Robert Harding Whittaker (December 27, 1920 – October 20, 1980) was an American

plant Plants are the eukaryotes that form the Kingdom (biology), kingdom Plantae; they are predominantly Photosynthesis, photosynthetic. This means that they obtain their energy from sunlight, using chloroplasts derived from endosymbiosis with c ...

ecologist Ecology () is the natural science of the relationships among living organisms and their environment. Ecology considers organisms at the individual, population, community, ecosystem, and biosphere levels. Ecology overlaps with the closely re ...

, active from the 1950s to the 1970s. He was the first to propose the five kingdom

taxonomic classification In biology, taxonomy () is the scientific study of naming, defining ( circumscribing) and classifying groups of biological organisms based on shared characteristics. Organisms are grouped into taxa (singular: taxon), and these groups are given ...

of the world's biota into the

Animalia Animals are multicellular, eukaryotic organisms in the Biology, biological Kingdom (biology), kingdom Animalia (). With few exceptions, animals heterotroph, consume organic material, Cellular respiration#Aerobic respiration, breathe oxygen, ...

, Plantae,

Fungi A fungus (: fungi , , , or ; or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and mold (fungus), molds, as well as the more familiar mushrooms. These organisms are classified as one ...

Protista A protist ( ) or protoctist is any Eukaryote, eukaryotic organism that is not an animal, Embryophyte, land plant, or fungus. Protists do not form a Clade, natural group, or clade, but are a Paraphyly, paraphyletic grouping of all descendants o ...

, and Monera in 1969.Hagen, Joel B. (2012)
Five kingdoms, more or less: Robert Whittaker and the broad classification of organisms
. ''BioScience'', 62 (1): 67-74. He also proposed the Whittaker

Biome A biome () is a distinct geographical region with specific climate, vegetation, and animal life. It consists of a biological community that has formed in response to its physical environment and regional climate. In 1935, Tansley added the ...

Classification, which categorized biome types upon two abiotic factors: temperature and precipitation. He proposed the concepts of

Alpha diversity In ecology, alpha diversity (α-diversity) is the mean species diversity in a site at a local scale. The term was introduced by R. H. WhittakerWhittaker, R. H. (1960) Vegetation of the Siskiyou Mountains, Oregon and California. Ecological Monogra ...

Beta diversity In ecology, beta diversity (β-diversity or true beta diversity) is the ratio between regional and local species diversity. The term was introduced by Robert Whittaker (ecologist), R. H. Whittaker together with the terms alpha diversity (α-diversi ...

, and

Gamma diversity In ecology, gamma diversity (γ-diversity) is the total species diversity in a landscape. The term was introduced by R. H. WhittakerWhittaker, R. H. (1960) Vegetation of the Siskiyou Mountains, Oregon and California. Ecological Monographs, 30, 279 ...

.Whittaker, R. H. (1960) Vegetation of the Siskiyou Mountains, Oregon and California. Ecological Monographs, 30, 279–338. Whittaker, R. H. (1972). Evolution and Measurement of Species Diversity. Taxon, 21, 213-251. Whittaker was elected to the

National Academy of Sciences The National Academy of Sciences (NAS) is a United States nonprofit, NGO, non-governmental organization. NAS is part of the National Academies of Sciences, Engineering, and Medicine, along with the National Academy of Engineering (NAE) and the ...

in 1974, received the Ecological Society of America's

Eminent Ecologist Award The Eminent Ecologist Award is prize awarded annually to a senior ecologist in recognition of an outstanding contribution to the science of ecology. The prize is awarded by the Ecological Society of America. According to the statutes, the recipient ...

in 1981, and was otherwise widely recognized and honored. He collaborated with many other ecologists including George Woodwell (Dartmouth), W. A. Niering, F. H. Bormann (Yale), and G. E. Likens (Cornell), and was particularly active in cultivating international collaborations

Early life

Born in

Wichita, Kansas Wichita ( ) is the List of cities in Kansas, most populous city in the U.S. state of Kansas and the county seat of Sedgwick County, Kansas, Sedgwick County. As of the 2020 United States census, 2020 census, the population of the city was 397, ...

, he obtained a B.A. at Washburn Municipal College (now

Washburn University Washburn University (WU), formally Washburn University of Topeka, is a public university in Topeka, Kansas, United States. It offers undergraduate and graduate programs as well as professional programs in law and business. The university enroll ...

) in

Topeka, Kansas Topeka ( ) is the capital city of the U.S. state of Kansas and the county seat of Shawnee County. It is along the Kansas River in the central part of Shawnee County, in northeastern Kansas, in the Central United States. As of the 2020 cen ...

, and, following

military service Military service is service by an individual or group in an army or other militia, air forces, and naval forces, whether as a chosen job (volunteer military, volunteer) or as a result of an involuntary draft (conscription). Few nations, such ...

, his Ph.D. in Biology at the

University of Illinois The University of Illinois Urbana-Champaign (UIUC, U of I, Illinois, or University of Illinois) is a public university, public land-grant university, land-grant research university in the Champaign–Urbana metropolitan area, Illinois, United ...

in 1948.

Career

Whittaker held teaching and research positions at Washington State College in

Pullman, Washington Pullman is the most populous city in Whitman County, located in southeastern Washington within the Palouse region of the Pacific Northwest. The population was 32,901 at the 2020 census, and estimated to be 32,508 in 2022. Originally founded as ...

from 1948 to 1951, and then moved Hanford Laboratories Aquatic Biology Unit near

Richland, Washington Richland () is a city in Benton County, Washington, United States. It is located in southeastern Washington at the confluence of the Yakima River, Yakima and the Columbia River, Columbia Rivers. As of the 2020 census, the city's population was ...

. In 1954, he was hired as an instructor in the Department of Biology of Brooklyn College, the City University of New York. In the 1960s, he worked at the

University of California, Irvine The University of California, Irvine (UCI or UC Irvine) is a Public university, public Land-grant university, land-grant research university in Irvine, California, United States. One of the ten campuses of the University of California system, U ...

and

Cornell University Cornell University is a Private university, private Ivy League research university based in Ithaca, New York, United States. The university was co-founded by American philanthropist Ezra Cornell and historian and educator Andrew Dickson W ...

Family

Whittaker married biochemist Clara Buehl (then a coworker at Hanford Laboratories) in 1952. They had three children. Clara was diagnosed with cancer in 1972, and she later died on December 31, 1976. Whittaker married graduate student Linda Olsvig in 1979. He too was diagnosed with lung cancer and died on October 20, 1980.

Works

*Robert H. Whittaker, ''Communities and Ecosystems'', Macmillan, 1970. Second edition 1975. *Robert H. Whittaker (Ed), ''Classification of Plant Communities, 1978 (Handbook of Vegetation Science)'', Kluwer Academic Publishers,

References

{{DEFAULTSORT:Whittaker, Robert Cornell University faculty American ecologists 1920 births 1980 deaths Ecological succession Scientists from Wichita, Kansas Plant ecologists Members of the United States National Academy of Sciences Deaths from lung cancer in New York (state) Brooklyn College faculty Presidents of the American Society of Naturalists