The rusty-margined guan (''Penelope superciliaris'') is a species of

bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...

in the family

Cracidae The chachalacas, guans, and curassows are birds in the Family (biology), family Cracidae. These are species of tropical and subtropical Central America, Central and South America. The range of one species, the plain chachalaca, just reaches south ...

, which includes the

chachalaca Chachalacas are galliform birds from the genus ''Ortalis''. These birds are found in wooded habitats in the far southern United States (Texas), Mexico, and Central and South America. They are social, can be very noisy and often remain fairly ...

guan Guan may refer to: * Guan (bird), any of a number of bird species of the family Cracidae, of South and Central America * Guan (surname), several similar Chinese surnames ** Guān, Chinese surname * Guan (state), ancient Chinese city-state * Guan ( ...

s, and

curassow Curassows are one of the three major groups of cracid birds. They comprise the largest-bodied species of the cracid family. Three of the four genera are restricted to tropical South America; a single species of ''Crax'' ranges north to Mexico. ...

s. It is found in

Argentina Argentina, officially the Argentine Republic, is a country in the southern half of South America. It covers an area of , making it the List of South American countries by area, second-largest country in South America after Brazil, the fourt ...

Bolivia Bolivia, officially the Plurinational State of Bolivia, is a landlocked country located in central South America. The country features diverse geography, including vast Amazonian plains, tropical lowlands, mountains, the Gran Chaco Province, w ...

Brazil Brazil, officially the Federative Republic of Brazil, is the largest country in South America. It is the world's List of countries and dependencies by area, fifth-largest country by area and the List of countries and dependencies by population ...

, and

Paraguay Paraguay, officially the Republic of Paraguay, is a landlocked country in South America. It is bordered by Argentina to the Argentina–Paraguay border, south and southwest, Brazil to the Brazil–Paraguay border, east and northeast, and Boli ...

Taxonomy and systematics

The

International Ornithological Committee The International Ornithologists' Union (IOU) is an international organization for the promotion of ornithology. It links basic and applied research and nurtures education and outreach activities. Specifically, the IOU organizes and funds global co ...

(IOC) and the ''Handbook of the Birds of the World'' taxonomies recognize the four rusty-margined guan subspecies listed in the box to the right. The

Clements taxonomy ''The Clements Checklist of Birds of the World'' is a book by Jim Clements which presents a list of the bird species of the world. The most recent printed version is the sixth edition (2007), but has been updated yearly, the last version in 202 ...

does not recognize ''P. s. pseudonyma'' as separate but includes its population in the nominate subspecies.HBW and BirdLife International (2020) ''Handbook of the Birds of the World and BirdLife International digital checklist of the birds of the world'' Version 5. Available at: http://datazone.birdlife.org/userfiles/file/Species/Taxonomy/HBW-BirdLife_Checklist_v5_Dec20.zip xls zipped 1 MBretrieved May 27, 2021Clements, J. F., T. S. Schulenberg, M. J. Iliff, S. M. Billerman, T. A. Fredericks, J. A. Gerbracht, D. Lepage, B. L. Sullivan, and C. L. Wood. 2021. The eBird/Clements checklist of Birds of the World: v2021. Downloaded from https://www.birds.cornell.edu/clementschecklist/download/ Retrieved August 25, 2021 Birds in Brazil's

Alagoas Alagoas () is one of the 27 federative units of Brazil and is situated in the eastern part of the Northeast Region, Brazil, Northeast Region. It borders: Pernambuco (N and NW); Sergipe (S); Bahia (SW); and the Atlantic Ocean (E). Its capital is ...

state are distinctive and might be a separate subspecies.del Hoyo, J. and G. M. Kirwan (2020). Rusty-margined Guan (''Penelope superciliaris''), version 1.0. In Birds of the World (J. del Hoyo, A. Elliott, J. Sargatal, D. A. Christie, and E. de Juana, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bow.rumgua1.01 retrieved September 27, 2021 The rusty-margined guan and the Marail guan (''P. marail'') form a

superspecies In biology, a species complex is a group of closely related organisms that are so similar in appearance and other features that the boundaries between them are often unclear. The taxa in the complex may be able to hybridize readily with each oth ...

.Remsen, J. V., Jr., J. I. Areta, E. Bonaccorso, S. Claramunt, A. Jaramillo, D. F. Lane, J. F. Pacheco, M. B. Robbins, F. G. Stiles, and K. J. Zimmer. Version 24 August 2021. A classification of the bird species of South America. American Ornithological Society. https://www.museum.lsu.edu/~Remsen/SACCBaseline.htm retrieved August 24, 2021

Description

The rusty-margined guan is long. The nominate subspecies weighs and ''P. s. jacupemba'' . The species' upperparts are dark olive to brownish with the eponymous reddish margins on the wing

covert Secrecy is the practice of hiding information from certain individuals or groups who do not have the "need to know", perhaps while sharing it with other individuals. That which is kept hidden is known as the secret. Secrecy is often controver ...

s and scapulars. The belly is also rusty. The face is pale with a

supercilium The supercilium is a plumage feature found on the heads of some bird species. It is a stripe which runs from the base of the bird's beak above its eye, finishing somewhere towards the rear of the bird's head.Dunn and Alderfer (2006), p. 10 Also k ...

whose color ranges from white to buff or ochre in the different subspecies. ''P. s. jacupemba'' paler overall than the nominate and ''P. s. major'' is darker.

Distribution and habitat

The nominate subspecies of rusty-margined guan is found in Amazonian Brazil south of the Amazon River. ''P. s. pseudonyma'' when treated separately is more westerly than the nominate

sensu stricto ''Sensu'' is a Latin word meaning "in the sense of". It is used in a number of fields including biology, geology, linguistics, semiotics, and law. Commonly it refers to how strictly or loosely an expression is used in describing any particular c ...

. ''P. s. jacupemba'' is found from central and southern Brazil into eastern Bolivia. ''P. s. major'' is found in extreme southeastern Brazil, eastern Paraguay, and extreme northeastern Argentina. The rusty-margined guan inhabits a wide variety of landscapes across its large range. Examples include the interior and edges of heavy forest, gallery forest, ''

restinga Restingas () are a distinct type of coastal tropical and subtropical moist broadleaf forest in eastern Brazil. They form on sandy, acidic, and nutrient-poor soils, and are characterized by medium-sized trees and shrubs adapted to the drier and n ...

'', woodlands in ''

cerrado The Cerrado () is a vast ecoregion of Tropics, tropical savanna in central Brazil, being present in the states of Goiás, Mato Grosso do Sul, Mato Grosso, Tocantins, Maranhão, Piauí, Bahia, Minas Gerais, São Paulo (state), São Paulo, Paraná ...

'' grasslands, ''

caatinga Caatinga () is a type of semi-arid tropical vegetation, and an ecoregion characterized by this vegetation in interior northeastern Brazil. The name "Caatinga" comes from the Tupi word '' ka'atinga'', meaning "white forest" or "white vegetat ...

'', mangroves, and ''

Eucalyptus ''Eucalyptus'' () is a genus of more than 700 species of flowering plants in the family Myrtaceae. Most species of ''Eucalyptus'' are trees, often Mallee (habit), mallees, and a few are shrubs. Along with several other genera in the tribe Eucalyp ...

'' plantations. It is generally a bird of the lowlands but can be found as high as in parts of Brazil.

Behavior

Movement

The rusty-margined guan is essentially sedentary, though in some areas birds roost in higher areas and move down to feed with an elevation change of up to . It seldom flies more than about and usually much less.

Feeding

The rusty-margined guan forages alone, in pairs, or in groups of up to 10 birds. It mainly feeds between about above the ground, but often feeds on the ground as well. Its diet is almost entirely fruit; one study identified the fruit of 117 plant species. It also eats small amounts of flowers, leaves, and insects.

Breeding

The rusty-margined guan's breeding season spans October to February in Argentina and August to February in southeastern Brazil; it has not been well defined elsewhere. Its nest is a platform made of sticks lined with live and dead leaves, usually built in a tree. The clutch size is three eggs.

Vocal and non-vocal sounds

The rusty-margined guan gives a wing-whirring display that has not well defined. It gives a "gruff barking" call, probably in alarm but possibly in other contexts as well.

Status

The

IUCN The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the status ...

has assessed the rusty-margined guan as being Near Threatened. It is reasonably common only in protected areas though even there poaching is a concern.

References

External links

Rusty-margined Guan videos
on the Internet Bird Collection

VIREO * ttps://web.archive.org/web/20070228130539/http://www.terrabrasil.org.br/ecosistema/fotos/aves05/jacu.jpg Photo-High Resbr>Article
– terrabrasil.org.br-''"Brazil Ecosystems"''

*Associação Mãe-da-lu

Photo, notes {{Taxonbar, from=Q1270743 rusty-margined guan Birds of the Caatinga Birds of the Cerrado Birds of the Pantanal Birds of Brazil Birds of Paraguay rusty-margined guan Taxonomy articles created by Polbot