The russet sparrow (''Passer cinnamomeus''), also called the cinnamon or cinnamon tree sparrow, is a

passerine A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped') which includes more than half of all bird species. Sometimes known as perching birds, passerines generally have an anisodactyl arrangement of their ...

bird of the sparrow

family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...

Passeridae. A stout little seed-eating bird with a thick

bill Bill(s) may refer to: Common meanings * Banknote, paper cash (especially in the United States) * Bill (law), a proposed law put before a legislature * Invoice, commercial document issued by a seller to a buyer * Bill, a bird or animal's beak Pl ...

, it has a body length of . Its

plumage Plumage () is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, there can b ...

is mainly warm

rufous Rufous () is a color that may be described as reddish-brown or brownish- red, as of rust or oxidised iron. The first recorded use of ''rufous'' as a color name in English was in 1782. However, the color is also recorded earlier in 1527 as a d ...

above and

grey Grey (more frequent in British English) or gray (more frequent in American English) is an intermediate color between black and white. It is a neutral or achromatic color, meaning that it has no chroma. It is the color of a cloud-covered s ...

below. It exhibits

sexual dimorphism Sexual dimorphism is the condition where sexes of the same species exhibit different Morphology (biology), morphological characteristics, including characteristics not directly involved in reproduction. The condition occurs in most dioecy, di ...

, with the plumage of both sexes patterned similarly to that of the corresponding sex of

house sparrow The house sparrow (''Passer domesticus'') is a bird of the Old World sparrow, sparrow family Passeridae, found in most parts of the world. It is a small bird that has a typical length of and a mass of . Females and young birds are coloured pa ...

. Its vocalisations are sweet and musical chirps, which when strung together form a song. Three

subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...

are recognised, differing chiefly in the yellowness of their underparts. The subspecies ''rutilans'' and ''intensior'' breed in parts of

eastern Asia East Asia is a geocultural region of Asia. It includes China, Japan, Mongolia, North Korea, South Korea, and Taiwan, plus two special administrative regions of China, Hong Kong and Macau. The economies of China, Japan, South Korea, and Taiwan ...

, where they are usually found in light

woodland A woodland () is, in the broad sense, land covered with woody plants (trees and shrubs), or in a narrow sense, synonymous with wood (or in the U.S., the '' plurale tantum'' woods), a low-density forest forming open habitats with plenty of sunli ...

, and the subspecies ''cinnamomeus'' breeds in the

Himalayas The Himalayas, or Himalaya ( ), is a mountain range in Asia, separating the plains of the Indian subcontinent from the Tibetan Plateau. The range has some of the Earth's highest peaks, including the highest, Mount Everest. More than list of h ...

, where it is usually associated with terrace cultivation. The russet sparrow is the typical sparrow of human habitations in towns where the house and

Eurasian tree sparrow The Eurasian tree sparrow (''Passer montanus'') is a passerine bird in the sparrow family with a rich chestnut crown and nape and a black patch on each pure white cheek. The sexes are similarly plumaged, and young birds are a duller version o ...

s are absent. In the southern part of its range, the russet sparrow prefers higher altitudes, but in the north it breeds by the sea. The russet sparrow is known well enough in the Himalayas to have a distinct name in some languages, and is depicted in

Japanese art Japanese art consists of a wide range of art styles and media that includes Jōmon pottery, ancient pottery, Japanese sculpture, sculpture, Ink wash painting, ink painting and Japanese calligraphy, calligraphy on silk and paper, Ukiyo-e, paint ...

. This sparrow feeds mainly on the

seed In botany, a seed is a plant structure containing an embryo and stored nutrients in a protective coat called a ''testa''. More generally, the term "seed" means anything that can be Sowing, sown, which may include seed and husk or tuber. Seeds ...

s of

herb Herbs are a widely distributed and widespread group of plants, excluding vegetables, with savory or aromatic properties that are used for flavoring and garnishing food, for medicinal purposes, or for fragrances. Culinary use typically distingu ...

s and

grain A grain is a small, hard, dry fruit (caryopsis) – with or without an attached husk, hull layer – harvested for human or animal consumption. A grain crop is a grain-producing plant. The two main types of commercial grain crops are cereals and ...

s, but it also eats

berries A berry is a small, pulpy, and often edible fruit. Typically, berries are juicy, rounded, brightly colored, sweet, sour or tart, and do not have a stone fruit, stone or pit (fruit), pit although many wikt:pip#Etymology 2, pips or seeds may be p ...

and

insect Insects (from Latin ') are Hexapoda, hexapod invertebrates of the class (biology), class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (Insect morphology#Head, head, ...

s, particularly during the

breeding season Seasonal breeders are animal species that successfully mate only during certain times of the year. These times of year allow for the optimization of survival of young due to factors such as ambient temperature, food and water availability, and ch ...

. This diet makes it a minor pest in agricultural areas, but also a predator of insect pests. While breeding, it is not social, as its nests are dispersed. It forms flocks when not breeding, although it associates with other bird species infrequently. In some parts of its range, the russet sparrow migrates, at least to lower altitudes. Its nest is located in a tree cavity, or a hole in a cliff or building. The male chooses the nest site before finding a mate and uses the nest for

courtship display A courtship display is a set of display behaviors in which an animal, usually a male, attempts to attract a mate; the mate exercises choice, so sexual selection acts on the display. These behaviors often include ritualized movement ("dances"), ...

. The typical

clutch A clutch is a mechanical device that allows an output shaft to be disconnected from a rotating input shaft. The clutch's input shaft is typically attached to a motor, while the clutch's output shaft is connected to the mechanism that does th ...

contains five or six whitish

eggs An egg is an organic vessel in which an embryo begins to develop. Egg, EGG or eggs may also refer to: Biology * Egg cell, the female reproductive cell (gamete) in oogamous organisms Food * Eggs as food Places * Egg, Austria * Egg, Switzerland ...

. Both sexes incubate and feed the young.

Taxonomy

The English ornithologist

John Gould John Gould (; 14 September 1804 – 3 February 1881) was an English ornithologist who published monographs on birds, illustrated by plates produced by his wife, Elizabeth Gould (illustrator), Elizabeth Gould, and several other artists, includ ...

described a specimen of the russet sparrow collected in the Himalayas at a meeting of the

Zoological Society of London The Zoological Society of London (ZSL) is a charity and organization devoted to the worldwide animal conservation, conservation of animals and their habitat conservation, habitats. It was founded in 1826. Since 1828, it has maintained London Zo ...

in December 1835 under the

binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin grammatical forms, altho ...

''Pyrgita cinnamomea''. His description was included in the volume of the society's proceedings for 1835 that was published on 8 April 1836. Dated 1835 on the title page but published in 1836. Its

specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...

comes from the

Neo-Latin Neo-LatinSidwell, Keith ''Classical Latin-Medieval Latin-Neo Latin'' in ; others, throughout. (also known as New Latin and Modern Latin) is the style of written Latin used in original literary, scholarly, and scientific works, first in Italy d ...

, "cinnamon-coloured". The russet sparrow was described as ''Fringilla rutilans'', by the Dutch zoologist

Coenraad Jacob Temminck Coenraad Jacob Temminck (; 31 March 1778 – 30 January 1858) was a Dutch people, Dutch patrician, Zoology, zoologist and museum director. Biography Coenraad Jacob Temminck was born on 31 March 1778 in Amsterdam in the Dutch Republic. Fro ...

, from a specimen collected in

Japan Japan is an island country in East Asia. Located in the Pacific Ocean off the northeast coast of the Asia, Asian mainland, it is bordered on the west by the Sea of Japan and extends from the Sea of Okhotsk in the north to the East China Sea ...

. Temminck's description is contained in his ''Nouveau recueil de planches coloriées d'oiseaux'' which was issued in 102 ''livraisons'' or parts between 1820 and 1839. It was at one time believed that the ''livraison'' containing the description of the russet sparrow was issued in 1835 but it has now been established that it appeared sometime in 1836 but as the precise date is not known under the rules of the

International Commission on Zoological Nomenclature The International Commission on Zoological Nomenclature (ICZN) is an organization dedicated to "achieving stability and sense in the scientific naming of animals". Founded in 1895, it currently comprises 26 commissioners from 20 countries. Orga ...

it is deemed to have been published on 31 December 1836. Most taxonomists give priority to Gould's publication and use the binomial name ''Passer cinnamomeus'' for the russet sparrow. The russet sparrow has usually been placed in the

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

Passer ''Passer'' is a genus of Old World sparrow, sparrows, also known as the true sparrows. The genus contains 28 species and includes the house sparrow and the Eurasian tree sparrow, two of the most common birds in the world. They are small birds wi ...

'', and within this genus it is seen as a part of the "

Palearctic The Palearctic or Palaearctic is a biogeographic realm of the Earth, the largest of eight. Confined almost entirely to the Eastern Hemisphere, it stretches across Europe and Asia, north of the foothills of the Himalayas, and North Africa. Th ...

black-bibbed sparrow" group, which includes the Eurasian tree sparrow as well as the house sparrow. It has generally been seen as a close relation of the house sparrow, and

Richard Meinertzhagen Colonel Richard Meinertzhagen, CBE, DSO (3 March 1878 – 17 June 1967) was a British soldier, intelligence officer, and ornithologist. He had a decorated military career spanning Africa and the Middle East. He was credited with creating an ...

even considered it to be the same species as the Somali sparrow, one of the house sparrow's closest relatives. However, studies of

mitochondrial DNA Mitochondrial DNA (mtDNA and mDNA) is the DNA located in the mitochondrion, mitochondria organelles in a eukaryotic cell that converts chemical energy from food into adenosine triphosphate (ATP). Mitochondrial DNA is a small portion of the D ...

indicate that the russet sparrow is an early offshoot or basal species among the Palearctic black-bibbed sparrows. While mitochrondrial DNA suggests

speciation Speciation is the evolutionary process by which populations evolve to become distinct species. The biologist Orator F. Cook coined the term in 1906 for cladogenesis, the splitting of lineages, as opposed to anagenesis, phyletic evolution within ...

in ''Passer'' occurred during the

Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...

and

Pliocene The Pliocene ( ; also Pleiocene) is the epoch (geology), epoch in the geologic time scale that extends from 5.33 to 2.58 British ornithologist J. Denis Summers-Smith estimated that the russet sparrow separated from the other Palearctic black-bibbed sparrows about 25,000 to 15,000 years ago, during the last glacial period. During this time, sparrows would have been isolated in ice-free refugia, such as the lower

Yangtze The Yangtze or Yangzi ( or ) is the longest river in Eurasia and the third-longest in the world. It rises at Jari Hill in the Tanggula Mountains of the Tibetan Plateau and flows including Dam Qu River the longest source of the Yangtze, i ...

valley, which Summers-Smith considered the most likely centre of evolution for the russet sparrow. Thirteen

have been described, but only three are widely recognised, these differing largely in the colour of their underparts. The subspecies ''Passer cinnamomea rutilans'' breeds in Japan,

Korea Korea is a peninsular region in East Asia consisting of the Korean Peninsula, Jeju Island, and smaller islands. Since the end of World War II in 1945, it has been politically Division of Korea, divided at or near the 38th parallel north, 3 ...

Taiwan Taiwan, officially the Republic of China (ROC), is a country in East Asia. The main geography of Taiwan, island of Taiwan, also known as ''Formosa'', lies between the East China Sea, East and South China Seas in the northwestern Pacific Ocea ...

, and southeastern and central

China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...

. The subspecies ''intensior'', described in 1922 by

Walter Rothschild Lionel Walter Rothschild, 2nd Baron Rothschild, Baron de Rothschild, (8 February 1868 – 27 August 1937) was a British banker, politician, zoologist, and soldier, who was a member of the Rothschild family. As a Zionist leader, he was present ...

from

Yunnan Yunnan; is an inland Provinces of China, province in Southwestern China. The province spans approximately and has a population of 47.2 million (as of 2020). The capital of the province is Kunming. The province borders the Chinese provinces ...

, breeds in southwest China and parts of

India India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...

Burma Myanmar, officially the Republic of the Union of Myanmar; and also referred to as Burma (the official English name until 1989), is a country in northwest Southeast Asia. It is the largest country by area in Mainland Southeast Asia and ha ...

Laos Laos, officially the Lao People's Democratic Republic (LPDR), is the only landlocked country in Southeast Asia. It is bordered by Myanmar and China to the northwest, Vietnam to the east, Cambodia to the southeast, and Thailand to the west and ...

, and

Vietnam Vietnam, officially the Socialist Republic of Vietnam (SRV), is a country at the eastern edge of mainland Southeast Asia, with an area of about and a population of over 100 million, making it the world's List of countries and depende ...

. In a large part of

Sichuan Sichuan is a province in Southwestern China, occupying the Sichuan Basin and Tibetan Plateau—between the Jinsha River to the west, the Daba Mountains to the north, and the Yunnan–Guizhou Plateau to the south. Its capital city is Cheng ...

''intensior'' intergrades with ''P. c. rutilans'', and a number of subspecies names have been proposed for the intergrades. The

nominate subspecies In biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. ...

''cinnamomeus'', described by Gould from the northwestern Himalayas, breeds from northern

Arunachal Pradesh Arunachal Pradesh (; ) is a States and union territories of India, state in northeast India. It was formed from the North-East Frontier Agency (NEFA) region, and India declared it as a state on 20 February 1987. Itanagar is its capital and la ...

Nuristan Nuristan, also spelled as Nurestan or Nooristan (Pashto: ; Katë: ), is one of the 34 provinces of Afghanistan, located in the eastern part of the country. It is divided into seven districts and is Afghanistan's least populous province, with a ...

Afghanistan Afghanistan, officially the Islamic Emirate of Afghanistan, is a landlocked country located at the crossroads of Central Asia and South Asia. It is bordered by Pakistan to the Durand Line, east and south, Iran to the Afghanistan–Iran borde ...

Description

The russet sparrow is a small chunky bird, with

warm

in overall colour. It is a medium-sized sparrow at and . It has a thick

suited to eating seeds, which is black on the breeding male, horn-coloured on the non-breeding male, and yellowish with a dark tip on the female. Wingspans for males range between and those for females from . Tail, bill, and tarsus lengths are , , and , respectively. Measurements for the russet sparrow vary geographically, between the three

, and also within the Himalayan subspecies ''cinnamomeus''. The subspecies ''cinnamomeus'' is generally larger than the others, and within this form there is a tendency for birds at higher altitudes to be larger, and a clinal variation in size with the smallest birds in the west of the range and the largest in the east. The iris is

chestnut The chestnuts are the deciduous trees and shrubs in the genus ''Castanea'', in the beech family Fagaceae. The name also refers to the edible nuts they produce. They are native to temperate regions of the Northern Hemisphere. Description ...

in colour. The legs of both sexes are pale brown to pinkish-brown. The flight of all sparrows is swift, and that of the russet sparrow is described as swifter and more direct than that of the

Plumage

The sexes differ, or are dimorphic, in their plumage, and have a similar pattern to that in the corresponding sex of

. There is some variation between the three subspecies, especially in the colouration of the underparts. Birds of the subspecies ''rutilans'' are off-white on their cheeks and the sides of their necks, and have pale grey underparts. Birds of the subspecies ''intensior'' have a pale yellow wash on their underparts and cheeks, as well as darker upperparts, while those of the subspecies ''cinnamomeus'' have a heavy yellow tinge to their underparts.

Moult In biology, moulting (British English), or molting (American English), also known as sloughing, shedding, or in many invertebrates, ecdysis, is a process by which an animal casts off parts of its body to serve some beneficial purpose, either at ...

is poorly recorded, the only records coming from

Sakhalin Sakhalin ( rus, Сахали́н, p=səxɐˈlʲin) is an island in Northeast Asia. Its north coast lies off the southeastern coast of Khabarovsk Krai in Russia, while its southern tip lies north of the Japanese island of Hokkaido. An islan ...

and

Himachal Pradesh Himachal Pradesh (; Sanskrit: ''himācāl prādes;'' "Snow-laden Mountain Province") is a States and union territories of India, state in the northern part of India. Situated in the Western Himalayas, it is one of the thirteen Indian Himalayan ...

. In Sakhalin, moult occurs in August and September, between the breeding season and migration. In Himachal Pradesh, aviculturalist G. A. Perreau observed captive and wild birds and reported that they were yellow from December to spring and whitish during the remainder of the year, a pattern which may be atypical. The breeding male is bright russet or cinnamon red on its upperparts from its

crown A crown is a traditional form of head adornment, or hat, worn by monarchs as a symbol of their power and dignity. A crown is often, by extension, a symbol of the monarch's government or items endorsed by it. The word itself is used, parti ...

to its rump, with a black streaking on its mantle. It has a small black bib and black around its eye, separated from the russet of its crown by a very thin white

supercilium The supercilium is a plumage feature found on the heads of some bird species. It is a stripe which runs from the base of the bird's beak above its eye, finishing somewhere towards the rear of the bird's head.Dunn and Alderfer (2006), p. 10 Also k ...

, a stripe running from the bill to the rear of the head. The side of its neck and cheek are off-white, and its underparts are pale grey or washed with yellow, varying geographically. Shoulders and greater

coverts A covert feather or tectrix on a bird is one of a set of feathers, called coverts (or ''tectrices''), which cover other feathers. The coverts help to smooth airflow over the wings and tail. Ear coverts The ear coverts are small feathers behind t ...

are chestnut, and its median coverts are black at the base with white at the tips. The rest of the wing is light brown with black tinges. Its tail is blackish brown, edged with ashy brown. The non-breeding male differs little from the breeding male, being paler with more orange upperparts. The only species with which the male is easily confused is the Eurasian tree sparrow, which differs in its black cheek spot and brown back. The female has mostly pale brown upperparts, and pale grey underparts, so it resembles the female house sparrow. It differs from the house sparrow in its slightly darker, russet-tinged plumage. It has a conspicuous cream supercilium from above its eye nearly around its head, and a bold dark brown stripe through its eye. It has mostly greyish brown wings and a back streaked with black and buff. The juvenile is similar to the female, though more pale and sandy. As the male reaches its first winter, it resembles the adult, differing in less bold chestnut and a dusky bib.

Voice

The russet sparrow's vocalisations are described by most sources as "the sweetest and most musical" of any sparrow. Its basic call is a ' or ', similar to that of other sparrows. This call is monosyllabic, unlike the house sparrow's ''chirrup'' call, and is softer than that of the other sparrows. This call is given as a flight call, or by displaying males. Recorded variations include a ' given by males at the nest and a trilled '. Sometimes the male strings calls together and sings them in a strident tone, to create a sort of short song, transcribed as ' or '. The song is interspersed with sibilant ' notes similar to those of the

white wagtail The white wagtail (''Motacilla alba'') is a small passerine bird in the family Motacillidae, which also includes pipits and longclaws. The species breeds in the Palearctic zone in most of Europe and Asia and parts of North Africa; it also has ...

. A thin ' vocalisation not unlike that of an

Indian robin The Indian robin (''Copsychus fulicatus'')Rasmussen & Anderton emend the species epithet from ''fulicata'' to ''fulicatus'' since ''Saxicola'' is masculine and the ''-oides'' ending is always masculine according to ICZN Code 30.1.4.4ICZN Code. Se ...

has been reported, but the context of this call is unrecorded. During territorial disputes, males give a rapid ' call.

Distribution and habitat

The russet sparrow is found in parts of eastern Asia and in the Himalayas. It is not known whether its distribution is continuous between these two areas, since the political sensitivity of the

Brahmaputra River The Brahmaputra is a trans-boundary river which flows through Southwestern China, Northeastern India, and Bangladesh. It is known as Brahmaputra or Luit in Assamese language, Assamese, Yarlung Tsangpo in Lhasa Tibetan, Tibetan, the Siang/Dihan ...

's valley near the China-India border makes the area less accessible to ornithologists. In the Himalayas the russet sparrow breeds from the far northeast of India through southeastern

Tibet Tibet (; ''Böd''; ), or Greater Tibet, is a region in the western part of East Asia, covering much of the Tibetan Plateau and spanning about . It is the homeland of the Tibetan people. Also resident on the plateau are other ethnic groups s ...

Bhutan Bhutan, officially the Kingdom of Bhutan, is a landlocked country in South Asia, in the Eastern Himalayas between China to the north and northwest and India to the south and southeast. With a population of over 727,145 and a territory of , ...

Sikkim Sikkim ( ; ) is a States and union territories of India, state in northeastern India. It borders the Tibet Autonomous Region of China in the north and northeast, Bhutan in the east, Koshi Province of Nepal in the west, and West Bengal in the ...

Nepal Nepal, officially the Federal Democratic Republic of Nepal, is a landlocked country in South Asia. It is mainly situated in the Himalayas, but also includes parts of the Indo-Gangetic Plain. It borders the Tibet Autonomous Region of China Ch ...

Uttarakhand Uttarakhand (, ), also known as Uttaranchal ( ; List of renamed places in India, the official name until 2007), is a States and union territories of India, state in North India, northern India. The state is bordered by Himachal Pradesh to the n ...

, and

Kashmir Kashmir ( or ) is the Northwestern Indian subcontinent, northernmost geographical region of the Indian subcontinent. Until the mid-19th century, the term ''Kashmir'' denoted only the Kashmir Valley between the Great Himalayas and the Pir P ...

and

in Afghanistan. Here, the russet sparrow makes short movements to lower altitudes between November and April in many areas. In eastern Asia, the russet sparrow is found in

, the

Kuril Islands The Kuril Islands or Kurile Islands are a volcanic archipelago administered as part of Sakhalin Oblast in the Russian Far East. The islands stretch approximately northeast from Hokkaido in Japan to Kamchatka Peninsula in Russia, separating the ...

, a small part of mainland Russia, Japan, southern Korea, and part of northern China, where it is mostly migratory. It is distributed through southern China and

, and the mountainous parts of Burma, southern

Northeast India Northeast India, officially the North Eastern Region (NER), is the easternmost region of India representing both a geographic and political Administrative divisions of India, administrative division of the country. It comprises eight States and ...

, Laos, and Vietnam, where it is mostly resident. The russet sparrow also occurs as a winter visitor in southern Japan, the further south of China, and northern Thailand. In eastern Asia, the autumn migration occurs between August and November. The russet sparrow appears to be abundant in most habitats across most of its very large range, and in some areas it is among the most common birds. In Southeast Asia, its range has contracted at lower elevations due to

global warming Present-day climate change includes both global warming—the ongoing increase in global average temperature—and its wider effects on Earth's climate system. Climate change in a broader sense also includes previous long-term changes ...

, but it has also moved higher at high elevations and it remains common. Although its global population has not been quantified, it is assessed for the

IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is an inventory of the global conservation status and extinction risk of biological ...

least concern A least-concern species is a species that has been evaluated and categorized by the International Union for Conservation of Nature (IUCN) as not being a focus of wildlife conservation because the specific species is still plentiful in the wil ...

for global extinction. Breeding takes place in

mountain A mountain is an elevated portion of the Earth's crust, generally with steep sides that show significant exposed bedrock. Although definitions vary, a mountain may differ from a plateau in having a limited summit area, and is usually higher t ...

s and uplands across most of the range. This preference for high altitudes is influenced by latitude: in the farthest south of its range it never breeds below

above mean sea level Height above mean sea level is a measure of a location's vertical distance (height, elevation or altitude) in reference to a vertical datum based on a historic mean sea level. In geodesy, it is formalized as orthometric height. The zero level ...

, but in the northernmost part of its range it often breeds by the sea. In eastern Asia, the russet sparrow prefers light

, but it is sometimes found in towns and agricultural areas. In Sakhalin, it breeds mostly in

riparian forest A riparian forest or riparian woodland is a forested or wooded area of land adjacent to a body of water such as a river, stream, pond, lake, marshland, estuary, canal, Sink (geography), sink, or reservoir. Due to the broad nature of the definitio ...

. In

Hokkaidō is the second-largest island of Japan and comprises the largest and northernmost prefecture, making up its own region. The Tsugaru Strait separates Hokkaidō from Honshu; the two islands are connected by railway via the Seikan Tunnel. The ...

, the russet sparrow finds a greater food supply for its young in more remote forests, and has greater reproductive success. In the Himalayas, it is strongly associated with terrace cultivation, and it probably only spread to the Himalayas when these agricultural practices arrived 3000 to 4000 years ago. In towns where it occurs alongside the house or tree sparrow, it is found in gardens and less built up areas. In those

hill station A hill station is a touristic town located at a higher elevation than the nearby plain or valley. The English term was originally used mostly in Western imperialism in Asia, colonial Asia, but also in Africa (albeit rarely), for towns founded by ...

s in India where both the house and the russet sparrow occur, the house sparrow breeds around more-built up areas and

bazaar A bazaar or souk is a marketplace consisting of multiple small Market stall, stalls or shops, especially in the Middle East, the Balkans, Central Asia, North Africa and South Asia. They are traditionally located in vaulted or covered streets th ...

s, whereas the russet sparrow is "rather more up market at the larger houses with gardens and open spaces". In towns where it is the only sparrow, it is not dependent on trees, breeding around houses and eating scraps in streets. In the winter, migratory birds occur in open cultivated land and riverine

grassland A grassland is an area where the vegetation is dominance (ecology), dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush (Juncaceae) can also be found along with variable proportions of legumes such as clover, and other Herbaceo ...

s, but are never far from shrubs or trees.

Behaviour

In many aspects of its behaviour, the russet sparrow is similar to the house and Eurasian tree sparrows. Like them, it feeds on the ground, but spends most of its time perching on branches. Unlike those species, it prefers open, exposed branches for perching. The russet sparrow is described as shy and wary by some observers, but J. Denis Summers-Smith found it approachable in Indian hill stations. Flocking russet sparrows feed close to the ground, moving forward as birds from the rear of a flock move to the front, in what is called "roller feeding". Outside its breeding season, the russet sparrow is gregarious and forms flocks to find food, though it infrequently associates with other birds. Wintering flocks tend to keep away from human habitation. The russet sparrow is also social at night during the winter, and it forms large communal roosts in trees and bushes. In the breeding season, the female roosts in the nests and the male nests in foliage nearby. The adult russet sparrow is mostly a seed-eater, eating the seeds of herbs and weeds as well as

rice Rice is a cereal grain and in its Domestication, domesticated form is the staple food of over half of the world's population, particularly in Asia and Africa. Rice is the seed of the grass species ''Oryza sativa'' (Asian rice)—or, much l ...

barley Barley (), a member of the grass family, is a major cereal grain grown in temperate climates globally. It was one of the first cultivated grains; it was domesticated in the Fertile Crescent around 9000 BC, giving it nonshattering spikele ...

, and other

grains A grain is a small, hard, dry fruit ( caryopsis) – with or without an attached hull layer – harvested for human or animal consumption. A grain crop is a grain-producing plant. The two main types of commercial grain crops are cereals and le ...

Berries A berry is a small, pulpy, and often edible fruit. Typically, berries are juicy, rounded, brightly colored, sweet, sour or tart, and do not have a stone fruit, stone or pit (fruit), pit although many wikt:pip#Etymology 2, pips or seeds may be p ...

, such as those of the kingore (certain ''

Berberis ''Berberis'' (), commonly known as barberry, is a large genus of deciduous and evergreen shrubs from tall, found throughout temperate and subtropical regions of the world (apart from Australia). Species diversity is greatest in South America a ...

'' spp.), are also eaten when available. Nestlings are fed mostly on

s, especially

caterpillar Caterpillars ( ) are the larval stage of members of the order Lepidoptera (the insect order comprising butterflies and moths). As with most common names, the application of the word is arbitrary, since the larvae of sawflies (suborder ...

s and

larva A larva (; : larvae ) is a distinct juvenile form many animals undergo before metamorphosis into their next life stage. Animals with indirect development such as insects, some arachnids, amphibians, or cnidarians typically have a larval phase ...

beetle Beetles are insects that form the Taxonomic rank, order Coleoptera (), in the superorder Holometabola. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 40 ...

s obtained on trees and flying insects caught by aerial pursuit. During the breeding season adults also consume insects. Mortality rates in the russet sparrow have not been studied, but it is known that many young birds die from '' Isospora'' infections, to which the bird has little resistance. Other recorded parasites of the russet sparrow include '' Protocalliphora'' blowflies, and '' Menacanthus''

chewing lice The Mallophaga are a possibly paraphyletic section of lice, known as chewing lice, biting lice, or bird lice, containing more than 3000 species. These lice are external parasites that feed mainly on birds, although some species also feed on mamma ...

Breeding

The russet sparrow's

is short, lasting about three months. The Himalayan subspecies ''cinnamomeus'' has been recorded breeding from April to August, the nominate subspecies breeds from May to July, and the subspecies ''intensior'' is believed to breed in March. During its breeding season, the russet sparrow is not gregarious, and its nests are dispersed uniformly rather than clustered. Its nests are most frequently built in tree cavities, often disused

woodpecker Woodpeckers are part of the bird family (biology), family Picidae, which also includes the piculets, wrynecks and sapsuckers. Members of this family are found worldwide, except for Australia, New Guinea, New Zealand, Madagascar and the extreme ...

nests. The russet sparrow has been recorded breeding alongside the Eurasian tree sparrow and

black kite The black kite (''Milvus migrans'') is a medium-sized bird of prey in the family Accipitridae, which also includes many other diurnal raptors. It is thought to be the world's most abundant species of Accipitridae, although some populations have ...

nests, taking advantage of the territorial defence of the kites to avoid nest predation. Other nest locations include the eaves of thatched roofs, stone walls and embankments, and electric junction boxes. In

, it nests in holes in the outer walls of monasteries, often alongside the Eurasian tree sparrow. In Sakhalin, it will sometimes build free-standing nests in bushes. The russet sparrow also uses the disused nests of the

red-rumped swallow The European red-rumped swallow (''Cecropis rufula'') is a small passerine bird in the swallow family Hirundinidae. It breeds in open hilly country of southern Europe and north Africa east to Iran, Pakistan and northwest India. During winter it m ...

, and a pair of russet sparrows has been recorded attempting to oust a pair of

black-crested tit The black-crested tit (''Periparus ater melanolophus''), also known as the spot-winged tit, is a bird in the family Paridae. It was formerly considered a species, but is now widely considered a subspecies of the coal tit. Distribution and habit ...

s from their nest. The male chooses the nest site, and uses it for his

, spending much time calling nearby. When a female comes near a male at his nest the male begins to display by raising his head, drooping his wings, pushing his chest forward, and lowering his tail. He then bows up and down in front of the female, who will lunge and then fly away if unreceptive. Both sexes take part in building the nest, which consists of a loose, untidy bunch of dry grass which fills the nesting cavity, lined with fur and feathers for warmth.

Eggs An egg is an organic vessel in which an embryo begins to develop. Egg, EGG or eggs may also refer to: Biology * Egg cell, the female reproductive cell (gamete) in oogamous organisms Food * Eggs as food Places * Egg, Austria * Egg, Switzerland ...

are elongated ovals, with a fine texture and a slight gloss. They are whitish in overall colour with a grey tinge and brown spots, streaks, or blotches. The average size of the egg is 19.2×14.2 millimetres (0.75×0.55 in). Eggs are similar to those of the Eurasian tree sparrow, differing in a duller colour and more narrow shape, though they cannot be separated from those of the other sparrows with certainty. Two

clutches A clutch is a mechanical device that allows an output shaft to be disconnected from a rotating input shaft. The clutch's input shaft is typically attached to a motor, while the clutch's output shaft is connected to the mechanism that does the ...

of four or sometimes five or six eggs are laid in a year. In Hokkaidō, clutches are laid between early May and early July, with two peaks in laying around mid May and late June. Both sexes incubate and feed the young, with the male often being more active in feeding the nestlings. In Hokkaidō, nestlings hatch weighing about , and fledge 14 or 15 days after hatching, weighing . The

common cuckoo The cuckoo, common cuckoo, European cuckoo or Eurasian cuckoo (''Cuculus canorus'') is a member of the cuckoo order of birds, Cuculiformes, which includes the Geococcyx, roadrunners, the ani (bird), anis and the coucals. This species is a widesp ...

has been recorded in old literature as a

brood parasite Brood may refer to: Nature * Brood, a collective term for offspring * Brooding, the incubation of bird eggs by their parents * Bee brood, the young of a beehive * Individual broods of North American periodical cicadas: ** Brood X, the largest ...

of the russet sparrow.

Relationships with humans

In parts of the range, the russet sparrow inhabits towns, and in most of its range, it occurs near cultivation, and is a minor pest of agriculture. Though it damages crops, it also feeds its nestlings largely on insect pests. In China, the russet sparrow has been recorded as a captive bird, kept with the Eurasian tree sparrow. In Japan it was eaten in the 1870s and sold in the Yokohama game market. The russet sparrow is known well enough in the Himalayas that in most languages it has a different vernacular name from the Eurasian tree sparrow. Examples of these vernacular names include in

Hindi Modern Standard Hindi (, ), commonly referred to as Hindi, is the Standard language, standardised variety of the Hindustani language written in the Devanagari script. It is an official language of India, official language of the Government ...

and in Tibetan. The Japanese artist

Hokusai , known mononymously as Hokusai, was a Japanese ukiyo-e artist of the Edo period, active as a painter and printmaker. His woodblock printing in Japan, woodblock print series ''Thirty-Six Views of Mount Fuji'' includes the iconic print ''The Gr ...

portrayed the russet sparrow, and due to this, it has appeared on

postage stamp A postage stamp is a small piece of paper issued by a post office, postal administration, or other authorized vendors to customers who pay postage (the cost involved in moving, insuring, or registering mail). Then the stamp is affixed to the f ...

s featuring Japanese art in Japan,

The Gambia The Gambia, officially the Republic of The Gambia, is a country in West Africa. Geographically, The Gambia is the List of African countries by area, smallest country in continental Africa; it is surrounded by Senegal on all sides except for ...

, and

Guyana Guyana, officially the Co-operative Republic of Guyana, is a country on the northern coast of South America, part of the historic British West Indies. entry "Guyana" Georgetown, Guyana, Georgetown is the capital of Guyana and is also the co ...

References

;Works cited * * * * * * * * * * * * * * * * * *

External links

Russet sparrow
at the Internet Bird Collection
Recording of the russet sparrow's calls
{{Taxonbar, from1=Q1316962, from2=Q28868046

russet sparrow The russet sparrow (''Passer cinnamomeus''), also called the cinnamon or cinnamon tree sparrow, is a passerine bird of the Old World sparrow, sparrow family (biology), family Passeridae. A stout little seed-eating bird with a thick beak, bill, ...

Birds of the Himalayas Birds of Eastern Himalaya Birds of East Asia