, unaware of Salgado's work and gave it a new definition as a stem clade. His definition was "All saltasaurids more closely related to ''Saltasaurus'' than to ''Opisthocoelicaudia
''Opisthocoelicaudia'' is a genus of sauropod dinosaur of the Late Cretaceous Period discovered in the Gobi Desert of Mongolia. The type species is ''Opisthocoelicaudia skarzynskii''. A well-preserved skeleton lacking only the head and neck was ...
''". In 2003, Jeffrey A. Wilson
Jeffrey A. Wilson, also known as JAW, is a Paleontology, paleontologist and professor of geology, geological sciences and assistant curator at the Museum of Paleontology at the University of Michigan.
His Doctor of Philosophy, doctoral dissert ...
and Paul Upchurch elaborated on this definition to "all Saltasauridae more closely related to ''Saltasaurus loricatus'' than to ''Opisthocoelicaudia skaryzinskii''".
Below is a cladogram by Villa ''et al.'' (2022), from the description of the European saltasaurine ''Abditosaurus
''Abditosaurus'' (meaning "forgotten lizard") is an extinct genus of titanosaurian sauropod dinosaur from the Late Cretaceous (Maastrichtian) Tremp Group of Catalonia, Spain. The type and only species is ''Abditosaurus kuehnei''. Phylogenetic ana ...
''.
References
{{Taxonbar, from=Q2535065
Cretaceous dinosaurs
Saltasaurids