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The Javan rusa or Sunda sambar (''Rusa timorensis'') is a large
deer A deer (: deer) or true deer is a hoofed ruminant ungulate of the family Cervidae (informally the deer family). Cervidae is divided into subfamilies Cervinae (which includes, among others, muntjac, elk (wapiti), red deer, and fallow deer) ...
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
native Native may refer to: People * '' Jus sanguinis'', nationality by blood * '' Jus soli'', nationality by location of birth * Indigenous peoples, peoples with a set of specific rights based on their historical ties to a particular territory ** Nat ...
to
Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania, between the Indian Ocean, Indian and Pacific Ocean, Pacific oceans. Comprising over List of islands of Indonesia, 17,000 islands, including Sumatra, ...
and
East Timor Timor-Leste, also known as East Timor, officially the Democratic Republic of Timor-Leste, is a country in Southeast Asia. It comprises the eastern half of the island of Timor, the coastal exclave of Oecusse in the island's northwest, and ...
. Introduced populations exist in a wide variety of locations in the Southern Hemisphere. ''Rusa'' is the Malay word for "deer" in general as well as lending to name of its own genus.


Taxonomy

Seven subspecies of the Javan rusa are recognised: *''R. t. timorensis'' (Timor rusa deer) –
Timor Timor (, , ) is an island at the southern end of Maritime Southeast Asia, in the north of the Timor Sea. The island is Indonesia–Timor-Leste border, divided between the sovereign states of Timor-Leste in the eastern part and Indonesia in the ...
. *''R. t. djonga'' – Muna and Butung Islands. *''R. t. floresiensis'' (Flores rusa deer) –
Flores Flores is one of the Lesser Sunda Islands, a group of islands in the eastern half of Indonesia. Administratively, it forms the largest island in the East Nusa Tenggara Province. The area is 14,250 km2. Including Komodo and Rinca islands ...
and other islands. *''R. t. macassaricus'' (Celebes rusa deer) –
Sulawesi Sulawesi ( ), also known as Celebes ( ), is an island in Indonesia. One of the four Greater Sunda Islands, and the List of islands by area, world's 11th-largest island, it is situated east of Borneo, west of the Maluku Islands, and south of Min ...
. *''R. t. moluccensis'' (Moluccan rusa deer) –
Maluku Islands The Maluku Islands ( ; , ) or the Moluccas ( ; ) are an archipelago in the eastern part of Indonesia. Tectonics, Tectonically they are located on the Halmahera Plate within the Molucca Sea Collision Zone. Geographically they are located in West ...
. *''R. t. renschi'' –
Bali Bali (English:; Balinese language, Balinese: ) is a Provinces of Indonesia, province of Indonesia and the westernmost of the Lesser Sunda Islands. East of Java and west of Lombok, the province includes the island of Bali and a few smaller o ...
. *''R. t. russa'' (Javan rusa deer) –
Java Java is one of the Greater Sunda Islands in Indonesia. It is bordered by the Indian Ocean to the south and the Java Sea (a part of Pacific Ocean) to the north. With a population of 156.9 million people (including Madura) in mid 2024, proje ...
.


Characteristics

The Javan rusa is dark blackish brown and has a gray forehead. Its back is almost black, the underparts and inner thighs are yellowish brown. The abdomen is lighter brown, and the tail tuft is dark blackish brown. The hair is coarse and longer on the chest than on the remaining body. Its ears are wide and a little shorter than the head. The
antler Antlers are extensions of an animal's skull found in members of the Cervidae (deer) Family (biology), family. Antlers are a single structure composed of bone, cartilage, fibrous tissue, skin, nerves, and blood vessels. They are generally fo ...
s are medium long and rather wide, the upper branch points forward.
Fawn A deer (: deer) or true deer is a hoofed ruminant ungulate of the family (biology), family Cervidae (informally the deer family). Cervidae is divided into subfamilies Cervinae (which includes, among others, muntjac, elk (wapiti), red deer, a ...
s are born without spots. Males are bigger than females; head-to-body length varies from , with a tail. Males weigh , female about .


Distribution and habitat

The Javan rusa natively occurs on the islands of Java, Bali and
Timor Timor (, , ) is an island at the southern end of Maritime Southeast Asia, in the north of the Timor Sea. The island is Indonesia–Timor-Leste border, divided between the sovereign states of Timor-Leste in the eastern part and Indonesia in the ...
in Indonesia. It has been introduced to
Irian Jaya New Guinea (; Hiri Motu: ''Niu Gini''; , fossilized , also known as Papua or historically ) is the world's second-largest island, with an area of . Located in Melanesia in the southwestern Pacific Ocean, the island is separated from Austral ...
,
Borneo Borneo () is the List of islands by area, third-largest island in the world, with an area of , and population of 23,053,723 (2020 national censuses). Situated at the geographic centre of Maritime Southeast Asia, it is one of the Greater Sunda ...
, the
Lesser Sunda Islands The Lesser Sunda Islands (, , ), now known as Nusa Tenggara Islands (, or "Southeast Islands"), are an archipelago in the Indonesian archipelago. Most of the Lesser Sunda Islands are located within the Wallacea region, except for the Bali pro ...
, Maluku, Sulawesi,
Pohnpei Pohnpei (formerly known as Ponape or Ascension, from Pohnpeian: "upon (''pohn'') a stone altar (''pei'')") is an island of the Senyavin Islands which are part of the larger Caroline Islands group. It belongs to Pohnpei State, one of the fou ...
,
Mauritius Mauritius, officially the Republic of Mauritius, is an island country in the Indian Ocean, about off the southeastern coast of East Africa, east of Madagascar. It includes the main island (also called Mauritius), as well as Rodrigues, Ag ...
,
Réunion Réunion (; ; ; known as before 1848) is an island in the Indian Ocean that is an overseas departments and regions of France, overseas department and region of France. Part of the Mascarene Islands, it is located approximately east of the isl ...
,
Fiji Fiji, officially the Republic of Fiji, is an island country in Melanesia, part of Oceania in the South Pacific Ocean. It lies about north-northeast of New Zealand. Fiji consists of an archipelago of more than 330 islands—of which about ...
,
Tonga Tonga, officially the Kingdom of Tonga, is an island country in Polynesia, part of Oceania. The country has 171 islands, of which 45 are inhabited. Its total surface area is about , scattered over in the southern Pacific Ocean. accordin ...
,
Samoa Samoa, officially the Independent State of Samoa and known until 1997 as Western Samoa, is an island country in Polynesia, part of Oceania, in the South Pacific Ocean. It consists of two main islands (Savai'i and Upolu), two smaller, inhabited ...
,
Vanuatu Vanuatu ( or ; ), officially the Republic of Vanuatu (; ), is an island country in Melanesia located in the South Pacific Ocean. The archipelago, which is of volcanic origin, is east of northern Australia, northeast of New Caledonia, east o ...
, the
Solomon Islands Solomon Islands, also known simply as the Solomons,John Prados, ''Islands of Destiny'', Dutton Caliber, 2012, p,20 and passim is an island country consisting of six major islands and over 1000 smaller islands in Melanesia, part of Oceania, t ...
, the
Christmas Island Christmas Island, officially the Territory of Christmas Island, is an States and territories of Australia#External territories, Australian external territory in the Indian Ocean comprising the island of the same name. It is about south o ...
, the
Cocos Islands The Cocos (Keeling) Islands (), officially the Territory of Cocos (Keeling) Islands (; ), are an Australian external territory in the Indian Ocean, comprising a small archipelago approximately midway between Australia and Sri Lanka and rel ...
,
Nauru Nauru, officially the Republic of Nauru, formerly known as Pleasant Island, is an island country and microstate in the South Pacific Ocean. It lies within the Micronesia subregion of Oceania, with its nearest neighbour being Banaba (part of ...
,
Mainland Australia Mainland Australia is the main landmass of the Australia (continent), Australian continent, excluding the Aru Islands Regency, Aru Islands, New Guinea, Tasmania, and other list of islands of Australia, Australian offshore islands. The landmass ...
,
New Caledonia New Caledonia ( ; ) is a group of islands in the southwest Pacific Ocean, southwest of Vanuatu and east of Australia. Located from Metropolitan France, it forms a Overseas France#Sui generis collectivity, ''sui generis'' collectivity of t ...
,
New Zealand New Zealand () is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and List of islands of New Zealand, over 600 smaller islands. It is the List of isla ...
,
Papua New Guinea Papua New Guinea, officially the Independent State of Papua New Guinea, is an island country in Oceania that comprises the eastern half of the island of New Guinea and offshore islands in Melanesia, a region of the southwestern Pacific Ocean n ...
,
New Britain New Britain () is the largest island in the Bismarck Archipelago, part of the Islands Region of Papua New Guinea. It is separated from New Guinea by a northwest corner of the Solomon Sea (or with an island hop of Umboi Island, Umboi the Dampie ...
, and New Ireland. The Javan rusa was introduced by the Dutch to
New Guinea New Guinea (; Hiri Motu: ''Niu Gini''; , fossilized , also known as Papua or historically ) is the List of islands by area, world's second-largest island, with an area of . Located in Melanesia in the southwestern Pacific Ocean, the island is ...
in the early 1900s. Since its introduction to the West Papuan lowlands, the species has become widely dispersed and is common in much of its new range. However its population in its native range has declined markedly by approximately 10,000 individuals in the past two decades and likely faces further decline. As a result it has been listed as vulnerable in its native range and was declared as a protected species under Indonesian law in 2018. During the 1980s and 1990s, protected areas such as national parks were used to control poaching and the effects of land conversion that destroyed grazing areas. However large numbers of Javan rusa died in
Baluran National Park Baluran National Park is located in Situbondo Regency, East Java, Indonesia. It has a relatively dry climate and mainly consists of savanna (40%), as well as lowland forests, mangrove forests and hills, with Mount Baluran (1,247m) as its highes ...
in Indonesia due to the loss of grazing area as a result of the invasive
thorny acacia ''Vachellia nilotica'', more commonly known as ''Acacia nilotica'', and by the vernacular names of gum arabic tree, babul, thorn mimosa, Egyptian acacia or thorny acacia, is a flowering tree in the family Fabaceae. It is native to Africa, the Mid ...
.


Ecology

Javan rusa are nocturnal, although they do graze during the day. They are rarely seen in the open and are very difficult to approach due to their keen senses and cautious instincts. The rusa deer is often found in small groups or pairs, although males are often seen alone. When alarmed, a rusa stag lets out an extremely loud honk. This is an alarm call and alerts any other deer in the vicinity. As with other deer species, Javan rusa mainly feed on grass, leaves, and fallen fruit. Most of their fluid requirements are met by the food they consume, so they hardly drink water.


Predators

The main predators of the Javan rusa includes
Javan leopard The Javan leopard (''Panthera pardus melas'') is a leopard subspecies confined to the Indonesian island of Java. It has been listed as Endangered on the IUCN Red List since 2021. The population is estimated at 188–571 mature individuals in 22 ...
,
Sunda clouded leopard The Sunda clouded leopard (''Neofelis diardi'') is a medium-sized wild cat native to Borneo and Sumatra. It is listed as Vulnerable on the IUCN Red List since 2015, as the total effective population probably consists of fewer than 10,000 mature ...
,
dhole The dhole ( ; ''Cuon alpinus'') is a canid native to South, East and Southeast Asia. It is anatomically distinguished from members of the genus ''Canis'' in several aspects: its skull is convex rather than concave in profile, it lacks a third ...
,
estuarine crocodile The saltwater crocodile (''Crocodylus porosus'') is a crocodilian native to saltwater habitats, brackish wetlands and freshwater rivers from India's east coast across Southeast Asia and the Sundaland to northern Australia and Micronesia. It h ...
,
reticulated python The reticulated python (''Malayopython reticulatus'') is a Pythonidae, python species native to South Asia, South and Southeast Asia. It is the world's List of largest snakes, longest snake, and the list of largest snakes, third heaviest snake. I ...
, and
Komodo dragon The Komodo dragon (''Varanus komodoensis''), also known as the Komodo monitor, is a large reptile of the monitor lizard family Varanidae that is endemic to the Indonesian islands of Komodo (island), Komodo, Rinca, Flores, Gili Dasami, and Gili ...
on the islands of
Rinca Rinca is a small island near Komodo and Flores island, East Nusa Tenggara, Indonesia, within the West Manggarai Regency. It is one of the three largest islands included in Komodo National Park. The island is famous for Komodo dragons, giant l ...
, Komodo, and Flores.


Reproduction

The Javan rusa mates around July and August, when stags contest by calling in a loud, shrill bark and dueling with the antlers. The doe gives birth to one or two calves after a gestation period of 8 months, at the start of spring. Calves are weaned at 6–8 months, and sexual maturity is attained at 3–5 years, depending on habitat conditions. Javan rusas live 15–20 years both in the wild and in captivity.


References


External links

* * {{Taxonbar, from=Q663412
Javan rusa The Javan rusa or Sunda sambar (''Rusa timorensis'') is a large deer species native to Indonesia and East Timor. Introduced populations exist in a wide variety of locations in the Southern Hemisphere. ''Rusa'' is the Malay word for "deer" in ...
Mammals of Indonesia Fauna of Java Mammals of Borneo Vulnerable fauna of Asia
Javan rusa The Javan rusa or Sunda sambar (''Rusa timorensis'') is a large deer species native to Indonesia and East Timor. Introduced populations exist in a wide variety of locations in the Southern Hemisphere. ''Rusa'' is the Malay word for "deer" in ...
Mammals of New Guinea