The rock partridge or common rock partridge (''Alectoris graeca'') is a
gamebird
Galliformes is an order of heavy-bodied ground-feeding birds that includes turkeys, chickens, quail, and other landfowl. Gallinaceous birds, as they are called, are important in their ecosystems as seed dispersers and predators, and are ofte ...
in the pheasant family,
Phasianidae
Phasianidae is a family (biology), family of heavy, ground-living birds, which includes pheasants, grouse, partridges, junglefowl, chickens, Turkey bird, turkeys, Old World quail, and peafowl. The family includes many of the most popular Game (hu ...
, of the order
Galliformes
Galliformes is an order (biology), order of heavy-bodied ground-feeding birds that includes turkey (bird), turkeys, chickens, Old World quail, quail, and other landfowl. Gallinaceous birds, as they are called, are important in their ecosystems ...
(gallinaceous
bird
Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
s). It is native to southern
Europe
Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...
, and is closely related and very similar to its eastern equivalent, the
chukar partridge
The chukar partridge (''Alectoris chukar''), or simply chukar, is a Palearctic upland Upland game, gamebird in the pheasant family Phasianidae. It has been considered to form a superspecies complex along with the rock partridge, Philby's partrid ...
, ''A. chukar''.
Habitat
It is a resident breeder in dry, open and often hilly country.
Breeding
It nests in a scantily lined ground scrape laying 5–21
egg
An egg is an organic vessel grown by an animal to carry a possibly fertilized egg cell (a zygote) and to incubate from it an embryo within the egg until the embryo has become an animal fetus that can survive on its own, at which point the ...
s.
Diet
The rock partridge takes a wide variety of seeds and some
insect
Insects (from Latin ') are Hexapoda, hexapod invertebrates of the class (biology), class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (Insect morphology#Head, head, ...
food.
Description
The rock partridge is a rotund bird, with a light brown back, grey breast and buff belly. The face is white with a black gorget. It has rufous-streaked flanks and red legs. When disturbed, it prefers to run rather than fly, but if necessary it flies a short distance on rounded wings. It is very similar to the chukar partridge, but is greyer on the back and has a white, not yellowish foreneck. The sharply defined gorget distinguishes this species from red-legged partridge. The song is a noisy ''ga-ga-ga-ga-chakera- chakera- chakera''.
Some confusion exists in the naming of this species because the name of the species called
stone partridge in English translates literally from other languages into English as "rock partridge". For instance, in Dutch the species is ''rotspatrijs'', in German – ''Felsenhenne'', in French – ''poulette de roche'' – all literal translations of "rock partridge". The confusion is further compounded as in some languages, ''
Alectoris
''Alectoris'' is a genus of partridges in the family Phasianidae, closely related to Old World quail (''Coturnix'' and relatives), snowcocks (''Tetraogallus''), partridge-francolins (''Pternistis''), bush quail (''Perdicula''), and sand and see-s ...
'' species are known by names that literally translate as stone partridge. In Dutch, ''steenpatrijs'', in German ''Steinhuhn'';
red-legged partridge (''A. rufa'') being ''rode steenpatrijs'' in Dutch. Further complication arises as, particularly within the US, the name "rock partridge" has been used for a variety of ''Alectoris'' species and hybrids. The international bird trade, for sport, aviaries' and meat, has led to misapplication of various of these common and scientific names.
Conservation
This species is declining in parts of its range due to
habitat loss
Habitat destruction (also termed habitat loss or habitat reduction) occurs when a natural habitat is no longer able to support its native species. The organisms once living there have either moved elsewhere, or are dead, leading to a decrease ...
and over-hunting. While populations are usually stable, the status of the
Sicilian population may be more precarious, and certainly deserves attention (Randi 2006).
Taxonomy
This species is closely related to the chukar,
Przevalski's, and
Philby's partridges, forming a
superspecies
In biology, a species complex is a group of closely related organisms that are so similar in appearance and other features that the boundaries between them are often unclear. The taxa in the complex may be able to hybridize readily with each oth ...
. The Western Mediterranean
red-legged and
Barbary partridges with their spotted neck collar are slightly more distant relatives. Nonetheless, although this species' range does not naturally overlap with that of its relatives, they co-occur where they have been introduced as gamebirds, for example in
North America
North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...
, the
United Kingdom
The United Kingdom of Great Britain and Northern Ireland, commonly known as the United Kingdom (UK) or Britain, is a country in Northwestern Europe, off the coast of European mainland, the continental mainland. It comprises England, Scotlan ...
,
New Zealand
New Zealand () is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and List of islands of New Zealand, over 600 smaller islands. It is the List of isla ...
, and
Russia
Russia, or the Russian Federation, is a country spanning Eastern Europe and North Asia. It is the list of countries and dependencies by area, largest country in the world, and extends across Time in Russia, eleven time zones, sharing Borders ...
, and in southeastern
France
France, officially the French Republic, is a country located primarily in Western Europe. Overseas France, Its overseas regions and territories include French Guiana in South America, Saint Pierre and Miquelon in the Atlantic Ocean#North Atlan ...
where red-legged partridges have been released. In these areas,
hybrids between this species, the chukar, and the red-legged partridge are usually found.(McGowan 1994, ''see also'' Randi 2006)
The rock partridge has four recognized
subspecies
In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
:
* ''A. g. graeca''
( Meisner, 1804) -
Serbia
, image_flag = Flag of Serbia.svg
, national_motto =
, image_coat = Coat of arms of Serbia.svg
, national_anthem = ()
, image_map =
, map_caption = Location of Serbia (gree ...
and
Albania
Albania ( ; or ), officially the Republic of Albania (), is a country in Southeast Europe. It is located in the Balkans, on the Adriatic Sea, Adriatic and Ionian Seas within the Mediterranean Sea, and shares land borders with Montenegro to ...
to
Greece
Greece, officially the Hellenic Republic, is a country in Southeast Europe. Located on the southern tip of the Balkan peninsula, it shares land borders with Albania to the northwest, North Macedonia and Bulgaria to the north, and Turkey to th ...
and
Bulgaria
Bulgaria, officially the Republic of Bulgaria, is a country in Southeast Europe. It is situated on the eastern portion of the Balkans directly south of the Danube river and west of the Black Sea. Bulgaria is bordered by Greece and Turkey t ...
* ''A. g. orlandoi''
(Priolo, 1984) -
Italy
Italy, officially the Italian Republic, is a country in Southern Europe, Southern and Western Europe, Western Europe. It consists of Italian Peninsula, a peninsula that extends into the Mediterranean Sea, with the Alps on its northern land b ...
* ''A. g. saxatilis''
( Bechstein, 1805) –
Alps
The Alps () are some of the highest and most extensive mountain ranges in Europe, stretching approximately across eight Alpine countries (from west to east): Monaco, France, Switzerland, Italy, Liechtenstein, Germany, Austria and Slovenia.
...
from
France
France, officially the French Republic, is a country located primarily in Western Europe. Overseas France, Its overseas regions and territories include French Guiana in South America, Saint Pierre and Miquelon in the Atlantic Ocean#North Atlan ...
to western
Croatia
Croatia, officially the Republic of Croatia, is a country in Central Europe, Central and Southeast Europe, on the coast of the Adriatic Sea. It borders Slovenia to the northwest, Hungary to the northeast, Serbia to the east, Bosnia and Herze ...
* ''A. g. whitakeri'' (
Schiebel, 1934) –
Sicily
Sicily (Italian language, Italian and ), officially the Sicilian Region (), is an island in the central Mediterranean Sea, south of the Italian Peninsula in continental Europe and is one of the 20 regions of Italy, regions of Italy. With 4. ...
* ''A. g. martelensis'' –
paleosubspecies
A chronospecies is a species derived from a sequential development pattern that involves continual and uniform changes from an extinct ancestral form on an evolutionary scale. The sequence of alterations eventually produces a population that is p ...
, only known from
fossil
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
s
The proposed subspecies from the Apennines, ''A. g. orlandoi''
Priolo, 1984, is of doubtful validity. It is usually included in ''saxatilis'', but apparently mostly derives from
Albania
Albania ( ; or ), officially the Republic of Albania (), is a country in Southeast Europe. It is located in the Balkans, on the Adriatic Sea, Adriatic and Ionian Seas within the Mediterranean Sea, and shares land borders with Montenegro to ...
n ''A. g. graeca''. These probably crossed the
Adriatic
The Adriatic Sea () is a body of water separating the Italian Peninsula from the Balkans, Balkan Peninsula. The Adriatic is the northernmost arm of the Mediterranean Sea, extending from the Strait of Otranto (where it connects to the Ionian Se ...
via a land-bridge during the
last ice age, to become isolated only with the sea levels rising at the beginning of the
Holocene
The Holocene () is the current geologic time scale, geological epoch, beginning approximately 11,700 years ago. It follows the Last Glacial Period, which concluded with the Holocene glacial retreat. The Holocene and the preceding Pleistocene to ...
c.12.000–10.000 years ago, with Alpine birds much less contributing to the Apennines population (Randi 2006).
Apennine birds are not consistently recognizable by external
morphology
Morphology, from the Greek and meaning "study of shape", may refer to:
Disciplines
*Morphology (archaeology), study of the shapes or forms of artifacts
*Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies, ...
, and are only weakly differentiated with regards to
mtDNA
Mitochondrial DNA (mtDNA and mDNA) is the DNA located in the mitochondria organelles in a eukaryotic cell that converts chemical energy from food into adenosine triphosphate (ATP). Mitochondrial DNA is a small portion of the DNA contained in ...
D-loop and
hypervariable control region sequence
In mathematics, a sequence is an enumerated collection of objects in which repetitions are allowed and order matters. Like a set, it contains members (also called ''elements'', or ''terms''). The number of elements (possibly infinite) is cal ...
s and
microsatellite
A microsatellite is a tract of repetitive DNA in which certain Sequence motif, DNA motifs (ranging in length from one to six or more base pairs) are repeated, typically 5–50 times. Microsatellites occur at thousands of locations within an organ ...
genotyping. As they nonetheless constitute a discrete subpopulation
evolving towards subspecies status, their population numbers could arguably deserve monitoring (Randi 2006).
References

* McGowan, Philip J. K. (1994): 11. Rock Partridge. ''In:'' del Hoyo, Josep; Elliott, Andrew & Sargatal, Jordi (editors): ''
Handbook of Birds of the World, Volume 2: New World Vultures to Guineafowl'': 485, plate 43. Lynx Edicions, Barcelona.
*Randi, Ettore (2006): Evolutionary and conservation genetics of the rock partridge, ''Alectoris graeca''. ''Acta Zoologica Sinica'' 52(Supplement): 370–374
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rock partridge
Birds of Southern Europe
rock partridge