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The rockhopper penguins are three closely related
taxa In biology, a taxon (back-formation from ''taxonomy''; : taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular name and ...
of crested penguins that have been traditionally treated as a single
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
and are sometimes split into three species. Not all experts agree on the classification of these penguins. Some consider all three as distinct species, some split the western and eastern forms into the southern rockhopper penguin and keep the northern rockhopper as distinct, while other experts consider all three potential varieties to be one species.


Appearance

Rockhopper penguins (Eudyptes chrysocome) are among the smaller species of penguin. After reaching full growth, they are about 20 inches or 50 centimetres in height. Males and females cannot be distinguished visually, so a DNA test is conducted by taking a feather from the bird to determine its gender. Like many penguins, rockhopper penguins have a white belly and the rest of their body is black. Some characteristics that differentiate them from the other penguins are their red eyes, orange beak, pink webbed feet, and the yellow and black spiky feathers they have on their head. Although their yellow and black spiky feathers differentiate them from other penguins, rockhopper penguin chicks do not have them; these feathers develop with age. Their orange beak is initially black, but as the penguins get older, their beaks turn orange. Due to the harsh rocky environment, they cannot slide on their bellies like most penguins, so they hop to get from one place to another, hence their name. The yellow and black ‘hair’ they have on the top of their head is similar to that of the Macaroni penguin (Eudyptes chrysolophus).


Subspecies

What separates rockhopper penguins into subspecies is their location of reproduction and reproductive behaviors. The difference in mating signals found between the subspecies ''E. chrysocome'' (southern) and ''E. moseleyi'' (northern) seems to have occurred quickly, thus these behavioral changes are enough to isolate these taxa. Southern rockhopper penguins are split into two subspecies and they are defined by their location of reproduction. The subspecies ''E. c. filholi'' (eastern) is known to reproduce in the sub-Antarctic around the
Prince Edward Islands The Prince Edward Islands are two small uninhabited subantarctic volcanic islands in the southern Indian Ocean that are administered by South Africa. They are named Marion Island (named after Marc-Joseph Marion du Fresne, 1724–1772) and P ...
,
Crozet Islands The Crozet Islands (; or, officially, ''Archipel Crozet'') are a sub-Antarctic archipelago of small islands in the southern Indian Ocean. They form one of the five administrative districts of the French Southern and Antarctic Lands. History ...
,
Kerguelen Islands The Kerguelen Islands ( or ; in French commonly ' but officially ', ), also known as the Desolation Islands (' in French), are a group of islands in the subantarctic, sub-Antarctic region. They are among the Extremes on Earth#Remoteness, most i ...
,
Heard Island The Territory of Heard Island and McDonald Islands (HIMI) is an Australian external territory comprising a volcanic group of mostly barren Antarctic islands, about two-thirds of the way from Madagascar to Antarctica. The group's overall land ...
,
Macquarie Island Macquarie Island is a subantarctic island in the south-western Pacific Ocean, about halfway between New Zealand and Antarctica. It has been governed as a part of Tasmania, Australia, since 1880. It became a Protected areas of Tasmania, Tasmania ...
and Campbell,
Auckland Auckland ( ; ) is a large metropolitan city in the North Island of New Zealand. It has an urban population of about It is located in the greater Auckland Region, the area governed by Auckland Council, which includes outlying rural areas and ...
and
Antipodes Islands The Antipodes Islands (, ) are inhospitable and uninhabited volcanic islands in subantarctic waters to the south of – and territorially part of – New Zealand. The archipelago lies to the southeast of Stewart Island / Rakiura, and to the ...
. The subspecies ''E. c. chrysocome'', which may be referred to the true southern subspecies, reproduces at offshore islands in southern
Chile Chile, officially the Republic of Chile, is a country in western South America. It is the southernmost country in the world and the closest to Antarctica, stretching along a narrow strip of land between the Andes, Andes Mountains and the Paci ...
,
Argentina Argentina, officially the Argentine Republic, is a country in the southern half of South America. It covers an area of , making it the List of South American countries by area, second-largest country in South America after Brazil, the fourt ...
and at the
Falkland Islands The Falkland Islands (; ), commonly referred to as The Falklands, is an archipelago in the South Atlantic Ocean on the Patagonian Shelf. The principal islands are about east of South America's southern Patagonian coast and from Cape Dub ...
. The Falkland Islands are known to have one of the largest populations of southern rockhopper penguins. The subspecies in the group are: *
Southern rockhopper penguin The western rockhopper penguin (''Eudyptes chrysocome''), traditionally known as the southern rockhopper penguin, is a species of rockhopper penguin that is sometimes considered distinct from the northern rockhopper penguin. It occurs in subanta ...
, ''Eudyptes (chrysocome) chrysocome'' *
Eastern rockhopper penguin The eastern rockhopper penguin (''Eudyptes filholi''), also known as the tawaki piki toka, is a crested penguin with yellow crest feathers. It was formerly treated as conspecific with the western rockhopper penguin with the name "southern rockh ...
, ''Eudyptes (chrysocome) filholi'' *
Northern rockhopper penguin The northern rockhopper penguin, Moseley's rockhopper penguin, or Moseley's penguin (''Eudyptes moseleyi'') is a penguin species native to the southern Indian Ocean, Indian and Atlantic Oceans. It is described as distinct from the southern rockho ...
, ''Eudyptes (chrysocome) moseleyi''


Habitat and distribution

Northern rockhopper penguins breed in cool temperate climates including on the islands of Gough and
Tristan da Cunha Tristan da Cunha (), colloquially Tristan, is a remote group of volcano, volcanic islands in the South Atlantic Ocean. It is one of three constituent parts of the British Overseas Territories, British Overseas Territory of Saint Helena, Ascensi ...
in the Atlantic Ocean and Saint Paul and Amsterdam Islands in the Indian Ocean. The southern rockhopper breeds on the
Falkland Islands The Falkland Islands (; ), commonly referred to as The Falklands, is an archipelago in the South Atlantic Ocean on the Patagonian Shelf. The principal islands are about east of South America's southern Patagonian coast and from Cape Dub ...
, Argentina and Chile, with breeding colonies around Cape Horn in South America, and Gough, Prince Edward, Marion, Crozet, Kerguelen, Heard, Macquarie, Campbell, Auckland and Antipodes Islands in the southern Indian Ocean. Eastern rockhopper penguins are mostly found breeding on Campbell Island in New Zealand, but their numbers have declined immensely. Rockhopper penguins usually make their habitat in rocky shorelines. They make nests and burrows in tall grasses called tussocks.


Diet

The rockhopper penguin's diet consists of krill and small crustaceans, which may include shrimp, crabs, lobsters or crayfish. They also eat squid and myctophid fish. Rockhopper penguins consume more krill than they do fish; their diet changes during migration and as the seasons change. Rockhopper penguins can be at sea for several days while hunting. They can dive up to for many minutes at a time while searching for prey.


Reproduction

Rockhoppers are the most widespread crested penguins.Hull, C., Hindell, M., Mar, K., Scofield, P., Wilson, J., & Lea, M. (n.d.). The breeding biology and factors affecting reproductive success in rockhopper penguins Eudyptes chrysocome at Macquarie Island. Polar Biology, 711-720. Their range goes from the Antarctic front to the South Atlantic and Indian Oceans. Because of how widespread they are, breeding varies among the penguins in different areas. Northern Penguins begin the mating process two months earlier than Southern ones.Jouventin, P., Cuthbert, R., & Ottvall, R. (n.d.). Genetic isolation and divergence in sexual traits: Evidence for the northern rockhopper penguin Eudyptes moseleyi being a sibling species. Molecular Ecology, 3413-3423. Two eggs are usually laid a few days apart during early November in hope that at least one will survive, and the second egg is usually larger than the first. They will use the same nest as they did in previous years as well. Eggs hatch about a month later and the mother will have food for the chick.Bingham, M. (n.d.). Rockhopper Penguin


Taxonomy

There are currently 19 species and six genera of living penguins. The rockhopper penguin has 3 subspecies: ''Eudyptes filholi'', the Eastern Rockhopper; ''Eudyptes mosleyi'', the Northern Rockhopper; and the ''Eudyptes chrysocome'', the Southern Rockhopper. They are separated by their locations of breeding. There are four other species in the genus ''Eudyptes'': ''E. pachyrhynchus'', ''E. robustus'', ''E. chrysolophus'', and ''E. schlegeli''. The three subspecies of the rockhopper penguin are believed to have split up because of latitude and watermasses rather than because of geographic distances. Research suggests that during the early Pleistocene, the Southern Ocean was cold and rockhopper populations from the Atlantic and Indian Oceans remained undifferentiated and lived in the same watermass. The mid-Pleistocene climatic transition was associated with the southward migration of frontal positions and islands became surrounded by subtropical water masses, resulting in a split between Northern and Southern rockhopper penguins. Many advance and retreat cycles of Patagonian icecaps during the late Pleistocene may have created a barrier to gene flow between Southern Pacific and Southern Atlantic populations, causing a split between Southern and Eastern rockhopper populations.


Predators

Adult Rockhopper penguins have no land-based predators. They are eaten when at sea by
leopard seal The leopard seal (''Hydrurga leptonyx''), also referred to as the sea leopard, is the second largest species of seal in the Antarctic (after the southern elephant seal). It is a top order predator, feeding on a wide range of prey including cep ...
s,
fur seal Fur seals are any of nine species of pinnipeds belonging to the subfamily Arctocephalinae in the family Otariidae. They are much more closely related to sea lions than Earless seal, true seals, and share with them external ears (Pinna (anatomy ...
s,
killer whale The orca (''Orcinus orca''), or killer whale, is a toothed whale and the largest member of the oceanic dolphin family. The only extant species in the genus '' Orcinus'', it is recognizable by its black-and-white-patterned body. A cosmopolit ...
s and blue sharks. Chicks and eggs are eaten by many birds including giant petrels, skuas, sheathbills and various types of
gull Gulls, or colloquially seagulls, are seabirds of the subfamily Larinae. They are most closely related to terns and skimmers, distantly related to auks, and even more distantly related to waders. Until the 21st century, most gulls were placed ...
s.


Conservation

Although the rockhopper penguin is one of the world's most numerous penguin populations, it is estimated that the population declined by some 30 percent over the final 30 years of the 20th century. The cause of this decline is mainly unknown, but scientists have speculated that humans are involved, mostly by commercial overfishing, oil exploitation, and pollution. Recent research has shown that rockhopper penguin population declines with changes in ocean surface temperature, indicating a direct impact from climate change.Dehnhard, N., Poisbleau, M., Demongin, L., Ludynia, K., Lecoq, M., Masello, J., & Quillfeldt, P. (2013). Survival of rockhopper penguins in times of global climate change. Aquatic Conservation: Marine and Freshwater Ecosystems, 23(5), 777-789. Retrieved October 23, 2014, from Wiley Online Library. These activities have depleted much of the rockhopper penguin's food supply and raised the
sea surface temperature Sea surface temperature (or ocean surface temperature) is the ocean temperature, temperature of ocean water close to the surface. The exact meaning of ''surface'' varies in the literature and in practice. It is usually between and below the sea ...
s, and because this species is geographically isolated to the tip of South America and the Falkland Islands and is very sensitive to food web changes, there is little foreseeable opportunity for the penguins to recover.Hilton, G., Thompson, D., Sagar, P., Cuthbert, R., Cheryl, Y., & Bury, S. (2006). A stable isotopic investigation into the causes of decline in a sub-Antarctic predator, the rockhopper penguin Eudyptes chrysocome. Global Change Biology, 2006(12), 611-625. Retrieved October 23, 2014, from Web of Science.


Oil spills

An oil spill followed the grounding of the MS ''Oliva'' off Nightingale Island in 2011, which resulted in thousands of Rockhopper penguins becoming oiled. Oil spills have impacted many penguin species since the conversion of shipping from sail and coal propulsion to liquid-based fuels in the early 20th century.


Gallery

File:Gorfou sauteur - Rockhopper Penguin.jpg,
Southern rockhopper penguin The western rockhopper penguin (''Eudyptes chrysocome''), traditionally known as the southern rockhopper penguin, is a species of rockhopper penguin that is sometimes considered distinct from the northern rockhopper penguin. It occurs in subanta ...
, ''Eudyptes (chrysocome) chrysocome'' File:Eudyptes moseleyi -Zoologischer Garten Berlin, Germany-8a.jpg,
Northern rockhopper penguin The northern rockhopper penguin, Moseley's rockhopper penguin, or Moseley's penguin (''Eudyptes moseleyi'') is a penguin species native to the southern Indian Ocean, Indian and Atlantic Oceans. It is described as distinct from the southern rockho ...
, ''Eudyptes (chrysocome) moseleyi'' File:MacquarieIslandRockhoppers.JPG,
Eastern rockhopper penguin The eastern rockhopper penguin (''Eudyptes filholi''), also known as the tawaki piki toka, is a crested penguin with yellow crest feathers. It was formerly treated as conspecific with the western rockhopper penguin with the name "southern rockh ...
, ''Eudyptes (chrysocome) filholi''


In films

Rockhoppers are seen in the films, '' The Pebble and the Penguin'', ''
Happy Feet ''Happy Feet'' is a 2006 animated jukebox musical comedy film directed and produced by George Miller and written by Miller, John Collee, Judy Morris and Warren Coleman. It stars the voices of Elijah Wood, Robin Williams, Brittany Murphy, Hug ...
'' and '' Happy Feet 2'', '' Surf's Up'' and '' Surf's Up 2: WaveMania'', and '' The Wild''.


See also

* Rockhopper *
Macaroni penguin The macaroni penguin (''Eudyptes chrysolophus'') is a species of penguin found from the Subantarctic to the Antarctic Peninsula. One of six species of crested penguin, it is very closely related to the royal penguin, and some authorities consid ...
*
Northern rockhopper penguin The northern rockhopper penguin, Moseley's rockhopper penguin, or Moseley's penguin (''Eudyptes moseleyi'') is a penguin species native to the southern Indian Ocean, Indian and Atlantic Oceans. It is described as distinct from the southern rockho ...


References


External links

{{Commons, Eudyptes chrysocome
Rockhopper penguin on PenguinWorld
Penguins Flightless birds