Saigini is a

tribe The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide usage of the term in English is in the discipline of anthropology. This definition is contested, in part due to confl ...

artiodactyl The even-toed ungulates (Artiodactyla , ) are ungulates—hoofed animals—which bear weight equally on two (an even number) of their five toes: the third and fourth. The other three toes are either present, absent, vestigial, or pointing poste ...

mammals Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fu ...

of the

Bovidae The Bovidae comprise the biological family of cloven-hoofed, ruminant mammals that includes cattle, bison, buffalo, antelopes, and caprines. A member of this family is called a bovid. With 143 extant species and 300 known extinct species ...

family, subfamily

Antilopinae The Antilopines are even-toed ungulates belonging to the subfamily Antilopinae of the family Bovidae. The members of tribe Antilopini include the gazelles, blackbucks, springboks, gerenuks, dibatags, and Central Asian gazelles, are often ...

, comprising two

species In biology, a species is the basic unit of Taxonomy (biology), classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of ...

of medium-sized

antelope The term antelope is used to refer to many species of even-toed ruminant that are indigenous to various regions in Africa and Eurasia. Antelope comprise a wastebasket taxon defined as any of numerous Old World grazing and browsing hoofed mamm ...

s that inhabit the

Eurasian steppe The Eurasian Steppe, also simply called the Great Steppe or the steppes, is the vast steppe ecoregion of Eurasia in the temperate grasslands, savannas and shrublands biome. It stretches through Hungary, Bulgaria, Romania, Moldova and Tra ...

Taxonomy

Description

The tribe was created in 1945 by the American

paleontologist Paleontology (), also spelled palaeontology or palæontology, is the scientific study of life that existed prior to, and sometimes including, the start of the Holocene epoch (roughly 11,700 years before present). It includes the study of foss ...

and theoretical

biologist A biologist is a scientist who conducts research in biology. Biologists are interested in studying life on Earth, whether it is an individual cell, a multicellular organism, or a community of interacting populations. They usually speciali ...

George Gaylord Simpson George Gaylord Simpson (June 16, 1902 – October 6, 1984) was an American paleontologist. Simpson was perhaps the most influential paleontologist of the twentieth century, and a major participant in the modern synthesis, contributing '' Tempo ...

. According to the Simpson classification the tribe comprises two

monospecific In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispe ...

genera Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial ...

: * Tribus Saigini ** Genus '' Pantholops'' *** ''

Pantholops hodgsonii The Tibetan antelope or chiru (''Pantholops hodgsonii'') (, pronounced ; ) is a medium-sized bovid native to the northeastern Tibetan plateau. Most of the population live within the Chinese border, while some scatter across India and Bhutan in ...

'' ** Genus ''

Saiga The saiga antelope (, ''Saiga tatarica''), or saiga, is a critically endangered antelope which during antiquity inhabited a vast area of the Eurasian steppe spanning the foothills of the Carpathian Mountains in the northwest and Caucasus in ...

'' *** ''

Saiga tatarica The saiga antelope (, ''Saiga tatarica''), or saiga, is a critically endangered antelope which during antiquity inhabited a vast area of the Eurasian steppe spanning the foothills of the Carpathian Mountains in the northwest and Caucasus in ...

A controversial taxonomy

''Saiga'' and ''Pantholops'' are problematic genera in taxonomy. ''Saiga'' was traditionally classified as a member of the tribe Saigini, within the subfamily

Caprinae The subfamily Caprinae, also sometimes referred to as the tribe Caprini, is part of the ruminant family Bovidae, and consists of mostly medium-sized bovids. A member of this subfamily is called a caprine, or, more informally, a goat-antelope ( ...

but some authors suggested that the genus ''Saiga'' was closer to the subfamily

. In 2000 Groves analyzed the morphological characters of ''

Procapra ''Procapra'' is a genus of Asian gazelles,Myers, P., R. Espinosa, C. S. Parr, T. Jones, G. S. Hammond, and T. A. Dewey. 2008. The Animal Diversity Web (online). Accessed May 31, 2008 at http://animaldiversity.org including three living species: * ...

'', '' Prodorcas'' and ''

'', and proposed three basal groups of Antilopinae, one of which included a clade ''Saiga'' + ''Procapra''.Groves, C. P. (2000). "Phylogenetic relationships within recent Antlopini (Bovidae). In: Elizabeth S. Vrba and George B. Schaller, eds. ''Antelopes, Deer, and Relatives: Fossil Record, Behavioral Ecology, Systematics, and Conservation''. New Haven and London: Yale University Press. , pp. 223-233. The genus ''

Panthalops The Tibetan antelope or chiru (''Pantholops hodgsonii'') (, pronounced ; ) is a medium-sized bovid native to the northeastern Tibetan plateau. Most of the population live within the Chinese border, while some scatter across India and Bhutan in ...

'' is monotypic (''P. hodgsonii''), and is sometimes included in the tribe Saigini on the basis of similar morphological features, most of which are

plesiomorphic In phylogenetics, a plesiomorphy ("near form") and symplesiomorphy are synonyms for an ancestral character shared by all members of a clade, which does not distinguish the clade from other clades. Plesiomorphy, symplesiomorphy, apomorphy, an ...

. However,

molecular A molecule is a group of two or more atoms held together by attractive forces known as chemical bonds; depending on context, the term may or may not include ions which satisfy this criterion. In quantum physics, organic chemistry, and bioch ...

and morphological findings suggest that ''Pantholops hodgsonii'' should be classified more correctly in the subfamily Caprinae, but the status of the

Tibetan gazelle The goa (''Procapra picticaudata''), also known as the Tibetan gazelle, is a species of antelope that inhabits the Tibetan plateau. Description The goa is a relatively small antelope, with slender and graceful bodies. Both males and females sta ...

in this subfamily remains uncertain. Data on

cytochrome b Cytochrome b within both molecular and cell biology, is a protein found in the mitochondria of eukaryotic cells. It functions as part of the electron transport chain and is the main subunit of transmembrane cytochrome bc1 and b6f complexes. ...

and small portions of 12S and 16S

ribosomal genes Ribosomal DNA (rDNA) is a DNA sequence that codes for ribosomal RNA. These sequences regulate transcription initiation and amplification, and contain both transcribed and non-transcribed spacer segments. In the human genome there are 5 chromo ...

suggest

phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ...

affinities of ''Saiga'' and ''Procapra'' with Antilopinae. However, these studies do not clarify the close relationships within this clade. Given this uncertainty, some authors have suggested that the Saigini taxon should be abandoned.

References

{{Authority control True antelopes Mammal tribes Taxa named by George Gaylord Simpson