Roystonea
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''Roystonea'' is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of eleven
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
of
monoecious Monoecy (; adj. monoecious ) is a sexual system in seed plants where separate male and female cones or flowers are present on the same plant. It is a monomorphic sexual system comparable with gynomonoecy, andromonoecy and trimonoecy, and contras ...
palms, native to the
Neotropics The Neotropical realm is one of the eight biogeographic realms constituting Earth's land surface. Physically, it includes the tropical terrestrial ecoregions of the Americas and the entire South American temperate zone. Definition In biogeog ...
, in the
Caribbean The Caribbean ( , ; ; ; ) is a region in the middle of the Americas centered around the Caribbean Sea in the Atlantic Ocean, North Atlantic Ocean, mostly overlapping with the West Indies. Bordered by North America to the north, Central America ...
, the adjacent coasts of
Florida Florida ( ; ) is a U.S. state, state in the Southeastern United States, Southeastern region of the United States. It borders the Gulf of Mexico to the west, Alabama to the northwest, Georgia (U.S. state), Georgia to the north, the Atlantic ...
in the
United States The United States of America (USA), also known as the United States (U.S.) or America, is a country primarily located in North America. It is a federal republic of 50 U.S. state, states and a federal capital district, Washington, D.C. The 48 ...
,
Mexico Mexico, officially the United Mexican States, is a country in North America. It is the northernmost country in Latin America, and borders the United States to the north, and Guatemala and Belize to the southeast; while having maritime boundar ...
,
Central America Central America is a subregion of North America. Its political boundaries are defined as bordering Mexico to the north, Colombia to the southeast, the Caribbean to the east, and the Pacific Ocean to the southwest. Central America is usually ...
and northern
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a considerably smaller portion in the Northern Hemisphere. It can also be described as the southern Subregion#Americas, subregion o ...
.Kew World Checklist of Selected Plant Families
/ref> Commonly known as the royal palms, the genus was named after Roy Stone, a
U.S. Army The United States Army (USA) is the primary land service branch of the United States Department of Defense. It is designated as the Army of the United States in the United States Constitution.Article II, section 2, clause 1 of the United Stat ...
engineer Engineers, as practitioners of engineering, are professionals who Invention, invent, design, build, maintain and test machines, complex systems, structures, gadgets and materials. They aim to fulfill functional objectives and requirements while ...
. It contains some of the most recognizable and commonly cultivated palms of tropical and subtropical regions.


Description

''Roystonea'' is a genus of large, unarmed, single-stemmed palms with
pinnate Pinnation (also called pennation) is the arrangement of feather-like or multi-divided features arising from both sides of a common axis. Pinnation occurs in biological morphology, in crystals, such as some forms of ice or metal crystals, and ...
leaves. The large stature and striking appearance of a ''Roystonea'' palm makes it a notable aspect of the landscape. The stems, which were compared to stone columns by
Louis Louis may refer to: People * Louis (given name), origin and several individuals with this name * Louis (surname) * Louis (singer), Serbian singer Other uses * Louis (coin), a French coin * HMS ''Louis'', two ships of the Royal Navy See also ...
and Elizabeth Agassiz in 1868, are smooth and columnar, although the trunks of '' R. altissima'' and '' R. maisiana'' are more slender than those of typical royal palms. Stems often are swollen and bulging along portions of their length, which may reflect years where growing conditions were better or worse than average. Leaf scars are often prominent along the stem, especially in young, rapidly growing individuals. Stem color ranges from gray-white to gray-brown except in '' R. violacea'', which have violet-brown or mauve stems. Royal palm, '' R. oleracea'', reaches heights of , but most species are in the range. The largest Royal palm is located in
Floresta Estadual Edmundo Navarro de Andrade Floresta may refer to: Geography * Floresta, Paraná, a city in the state of Paraná, Brazil * Floresta, Pernambuco, a city in the state of Pernambuco, Brazil * Floresta, Rio Grande do Sul Floresta (meaning ''Forest'' in Portuguese) is a neigh ...
in
Rio Claro, São Paulo Rio Claro is a city in the state of São Paulo in Brazil. The elevation is 613 m. It was incorporated as the village of '' São João Batista do Ribeirão Claro'' in 1827, and this incorporation is celebrated every year on June 24 as a municipal h ...
, Brazil with 42.4 m and was discovered by Vincent Ferh and Mauro Galetti ''Roystonea'' leaves consist of a sheathing leaf base, a
petiole Petiole may refer to: *Petiole (botany), the stalk of a leaf, attaching the blade to the stem *Petiole (insect anatomy) In entomology, petiole is the technical term for the narrow waist of some hymenopteran insects, especially ants, bees, and ...
, and a
rachis In biology, a rachis (from the [], "backbone, spine") is a main axis or "shaft". In zoology and microbiology In vertebrates, ''rachis'' can refer to the series of articulated vertebrae, which encase the spinal cord. In this case the ''rachi ...
. The leaf base forms a distinctive green sheath around the uppermost portion of the trunk. Known as the crownshaft, this sheath extends down the trunk. The petiole connects the lead base with the rachis. The American botanist Scott Zona only reported petiole lengths for three of the 10 species, ranging from . The rachis is pinnately divided and ranges from long. The leaf segments themselves range in length from in ''R. altissima'' up to as much as in '' R. lenis''. They are arranged in two or three planes along the rachis. Many authors have reported that the leaves ''R. oleracea'' are arranged in a single plane, but Zona reported that this is not the case. These plants have the ability to easily release their leaves in strong winds, a supposed adaption serving to prevent toppling during
hurricane A tropical cyclone is a rapidly rotating storm system with a low-pressure area, a closed low-level atmospheric circulation, strong winds, and a spiral arrangement of thunderstorms that produce heavy rain and squalls. Depending on its ...
s.
Inflorescence In botany, an inflorescence is a group or cluster of flowers arranged on a plant's Plant stem, stem that is composed of a main branch or a system of branches. An inflorescence is categorized on the basis of the arrangement of flowers on a mai ...
s occur beneath the crownshaft, emerging from a narrow, horn-shaped
bract In botany, a bract is a modified or specialized leaf, associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves in size, color, shape or texture. They also lo ...
. The
flower Flowers, also known as blooms and blossoms, are the reproductive structures of flowering plants ( angiosperms). Typically, they are structured in four circular levels, called whorls, around the end of a stalk. These whorls include: calyx, m ...
s on the branched
panicle In botany, a panicle is a much-branched inflorescence. (softcover ). Some authors distinguish it from a compound spike inflorescence, by requiring that the flowers (and fruit) be pedicellate (having a single stem per flower). The branches of a p ...
s are usually white,
unisexual Dioecy ( ; ; adj. dioecious, ) is a characteristic of certain species that have distinct unisexual individuals, each producing either male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproduction is ...
, and contain both sexes. The
fruit In botany, a fruit is the seed-bearing structure in flowering plants (angiosperms) that is formed from the ovary after flowering. Fruits are the means by which angiosperms disseminate their seeds. Edible fruits in particular have long propaga ...
is an oblong or globose
drupe In botany, a drupe (or stone fruit) is a type of fruit in which an outer fleshy part (exocarp, or skin, and mesocarp, or flesh) surrounds a single shell (the ''pip'' (UK), ''pit'' (US), ''stone'', or ''pyrena'') of hardened endocarp with a seed ...
long and deep purple when ripe. Some species so closely resemble one another that scientific differentiation is by inflorescence detail; flower size, colour, etc.


Taxonomy

''Roystonea'' is placed in the
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end botanical subfamily names with "-oideae", and zo ...
Arecoideae and the
tribe The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide use of the term in English is in the discipline of anthropology. The definition is contested, in part due to conflict ...
Roystoneae, which only contains ''Roystonea''. The placement of ''Roystonea'' within the Arecoideae is uncertain; a
phylogeny A phylogenetic tree or phylogeny is a graphical representation which shows the evolutionary history between a set of species or Taxon, taxa during a specific time.Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, M ...
based on
plastid A plastid is a membrane-bound organelle found in the Cell (biology), cells of plants, algae, and some other eukaryotic organisms. Plastids are considered to be intracellular endosymbiotic cyanobacteria. Examples of plastids include chloroplasts ...
DNA failed to resolve the position of the genus within the Arecoideae. As of 2008, there appear to be no molecular phylogenetic studies of ''Roystonea''. One species is known only from two
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
ized flowers preserved in
Dominican amber Dominican amber is amber from the Dominican Republic derived from resin of the extinct tree '' Hymenaea protera''. Dominican amber differentiates itself from Baltic amber by being nearly always transparent, and it has a higher number of fossil in ...
which were described in 2002.


Species

Accepted species:Carnevali, G., J. L. Tapia-Muñoz, R. Duno de Stefano & I. M. Ramírez Morillo. 2010. Flora Ilustrada de la Peninsula Yucatán: Listado Florístico 1–326


Distribution

''Roystonea'' has a circum-Caribbean distribution which ranges from southern Florida in the north, to southern Mexico, Honduras and Nicaragua in the east and Venezuela and Colombia in the south. Species are found throughout the Caribbean, although only Jamaica and Hispaniola (with two native species) and Cuba (with five native species) have more than one native species. A few species are planted in
Tunisia Tunisia, officially the Republic of Tunisia, is a country in the Maghreb region of North Africa. It is bordered by Algeria to the west and southwest, Libya to the southeast, and the Mediterranean Sea to the north and east. Tunisia also shares m ...
as well.


Uses

Royal palms are widely planted for decorative purposes throughout their native region, and elsewhere in the
tropics The tropics are the regions of Earth surrounding the equator, where the sun may shine directly overhead. This contrasts with the temperate or polar regions of Earth, where the Sun can never be directly overhead. This is because of Earth's ax ...
and
subtropics The subtropical zones or subtropics are geographical and climate zones immediately to the north and south of the tropics. Geographically part of the temperate zones of both hemispheres, they cover the middle latitudes from to approximately ...
. Royal palms are very fond of water and thrive on supplemental
irrigation Irrigation (also referred to as watering of plants) is the practice of applying controlled amounts of water to land to help grow crops, landscape plants, and lawns. Irrigation has been a key aspect of agriculture for over 5,000 years and has bee ...
. They also do better in a soil with much
humus In classical soil science, humus is the dark organic matter in soil that is formed by the decomposition of plant and animal matter. It is a kind of soil organic matter. It is rich in nutrients and retains moisture in the soil. Humus is the Lati ...
. Though mainly a decorative plant, royal palms do have some minor agricultural uses. The heart of the palm is used to make salad in some parts of the Caribbean, and its seeds can be used as substitutes for coffee beans. Royal palm seeds were widely used in Cuba to feed pigs at least up to the 1940s and 1950s. The meat of pigs raised with royal palm seeds was said to be the very best. The lard obtained from pigs fattened or raised with royal palm seeds was said to exhibit a grainy texture, and by inference, to have been the best lard to consume. The seeds generally were obtained by men who specialized in climbing the royal palms using a set of two ropes looped around the stem, with two loops around the climber's legs to support himself. Once the climber reached the seed clumps, he would tie the clumps that were mature, cut them, and let them down by a rope supported from other seed clumps.


References


External links


Kew Palms Checklist: ''Roystonea''

Flora of North America: ''Roystonea''
* {{Taxonbar, from1=Q309496, from2=Q19450725 Arecaceae genera Flora of the Neotropical realm National symbols of Haiti