The Russulales are an

order Order, ORDER or Orders may refer to: * A socio-political or established or existing order, e.g. World order, Ancien Regime, Pax Britannica * Categorization, the process in which ideas and objects are recognized, differentiated, and understood ...

of the

Agaricomycetes The Agaricomycetes are a class of fungi in the division Basidiomycota. The taxon is roughly identical to that defined for the Homobasidiomycetes (alternatively called holobasidiomycetes) by Hibbett & Thorn, with the inclusion of Auriculariales ...

, (which include the

agaric An agaric () is a type of fungal fruiting body characterized by the presence of a pileus (cap) that is clearly differentiated from the stipe (stalk), with lamellae (gills) on the underside of the pileus. It is a type of mushroom (or toadstool) ...

genera ''

Russula ''Russula'' is a very large genus composed of around 750 worldwide species of fungi. The genus was described by Christian Hendrik Persoon in 1796. The mushrooms are fairly large, and brightly colored – making them one of the most recognizable ...

'' and ''

Lactarius ''Lactarius'' is a genus of mushroom-producing, ectomycorrhizal fungi, containing several Edible mushroom, edible species. The species of the genus, common name, commonly known as milk-caps, are characterized by the milky fluid ("latex") they ex ...

'' and their polyporoid and corticioid relatives). According to the ''Dictionary of the Fungi'' (10th edition, 2008), the order consists of 12

families Family (from ) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictability, structure, and safety as ...

, 80

genera Genus (; : genera ) is a taxonomic rank above species and below family as used in the biological classification of living and fossil organisms as well as viruses. In binomial nomenclature, the genus name forms the first part of the binomial s ...

, and 1767

species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...

. According to ''Species Fungorum'' (January 2016), the order contains 13 families, 117 genera (16 not assigned to a family), and 3,060 species. Russuloid agarics represent an independent evolutionary line of agarics, not directly related to the

Agaricales The Agaricales are an order (biology), order of fungi in the division (mycology), division Basidiomycota. As originally conceived, the order contained all the agarics (gilled mushrooms), but subsequent research has shown that not all agarics are ...

. This group also includes a number of russuloid

hypogeous Hypogeal, hypogean, hypogeic and hypogeous (; ) are biological terms describing an organism's activity below the soil surface. In botany, a seed is described as showing hypogeal germination when the cotyledons of the germinating seed remain no ...

fungi,

polypores Polypores, also called bracket or shelf fungi, are a morphological group of basidiomycete-like gilled mushrooms and hydnoid fungi that form large fruiting bodies called conks, which are typically woody, circular, shelf- or bracket-shaped, ...

such as ''

Bondarzewia ''Bondarzewia'' is a widely distributed genus of fungi in the family Bondarzewiaceae. The genus was circumscribed by mycologist Rolf Singer Rolf Singer (June 23, 1906 – January 18, 1994) was a German mycologist and Taxonomy (biology), taxo ...

'', some

tooth fungi The hydnoid fungi are a group of fungi in the Basidiomycota with basidiocarps (fruit bodies) producing spores on pendant, tooth-like or spine-like projections. They are colloquially called tooth fungi. Originally such fungi were referred to the ge ...

(e.g. '' Auriscalpium vulgare''), and

club fungi The clavarioid fungi are a group of fungi in the Basidiomycota typically having erect, simple or branched basidiocarps (fruit bodies) that are formed on the ground, on decaying vegetation, or on dead wood. They are colloquially called club fungi a ...

e.g. '' Artomyces''.

Basidiospores A basidiospore is a reproductive spore produced by basidiomycete fungi, a grouping that includes mushrooms, shelf fungi, rusts, and smuts. Basidiospores typically each contain one haploid nucleus that is the product of meiosis, and they are pr ...

in this group are typically ornamented with

amyloid Amyloids are aggregates of proteins characterised by a fibrillar morphology of typically 7–13 nm in diameter, a β-sheet secondary structure (known as cross-β) and ability to be stained by particular dyes, such as Congo red. In the human ...

warts or reticulation but a few exceptions are known, e.g. '' Heterobasidion annosum''. The genus ''

Clavicorona ''Clavicorona'' is a fungal genus in the family Clavariaceae. The genus was circumscribed by Maxwell Stanford Doty in 1947, who included the species ''C. pyxidata'', ''C. cristata'', ''C. taxophila'', and ''C. candelabrum'' ...

'' was often treated in the Russulales, but its

type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...

, '' C. taxophila'', is in the

. The remaining species are retained in the Russulales in the genus '' Artomyces''.

Genera ''Incertae sedis''

There are several genera classified in the Russulales that are i) poorly known, ii) have not been subjected to DNA analysis, or iii) if analysed phylogenetically do not group with as yet named or identified families, and have not been assigned to a specific family (i.e., ''

Incertae sedis or is a term used for a taxonomy (biology), taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertainty ...

'' with respect to familial placement). These include: *'' Aleurocystidiellum'' *'' Dentipellopsis'' *'' Gloeoasterostroma'' *'' Gloeohypochnicium'' *''

Gloeopeniophorella ''Gloeopeniophorella'' is a genus of crust-like, wood-decaying fungi in the family Russulaceae. It contains six known species. ''Gloeopeniophorella'' was first described by Brazilian mycologist Johannes Rick Johannes (João Evangelista) Rick (1 ...

'' *'' Haloaleurodiscus'' *'' Neoalbatrellus'' *'' Polypus'' *'' Scopulodontia'' *'' Scytinostromella'' *'' Xeroceps''

References

External links

Russulales News

{{Authority control Basidiomycota orders