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''Gaultheria shallon'' is an
evergreen In botany, an evergreen is a plant which has Leaf, foliage that remains green and functional throughout the year. This contrasts with deciduous plants, which lose their foliage completely during the winter or dry season. Consisting of many diffe ...
shrub A shrub or bush is a small to medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees by their multiple ...
in the heather
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
(
Ericaceae The Ericaceae () are a Family (biology), family of flowering plants, commonly known as the heath or heather family, found most commonly in acidic and infertile growing conditions. The family is large, with about 4,250 known species spread acros ...
), native to western North America. Common names include salal (), shallon, or (mainly in Britain) gaultheria.


Description

''Gaultheria shallon'' is tall, sprawling to erect. It is loosely to densely branched and often forms dense, nearly impenetrable thickets. The
twig A twig is a thin, often short, branch of a tree or Bush (plant), bush. The buds on the twig are an important diagnostic characteristic, as are the abscission scars where the leaves have fallen away. The color, texture, and patterning of the t ...
s are reddish-brown, with shredding bark. Twigs can live up to 16 years or more, but bear
leaves A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, ...
only the first few years. The leaves are alternate,
evergreen In botany, an evergreen is a plant which has Leaf, foliage that remains green and functional throughout the year. This contrasts with deciduous plants, which lose their foliage completely during the winter or dry season. Consisting of many diffe ...
, leathery, thick and egg shaped. They are shiny and dark green on the upper surface, and rough and lighter green on the lower. Each finely and sharply
serrate Serration is a saw-like appearance or a row of sharp or tooth-like projections. A serrated cutting edge has many small points of contact with the material being cut. By having less contact area than a smooth blade or other edge, the applied pr ...
leaf is long. Each leaf generally lives for 2 to 4 years before it is replaced. The
inflorescence In botany, an inflorescence is a group or cluster of flowers arranged on a plant's Plant stem, stem that is composed of a main branch or a system of branches. An inflorescence is categorized on the basis of the arrangement of flowers on a mai ...
of flowers consists of a
bract In botany, a bract is a modified or specialized leaf, associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves in size, color, shape or texture. They also lo ...
eate
raceme A raceme () or racemoid is an unbranched, indeterminate growth, indeterminate type of inflorescence bearing flowers having short floral stalks along the shoots that bear the flowers. The oldest flowers grow close to the base and new flowers are ...
, one-sided, with 5–15 flowers at the ends of branches. Each flower is composed of a deeply five-parted, glandular-haired
calyx CALYX, Inc. is a non-profit publisher of art and literature by women founded in 1976 based in Corvallis, Oregon. CALYX publishes both '' CALYX, A Journal of Art and Literature by Women'' twice a year and CALYX Books, which publishes one to three ...
and an urn-shaped pink to white, glandular to hairy, five-lobed
petal Petals are modified leaves that form an inner whorl surrounding the reproductive parts of flowers. They are often brightly coloured or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''corol ...
s (corolla), long. The
fruit In botany, a fruit is the seed-bearing structure in flowering plants (angiosperms) that is formed from the ovary after flowering. Fruits are the means by which angiosperms disseminate their seeds. Edible fruits in particular have long propaga ...
is reddish to blue, rough-surfaced, covered in tiny hairs, nearly spherical and 6 to 10 mm in diameter. The fruits are 'pseudoberries', or capsules made up of a fleshy outer calyx, and each fruit contains an average of 126 brown, reticulate
seed In botany, a seed is a plant structure containing an embryo and stored nutrients in a protective coat called a ''testa''. More generally, the term "seed" means anything that can be Sowing, sown, which may include seed and husk or tuber. Seeds ...
s approximately 0.1 mm in length. These berries are also edible.


Etymology

Lewis and Clark Lewis may refer to: Names * Lewis (given name), including a list of people with the given name * Lewis (surname), including a list of people with the surname Music * Lewis (musician), Canadian singer * " Lewis (Mistreated)", a song by Radiohe ...
reported the local
Chinook Jargon Chinook Jargon (' or ', also known simply as ''Chinook'' or ''Jargon'') is a language originating as a pidgin language, pidgin trade language in the Pacific Northwest. It spread during the 19th century from the lower Columbia River, first to othe ...
name of the omnipresent evergreen shrub to be ''shallon'', ''shelwel'', or ''shellwell'', but when Scottish naturalist David Douglas arrived at Fort George in April 1825 he noted that it was not called ''shallon'' but rather ''salal or sallal''. The genus ''Gaultheria'' was named by
Pehr Kalm Pehr Kalm (6 March 1716 – 16 November 1779), also known as Peter Kalm, was a Swedish-Finnish List of explorers, explorer, botany, botanist, natural history, naturalist, and Agricultural economics, agricultural economist. He was one of the most ...
for his guide in Canada, fellow botanist
Jean François Gaultier Jean François Gaultier (6 October 1708 in La Croix-Avranchin – 10 July 1756 in Quebec) was a French physician and botanist. He was the king's physician for New France and was the regular physician at the Hôtel-Dieu de Québec. Gaultier, bein ...
. In the
Squamish language Squamish ( ; ', ''sníchim'' meaning "language") is a Coast Salish language spoken by the Squamish people of the Pacific Northwest. It is spoken in southwestern British Columbia, Canada, centred on their reserve communities in Squamish, Nort ...
, the fruits are called t’áḵa7 and the bush is called t’áḵa7áy̓. In the
Saanich dialect Saanich (also Sənčáθən, written as in Saanich orthography, ) is the language of the First Nations Saanich people in the Pacific Northwest region of northwestern North America. Saanich is a Coast Salishan language in the Northern Straits ...
, salal berries are called DAḴE In the Lushootseed language, the month of August is named pədt̕aqaʔ, literally 'time of the salal.'


Distribution

Salal is found in Southeast
Alaska Alaska ( ) is a non-contiguous U.S. state on the northwest extremity of North America. Part of the Western United States region, it is one of the two non-contiguous U.S. states, alongside Hawaii. Alaska is also considered to be the north ...
,
British Columbia British Columbia is the westernmost Provinces and territories of Canada, province of Canada. Situated in the Pacific Northwest between the Pacific Ocean and the Rocky Mountains, the province has a diverse geography, with rugged landscapes that ...
,
Washington Washington most commonly refers to: * George Washington (1732–1799), the first president of the United States * Washington (state), a state in the Pacific Northwest of the United States * Washington, D.C., the capital of the United States ** A ...
and
Oregon Oregon ( , ) is a U.S. state, state in the Pacific Northwest region of the United States. It is a part of the Western U.S., with the Columbia River delineating much of Oregon's northern boundary with Washington (state), Washington, while t ...
and along much of the
California California () is a U.S. state, state in the Western United States that lies on the West Coast of the United States, Pacific Coast. It borders Oregon to the north, Nevada and Arizona to the east, and shares Mexico–United States border, an ...
coast. It grows as far north as
Baranof Island Baranof Island is an island in the northern Alexander Archipelago in the Alaska Panhandle, in Alaska. The name "Baranof" was given to the island in 1805 by Imperial Russian Navy captain Yuri Lisyansky, U. F. Lisianski in honor of Alexander Andrey ...
,
Alaska Alaska ( ) is a non-contiguous U.S. state on the northwest extremity of North America. Part of the Western United States region, it is one of the two non-contiguous U.S. states, alongside Hawaii. Alaska is also considered to be the north ...
.
Western poison oak ''Toxicodendron diversilobum'' (syn. ''Rhus diversiloba''), commonly named Pacific poison oak or western poison oak, is a woody vine or shrub in the sumac family, Anacardiaceae. It is widely distributed in western North America, inhabiting co ...
is a common associate in the
California Coast Ranges The Coast Ranges of California span from Del Norte County, California, Del Norte or Humboldt County, California, south to Santa Barbara County, California, Santa Barbara County. The other three coastal California mountain ranges are the Trans ...
.


In Europe

The species was introduced to Britain in 1828 by David Douglas, who intended the plant to be used as an
ornamental Ornamental may refer to: *Ornamental grass, a type of grass grown as a decoration *Ornamental iron, mild steel that has been formed into decorative shapes, similar to wrought iron work *Ornamental plant, a plant that is grown for its ornamental qua ...
. It is believed to have been planted as cover for
pheasant Pheasants ( ) are birds of several genera within the family Phasianidae in the order Galliformes. Although they can be found all over the world in introduced (and captive) populations, the pheasant genera's native range is restricted to Eura ...
s on shooting estates.


Ecology


Preferred sites

''G. shallon'' grows in moist to dry, montane to lowland coastal conifer forests as well as forested peatland and
swamp A swamp is a forested wetland.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p. Swamps are considered to be transition zones because both land and water play a role in ...
s throughout the Pacific Northwest down to Southern California. It is a common coniferous forest understory species where shade is not heavy and may dominate large areas with its spreading
rhizome In botany and dendrology, a rhizome ( ) is a modified subterranean plant stem that sends out roots and Shoot (botany), shoots from its Node (botany), nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from ...
s. Individual plants may live for hundreds of years, spreading from their rhizomes. The plant is sensitive to frost. It is tolerant of salt spray and grows well on stabilized dunes, exposed slopes, and bluffs near the ocean, as well as in coastal brushfields and coastal
shore pine ''Pinus contorta'', with the common names lodgepole pine and shore pine, and also known as twisted pine, and contorta pine, is a common tree in western North America. It is common near the ocean shore and in dry montane forests to the subalpine, ...
and
Sitka spruce ''Picea sitchensis'', the Sitka spruce, is a large, coniferous, evergreen tree growing to just over tall, with a trunk diameter at breast height that can exceed 5 m (16 ft). It is by far the largest species of spruce and the fifth- ...
forests. Salal tolerates moderate shade and does well in sun. It grows in a variety of soils but prefers moist sandy or peaty soils, and commonly establishes on decaying wood and stumps or as an
epiphyte An epiphyte is a plant or plant-like organism that grows on the surface of another plant and derives its moisture and nutrients from the air, rain, water (in marine environments) or from debris accumulating around it. The plants on which epiphyt ...
in extremely humid areas. Conifer regeneration is typically better on sites dominated by salal than on sites dominated by the western swordfern or
vine maple ''Acer circinatum'', or vine maple, is a species of maple native to northwestern North America. Vine maple typically grows as a low-elevation coastal tree in temperate areas of high precipitation such as the west coast of Oregon and northern Cal ...
, although it still competes with conifer seedlings for water, nutrients, and space. There are mixed reports on its impact on
Douglas-fir The Douglas fir (''Pseudotsuga menziesii'') is an evergreen conifer species in the pine family, Pinaceae. It is the tallest tree in the Pinaceae family. It is native to western North America and is also known as Douglas-fir, Douglas spruce, Or ...
regrowth; on some sites it may hinder it, but on others it may add nutrients to the soil and actually increase conifer regeneration.


Use by animals

The leaves are browsed by
deer A deer (: deer) or true deer is a hoofed ruminant ungulate of the family Cervidae (informally the deer family). Cervidae is divided into subfamilies Cervinae (which includes, among others, muntjac, elk (wapiti), red deer, and fallow deer) ...
and
elk The elk (: ''elk'' or ''elks''; ''Cervus canadensis'') or wapiti, is the second largest species within the deer family, Cervidae, and one of the largest terrestrial mammals in its native range of North America and Central and East Asia. ...
, and it is an important winter food for those species. Browsing is heaviest when other low-growing species become covered in snow; in Western Washington salal leaves composed 30.4% of deer diet by volume in January, compared to only 0.5% in June. The leaves have relatively low nutritional value, and deer which feed exclusively on them have shown signs of
malnutrition Malnutrition occurs when an organism gets too few or too many nutrients, resulting in health problems. Specifically, it is a deficiency, excess, or imbalance of energy, protein and other nutrients which adversely affects the body's tissues a ...
, cementing their status as a winter and emergency food for
ungulate Ungulates ( ) are members of the diverse clade Euungulata ("true ungulates"), which primarily consists of large mammals with Hoof, hooves. Once part of the clade "Ungulata" along with the clade Paenungulata, "Ungulata" has since been determined ...
s.
Beavers Beavers (genus ''Castor'') are large, semiaquatic rodents of the Northern Hemisphere. There are two existing species: the North American beaver (''Castor canadensis'') and the Eurasian beaver (''C. fiber''). Beavers are the second-large ...
and the white-footed vole are known to feed on salal leaves, as are domestic
goat The goat or domestic goat (''Capra hircus'') is a species of Caprinae, goat-antelope that is mostly kept as livestock. It was domesticated from the wild goat (''C. aegagrus'') of Southwest Asia and Eastern Europe. The goat is a member of the ...
s and
sheep Sheep (: sheep) or domestic sheep (''Ovis aries'') are a domesticated, ruminant mammal typically kept as livestock. Although the term ''sheep'' can apply to other species in the genus '' Ovis'', in everyday usage it almost always refers to d ...
in some areas. The
band-tailed pigeon The band-tailed pigeon (''Patagioenas fasciata'') is a pigeon native to the Americas, and the largest pigeon native to North America. They are a native species throughout the Southwestern United States and Mexico, extending south to Peru. Taxo ...
,
wrentit The wrentit (''Chamaea fasciata'') is a small bird that lives in chaparral, California oak woodland, oak woodlands, and bushland on the western coast of North America. It is the Monotypic taxon, only species in the genus ''Chamaea''. Its systema ...
, various species of
grouse Grouse are a group of birds from the order (biology), order Galliformes, in the family (biology), family Phasianidae. Grouse are presently assigned to the Tribe (biology), tribe Tetraonini (formerly the subfamily Tetraoninae and the family Tetr ...
, and numerous songbirds consume the fruit, as do mammals such as the
red squirrel The red squirrel (''Sciurus vulgaris''), also called Eurasian red squirrel, is a species of tree squirrel in the genus ''Sciurus''. It is an arboreal and primarily herbivorous rodent and common throughout Eurasia. Taxonomy There have been ...
,
black bear Black bear or Blackbear may refer to: Animals * American black bear (''Ursus americanus''), a North American bear species * Asian black bear (''Ursus thibetanus''), an Asian bear species Music * Black Bear (band), a Canadian First Nations group ...
, black-tailed deer, Townsend's chipmunk, and the
Douglas squirrel The Douglas squirrel (''Tamiasciurus douglasii'') is a pine squirrel found in western North America, from the Pacific Northwest (including the northwestern coastal states of the United States as well as the southwestern coast of British Columb ...
. Salal flowers are pollinated by hummingbirds, bees, and flies, and are browsed by deer. It provides important cover and hiding places for a variety of species, from large ungulates to small birds and mammals.


Fire ecology

Salal is adapted to a
fire regime A fire regime is the pattern, frequency, and intensity of the bushfires and wildfires that prevail in an area over long periods of time. It is an integral part of fire ecology, and renewal for certain types of ecosystems. A fire regime describes th ...
of infrequent fires, from intervals of 50 to 500+ years. Although aboveground portions of the plant may be consumed, rhizomes in the soil will survive and resprout after light to moderate fires. Severe fires on dry, shallow soil will penetrate into the roots and kill the plant.


Invasiveness

It readily colonizes
heathland A heath () is a shrubland habitat found mainly on free-draining infertile, acidic soils and is characterised by open, low-growing woody vegetation. Moorland is generally related to high-ground heaths with—especially in Great Britain—a coole ...
and acidic woodland habitats in southern England, often forming very tall and dense evergreen stands which smother other vegetation. Although heathland managers widely regard it as a problem weed on unmanaged heathland, it is readily browsed by cattle (especially in winter), so where traditional grazing management has been restored, the dense stands become broken up and the plant becomes a more scattered component of the heathland vegetation. As an
invasive species An invasive species is an introduced species that harms its new environment. Invasive species adversely affect habitats and bioregions, causing ecological, environmental, and/or economic damage. The term can also be used for native spec ...
, salal is resistant to many
herbicide Herbicides (, ), also commonly known as weed killers, are substances used to control undesired plants, also known as weeds.EPA. February 201Pesticides Industry. Sales and Usage 2006 and 2007: Market Estimates. Summary in press releasMain page f ...
s because of its waxy foliage, although Garlon and Silvex can be effective when properly applied. Attempts at mechanical removal may break up rhizomes and stimulate new growth, thus increasing salal cover.


Uses

The dark blue berries and young leaves are edible and efficient
appetite suppressant An anorectic is a drug that reduces appetite, resulting in lower food consumption, leading to weight loss. These substances work by affecting the central nervous system or certain neurotransmitters to create a feeling of fullness or reduce the desi ...
s, both with a unique flavor. The berries were a significant food resource for some Native American tribes, who ate them fresh and dried them into cakes. They were also used as a
sweetener A sweetener is a substance added to food or drink to impart the flavor of sweetness, either because it contains a type of sugar, or because it contains a sweet-tasting sugar substitute. Various natural non-sugar sweeteners (NSS) and artificial s ...
, and the
Haida Haida may refer to: Haida people Many uses of the word derive from the name of an indigenous people of the Pacific Northwest Coast of North America. * Haida people, an Indigenous ethnic group of North America (Canada) ** Council of the Haida Nati ...
used them to thicken salmon eggs. The leaves of the plant were also sometimes used to flavor fish soup. More recently, the berries are used locally in jams, preserves, and pies. They are often combined with
Oregon-grape ''Berberis aquifolium'', the Oregon grape or holly-leaved barberry, is a North American species of flowering plant in the family Berberidaceae. It is an evergreen shrub growing up to tall and wide, with pinnate leaves consisting of spiny leafle ...
because salal's mild sweetness partially masks the former's tartness. Salal is widely cultivated as an ornamental both within and outside of its native range, useful for ground cover and landscaping. In the Pacific Northwest, the harvesting of ''G. shallon'' is the heart of a large industry which supplies cut evergreens worldwide for use in
floral arrangement Floral design or flower arrangement is the art of using plant material and flowers to create an eye-catching and balanced composition or display. Evidence of refined floral design is found as far back as the culture of ancient Egypt. Floral desi ...
s. It is used in
native plant In biogeography, a native species is indigenous to a given region or ecosystem if its presence in that region is the result of only local natural evolution (though often popularised as "with no human intervention") during history. The term is equi ...
gardens and sold as "Lemon Leaf".


Medicinal properties

The species has been used for its medicinal properties by local natives for generations, although its medicinal uses are not widely known. The leaves have an
astringent An astringent (sometimes called adstringent) is a chemical that shrinks or constricts body tissues. The word derives from the Latin '' adstringere'', which means "to bind fast". Astringency, the dry, puckering or numbing mouthfeel caused by t ...
effect, making it an effective
anti-inflammatory Anti-inflammatory is the property of a substance or treatment that reduces inflammation, fever or swelling. Anti-inflammatory drugs, also called anti-inflammatories, make up about half of analgesics. These drugs reduce pain by inhibiting mechan ...
and anti-cramping herb. Leaves prepared in a tea or tincture are thought to decrease internal inflammation (such as bladder inflammation), stomach or duodenal ulcers,
heartburn Heartburn is a burning sensation felt behind the breastbone. It is a symptom that is commonly linked to acid reflux and is often triggered by food, particularly fatty, sugary, spicy, chocolate, citrus, onion-based and tomato-based products. Ly ...
, indigestion, sinus inflammation, diarrhea, moderate fever, inflamed/irritated throat, and menstrual cramps. A
poultice A poultice or cataplasm, also called a fomentation, is a soft moist mass, often heated and medicated, that is applied to the skin to reduce inflammation, soothe pain, promote healing, or otherwise treat wounds or ailments. Soft materials like cer ...
of the leaf can be used externally to ease discomfort from insect bites and stings.


References


External links

* {{Taxonbar, from=Q941600 shallon Flora of the West Coast of the United States Flora of Alaska Flora of British Columbia Flora of Oregon Flora of Washington (state) Flora of California Cuisine of the Western United States Garden plants of North America Groundcovers Canadian cuisine Plants used in Native American cuisine