Saccolaimus Mixtus
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The Papuan sheath-tailed bat (''Saccolaimus mixtus'') is a species of
bat Bats are flying mammals of the order Chiroptera (). With their forelimbs adapted as wings, they are the only mammals capable of true and sustained flight. Bats are more agile in flight than most birds, flying with their very long spread-out ...
in the family
Emballonuridae Emballonuridae is a family of microbats, many of which are referred to as sac-winged or sheath-tailed bats. They are widely distributed in tropical and subtropical regions around the world. The earliest fossil records are from the Eocene. Desc ...
which occurs at the
Cape York Peninsula The Cape York Peninsula is a peninsula located in Far North Queensland, Australia. It is the largest wilderness in northern Australia.Mittermeier, R.E. et al. (2002). Wilderness: Earth's last wild places. Mexico City: Agrupación Sierra Madre, ...
and
New Guinea New Guinea (; Hiri Motu: ''Niu Gini''; , fossilized , also known as Papua or historically ) is the List of islands by area, world's second-largest island, with an area of . Located in Melanesia in the southwestern Pacific Ocean, the island is ...
. The poorly known species hunts in open forests for night flying insects.


Taxonomy

The first description was published by
Ellis Le Geyt Troughton Ellis Le Geyt Troughton (born in Sydney on 29 April 1893; died 30 November 1974) was an Australian zoologist and mammalogist. Biography Ellis Troughton began to exercise his interest in mammals at fourteen years of age, taking a role at the A ...
in 1925. Troughton found features typical of the genus, but notes the clearly defined wing pouches that characteristic of the genus ''
Taphozous ''Taphozous'' is a genus of the family Emballonuridae. The wide distribution of the genus includes several regions of Australia, Indonesia, Papua New Guinea and Africa. ''Taphozous'' comes from the Greek τάφος, meaning "a tomb". The common n ...
''; the author makes reference to a note by
Edward Ramsay Edward Bannerman Ramsay, (17 January 1793– 27 December 1872), usually referred to as Dean Ramsay, was a clergyman of the Scottish Episcopal Church, and Dean of Edinburgh in that communion from 1841, has a place in literature through his ''R ...
in 1878 on an undiagnosed species of that genus which resembled specimens found at Cape York. Troughton describes the holotype, a male, and two other specimens, a series purchased from Kendal Broadbent who had obtained them at
Port Moresby (; Tok Pisin: ''Pot Mosbi''), also referred to as Pom City or simply Moresby, is the capital and largest city of Papua New Guinea. It is one of the largest cities in the southwestern Pacific (along with Jayapura) outside of Australia and New ...
in 1878. The name was assessed in a revision of the genus in 1991, replacing the previously recognised combination ''Taphozous mixtus'' (Troughton) 1925. The number of specimens available has been limited to another two collected before 1973 in New Guinea, and some 25 specimens obtained from the Australian continent. The epithet ''mixtus'', meaning intermediate or mixed, is derived from Latin. The common names of the species includes Papuan sheath-tailed bat and Troughton's pouched bat. The proposed vernacular for the Australian region is Cape York sheath-tailed bat Also cited are the names New Guinea sheathtail bat, wing-pouched saccolaimus and allied freetail bat.


Description

A species of '' Saccolaimus'' with dark hoary grey fur for across the back, the colour is darkest at the head and shoulder, the ventral side is pale buff-grey. The range of measurements of the forearm are 62 to 68 millimetres, the total length of head and body is 72 to 77 mm. It weighs around 24 grams. The ears are around 18 mm from the notch at the head to the tip, the bare skin of these, along with the muzzle and other parts of the face, is a dark brown colour. The pouch at the wing, a feature of related species, has a lining of whitish hairs. A throat pouch is also found on the species. this is quite prominent in males and presented as a bare patch of skin in females. The first harmonic of the call of the ''S. Mixtus'' is audible, their signals are recorded at 9 kilohertz, distinguishable in visual analysis from ''
Saccolaimus flaviventris The yellow-bellied sheath-tailed bat (''Saccolaimus flaviventris''), also known as the yellow-bellied sheathtail or yellow-bellied pouched bat, is a microbat species of the family Emballonuridae found extensively in Australia and less commonly in ...
'' (yellow-bellied) and closely resembling '' S. saccolaimus'' (naked-rumped pouched bat). Their rapid wing beats, small size and ventral colour allows them to be distinguished in flight from known species by workers spotting with torchlight. They resemble another species also found in Australia ''S. flaviventris'', which is discernible by its greater size, darker
pelage A fur is a soft, thick growth of hair that covers the skin of almost all mammals. It consists of a combination of oily guard hair on top and thick underfur beneath. The guard hair keeps moisture from reaching the skin; the underfur acts as an ...
and the presence of a developed throat pouch in the female.


Distribution and habitat

The distribution range extends from northern
Australia Australia, officially the Commonwealth of Australia, is a country comprising mainland Australia, the mainland of the Australia (continent), Australian continent, the island of Tasmania and list of islands of Australia, numerous smaller isl ...
to the southwest of
Papua New Guinea Papua New Guinea, officially the Independent State of Papua New Guinea, is an island country in Oceania that comprises the eastern half of the island of New Guinea and offshore islands in Melanesia, a region of the southwestern Pacific Ocean n ...
. The occurrence at Cape York is only recorded at a few locations, seven collected at
Weipa Weipa () is a coastal mining town in the local government area of Weipa Town in Queensland. It is one of the largest towns on the Cape York Peninsula. It exists because of the enormous bauxite deposits along the coast. The Port of Weipa is main ...
and several more at Brown's Creek near the
Pascoe River The Pascoe River is a river in Far North Queensland, Australia. The headwaters rise under Mount Yangee in the Table Range, part of the Great Dividing Range at the northern end of Cape York Peninsula. The river initially flows south then west p ...
at the north of the Cape York district. ''S. mixtus'' is associated with forest of western Queensland dominated by the stringybark ''
Eucalyptus tetrodonta ''Eucalyptus tetrodonta'', commonly known as Darwin stringybark or messmate, is a species of medium-sized to tall tree that is endemic to northern Australia. It has rough, stringy or fibrous bark on the trunk and branches, lance-shaped leave ...
'', and is recorded in small groups occupying dead specimens of that tree. The little information on their life history and habitat is derived from observations in New Guinea, where it is associated with roosts limestone caves and forages over the canopy of the forest. Their daytime roosts in Australia are
tree hollows A tree hollow or tree hole is a semi-enclosed cavity which has naturally formed in the trunk or branch of a tree. They are found mainly in old trees, whether living or not. Hollows form in many species of trees. They are a prominent feature of n ...
. They appear to favour hunting over the canopy and at open flight paths over streams and clearings. Surveys undertaken in Australia have suggested evidence of a greater range than was previously established, extending the area it may occur to southern parts and western regions of Cape York and at islands to the north. The species has been reported at
Pormpuraaw, Queensland Pormpuraaw (pronounced ''porm-pure-ow'') is a coastal town and a locality in the Aboriginal Shire of Pormpuraaw, Queensland, Australia. Pormpuraaw is an Aboriginal community situated on the west coast of Cape York Peninsula approximately halfw ...
.


Conservation status

The
IUCN The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the status ...
notes the population of ''Saccolaimus mixtus'' as declining and is classified as near threatened by
habitat loss Habitat destruction (also termed habitat loss or habitat reduction) occurs when a natural habitat is no longer able to support its native species. The organisms once living there have either moved elsewhere, or are dead, leading to a decrease ...
in 2017, the work group amending the previous listing as data deficient in 2008. The population in Australia is threatened by the activities of
bauxite mining Bauxite () is a sedimentary rock with a relatively high aluminium content. It is the world's main source of aluminium and gallium. Bauxite consists mostly of the aluminium minerals gibbsite (), boehmite (γ-AlO(OH)), and diaspore (α-AlO(OH) ...
and inappropriate fire regimes. The effects of land clearing and other ecological changes in New Guinea had not been assessed, but are intensive at the type locality around Port Moresby, and new records in the decades preceding the reassessment were absent. The status listed by the Queensland state government is least concern.


References

{{Taxonbar, from=Q1834270 Emballonuridae Bats of Oceania Bats of Australia Mammals of Papua New Guinea Vulnerable fauna of Australia Vulnerable fauna of Oceania Mammals described in 1925 Taxonomy articles created by Polbot Taxa named by Ellis Le Geyt Troughton Bats of New Guinea