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The roadrunners (genus ''Geococcyx''), also known as chaparral birds or chaparral cocks, are two species of fast-running ground
cuckoo Cuckoos are birds in the Cuculidae ( ) family, the sole taxon in the order Cuculiformes ( ). The cuckoo family includes the common or European cuckoo, roadrunners, koels, malkohas, couas, coucals, and anis. The coucals and anis are somet ...
s with long tails and crests. They are found in the southwestern and south-central United States,
Mexico Mexico, officially the United Mexican States, is a country in North America. It is the northernmost country in Latin America, and borders the United States to the north, and Guatemala and Belize to the southeast; while having maritime boundar ...
and
Central America Central America is a subregion of North America. Its political boundaries are defined as bordering Mexico to the north, Colombia to the southeast, the Caribbean to the east, and the Pacific Ocean to the southwest. Central America is usually ...
, usually in the desert. Although capable of flight, roadrunners generally run away from predators. On the ground, some have been measured at .


Species

The subfamily Neomorphinae, the New World ground cuckoos, includes 11 species of birds, while the genus ''Geococcyx'' has just two:


Morphology

The roadrunner generally ranges in size from from tail to
beak The beak, bill, or rostrum is an external anatomical structure found mostly in birds, but also in turtles, non-avian dinosaurs and a few mammals. A beak is used for pecking, grasping, and holding (in probing for food, eating, manipulating and ...
. The average weight is about . The roadrunner is a slender, black-brown and white-streaked ground bird with a distinctive head crest. It has long legs, strong feet, and an oversized dark bill. The tail is broad with white tips on the three outer tail feathers. The bird has a bare patch of skin behind each eye; this patch is shaded blue anterior to red posterior. The lesser roadrunner is slightly smaller, not as streaky, and has a smaller bill. Both the lesser roadrunner and the greater roadrunner leave behind very distinct "X" track marks appearing as if they are travelling in both directions. Roadrunners and other members of the cuckoo family have zygodactyl feet. The roadrunner can run at speeds of up to and generally prefer sprinting to flying, though it will fly to escape
predator Predation is a biological interaction in which one organism, the predator, kills and eats another organism, its prey. It is one of a family of common List of feeding behaviours, feeding behaviours that includes parasitism and micropredation ...
s. During flight, the short, rounded wings reveal a white crescent in the primary feathers.


Vocalization

The roadrunner has a slow and descending dove-like "coo". It also makes a rapid, vocalized clattering sound with its beak.


Geographic range

Roadrunners inhabit the
Southwestern United States The Southwestern United States, also known as the American Southwest or simply the Southwest, is a geographic and cultural list of regions of the United States, region of the United States that includes Arizona and New Mexico, along with adjacen ...
, to parts of Missouri, Arkansas, and Louisiana, as well as
Mexico Mexico, officially the United Mexican States, is a country in North America. It is the northernmost country in Latin America, and borders the United States to the north, and Guatemala and Belize to the southeast; while having maritime boundar ...
and
Central America Central America is a subregion of North America. Its political boundaries are defined as bordering Mexico to the north, Colombia to the southeast, the Caribbean to the east, and the Pacific Ocean to the southwest. Central America is usually ...
. They live in
arid Aridity is the condition of geographical regions which make up approximately 43% of total global available land area, characterized by low annual precipitation, increased temperatures, and limited water availability.Perez-Aguilar, L. Y., Plata ...
lowland or mountainous
shrubland Shrubland, scrubland, scrub, brush, or bush is a plant community characterized by vegetation dominance (ecology), dominated by shrubs, often also including grasses, herbaceous plant, herbs, and geophytes. Shrubland may either occur naturally o ...
or
woodland A woodland () is, in the broad sense, land covered with woody plants (trees and shrubs), or in a narrow sense, synonymous with wood (or in the U.S., the '' plurale tantum'' woods), a low-density forest forming open habitats with plenty of sunli ...
. They are non-migratory, staying in their breeding area year-round. The greater roadrunner is not currently considered threatened in the US, but is habitat-limited.


Food and foraging habits

The roadrunner is an opportunistic
omnivore An omnivore () is an animal that regularly consumes significant quantities of both plant and animal matter. Obtaining energy and nutrients from plant and animal matter, omnivores digest carbohydrates, protein, fat, and fiber, and metabolize t ...
. Its diet normally consists of
insect Insects (from Latin ') are Hexapoda, hexapod invertebrates of the class (biology), class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (Insect morphology#Head, head, ...
s (such as
grasshopper Grasshoppers are a group of insects belonging to the suborder Caelifera. They are amongst what are possibly the most ancient living groups of chewing herbivorous insects, dating back to the early Triassic around 250 million years ago. Grassh ...
s,
crickets Crickets are orthopteran insects which are related to bush crickets and more distantly, to grasshoppers. In older literature, such as Imms,Imms AD, rev. Richards OW & Davies RG (1970) ''A General Textbook of Entomology'' 9th Ed. Methuen 886 ...
,
caterpillar Caterpillars ( ) are the larval stage of members of the order Lepidoptera (the insect order comprising butterflies and moths). As with most common names, the application of the word is arbitrary, since the larvae of sawflies (suborder ...
s, and
beetle Beetles are insects that form the Taxonomic rank, order Coleoptera (), in the superorder Holometabola. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 40 ...
s), small
reptile Reptiles, as commonly defined, are a group of tetrapods with an ectothermic metabolism and Amniotic egg, amniotic development. Living traditional reptiles comprise four Order (biology), orders: Testudines, Crocodilia, Squamata, and Rhynchocepha ...
s (such as
lizard Lizard is the common name used for all Squamata, squamate reptiles other than snakes (and to a lesser extent amphisbaenians), encompassing over 7,000 species, ranging across all continents except Antarctica, as well as most Island#Oceanic isla ...
s and
snake Snakes are elongated limbless reptiles of the suborder Serpentes (). Cladistically squamates, snakes are ectothermic, amniote vertebrates covered in overlapping scales much like other members of the group. Many species of snakes have s ...
s, including
rattlesnake Rattlesnakes are venomous snakes that form the genus, genera ''Crotalus'' and ''Sistrurus'' of the subfamily Crotalinae (the pit vipers). All rattlesnakes are vipers. Rattlesnakes are predators that live in a wide array of habitats, hunting sm ...
s),
rodent Rodents (from Latin , 'to gnaw') are mammals of the Order (biology), order Rodentia ( ), which are characterized by a single pair of continuously growing incisors in each of the upper and Mandible, lower jaws. About 40% of all mammal specie ...
s and other small
mammal A mammal () is a vertebrate animal of the Class (biology), class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three ...
s,
spider Spiders (order (biology), order Araneae) are air-breathing arthropods that have eight limbs, chelicerae with fangs generally able to inject venom, and spinnerets that extrude spider silk, silk. They are the largest order of arachnids and ran ...
s (including
tarantula Tarantulas comprise a group of large and often hairy spiders of the family Theraphosidae. , 1,100 species have been identified, with 166 genera. The term "tarantula" is usually used to describe members of the family Theraphosidae, although ...
s),
scorpion Scorpions are predatory arachnids of the Order (biology), order Scorpiones. They have eight legs and are easily recognized by a pair of Chela (organ), grasping pincers and a narrow, segmented tail, often carried in a characteristic forward cur ...
s,
centipede Centipedes (from Neo-Latin , "hundred", and Latin , "foot") are predatory arthropods belonging to the class Chilopoda (Ancient Greek , ''kheilos'', "lip", and Neo-Latin suffix , "foot", describing the forcipules) of the subphylum Myriapoda, ...
s,
snail A snail is a shelled gastropod. The name is most often applied to land snails, terrestrial molluscs, terrestrial pulmonate gastropod molluscs. However, the common name ''snail'' is also used for most of the members of the molluscan class Gas ...
s, small birds (and nestlings), eggs, and fruits and seeds like those from
prickly pear cactus ''Opuntia'', commonly called the prickly pear cactus, is a genus of flowering plants in the cactus family Cactaceae, many known for their flavorful fruit and showy flowers. Cacti are native to the Americas, and are well adapted to arid climat ...
es and
sumac Sumac or sumach ( , )—not to be confused with poison sumac—is any of the roughly 35 species of flowering plants in the genus ''Rhus'' (and related genera) of the cashew and mango tree family, Anacardiaceae. However, it is '' Rhus coriaria ...
s. The lesser roadrunner eats mainly insects. The roadrunner forages on the ground and, when hunting, usually runs after prey from under cover. It may leap to catch insects, and commonly batters certain prey against the ground. The roadrunner is one of the few animals that preys upon rattlesnakes; it is also the only real predator of tarantula hawk wasps.


Behavior and breeding

The roadrunner usually lives alone or in pairs. Breeding pairs are monogamous and mate for life, and pairs may hold a territory all year. During the
courtship display A courtship display is a set of display behaviors in which an animal, usually a male, attempts to attract a mate; the mate exercises choice, so sexual selection acts on the display. These behaviors often include ritualized movement ("dances"), ...
, the male bows, alternately lifting and dropping his wings and spreading his tail. He parades in front of the female with his head high and his tail and wings drooped, and may bring an offering of food. The reproductive season is spring to mid-summer (depending on geographic location and species). The roadrunner's
nest A nest is a structure built for certain animals to hold Egg (biology), eggs or young. Although nests are most closely associated with birds, members of all classes of vertebrates and some invertebrates construct nests. They may be composed of ...
is often composed of sticks, and may sometimes contain leaves, feathers, snakeskins, or dung. It is commonly placed above ground level"Usually 1-3 meters above ground; infrequently higher than 3 meters (Hughes 1996)." in a low tree, bush, or
cactus A cactus (: cacti, cactuses, or less commonly, cactus) is a member of the plant family Cactaceae (), a family of the order Caryophyllales comprising about 127 genera with some 1,750 known species. The word ''cactus'' derives, through Latin, ...
. Roadrunner eggs are generally white. The greater roadrunner generally lays 2–6 eggs per
clutch A clutch is a mechanical device that allows an output shaft to be disconnected from a rotating input shaft. The clutch's input shaft is typically attached to a motor, while the clutch's output shaft is connected to the mechanism that does th ...
, but the lesser roadrunner's clutches are typically smaller. Hatching is asynchronous. Both sexes incubate the nest (with males incubating the nest at night) and feed the hatchlings. For the first one to two weeks after the young hatch, one parent remains at the nest. The young leave the nest at two to three weeks old, foraging with parents for a few days after.


Thermoregulation

During the cold desert night, the roadrunner lowers its body temperature slightly, going into a slight
torpor Torpor is a state of decreased physiological activity in an animal, usually marked by a reduced body temperature and metabolic rate. Torpor enables animals to survive periods of reduced food availability. The term "torpor" can refer to the ti ...
to conserve energy. To warm itself during the day, the roadrunner exposes dark patches of skin on its back to the sun.


Indigenous lore

The
Hopi The Hopi are Native Americans who primarily live in northeastern Arizona. The majority are enrolled in the Hopi Tribe of Arizona and live on the Hopi Reservation in northeastern Arizona; however, some Hopi people are enrolled in the Colorado ...
and other
Pueblo Pueblo refers to the settlements of the Pueblo peoples, Native American tribes in the Southwestern United States, currently in New Mexico, Arizona, and Texas. The permanent communities, including some of the oldest continually occupied settlement ...
tribes believed roadrunners were ''
medicine Medicine is the science and Praxis (process), practice of caring for patients, managing the Medical diagnosis, diagnosis, prognosis, Preventive medicine, prevention, therapy, treatment, Palliative care, palliation of their injury or disease, ...
'' birds, capable of warding off evil spirits. The X-shaped footprints of roadrunners were seen as sacred symbols, believed to confuse evil spirits by concealing the bird's direction of travel. Stylized roadrunner tracks have been found in the rock art of ancestral Southwestern tribes like the Mogollon cultures. Roadrunner feathers were used to decorate Pueblo cradleboards for spiritual protection. Among Mexican Indian and American Indian tribes, such as the Pima, seeing a roadrunner is considered good luck. While some Mexican tribes revered the roadrunner and never killed it, most used its meat as a folk remedy for illness or to boost stamina and strength. Central American Indigenous peoples have various beliefs about the roadrunner. The Ch’orti’, known to call it t’unk’u’x or mu’, have taboos against harming the bird. The Ch'ol
Maya Maya may refer to: Ethnic groups * Maya peoples, of southern Mexico and northern Central America ** Maya civilization, the historical civilization of the Maya peoples ** Mayan languages, the languages of the Maya peoples * Maya (East Africa), a p ...
believe roadrunners possess special powers, calling it ajkumtz’u’ due to its call, which is believed to induce tiredness in listeners. The word for roadrunner in the O'odham language is , which is the name of a transit center in
Tucson, Arizona Tucson (; ; ) is a city in Pima County, Arizona, United States, and its county seat. It is the second-most populous city in Arizona, behind Phoenix, Arizona, Phoenix, with a population of 542,630 in the 2020 United States census. The Tucson ...
. In the O'odham tradition, the roadrunner is also credited with bringing fire to the people.


In media

The roadrunner is the state bird of New Mexico. The roadrunner was made popular by the
Warner Bros. Warner Bros. Entertainment Inc. (WBEI), commonly known as Warner Bros. (WB), is an American filmed entertainment studio headquartered at the Warner Bros. Studios complex in Burbank, California and the main namesake subsidiary of Warner Bro ...
cartoon characters Wile E. Coyote and the Road Runner, created in 1949, and the subject of a long-running series of theatrical cartoon shorts. In each episode, the cunning, insidious, and constantly hungry Wile E. Coyote repeatedly attempts to catch and subsequently eat the Road Runner, but is never successful. The cartoons led to a misconception that the call of the roadrunner is "meep, meep" because the roadrunner in this cartoon series made that sound instead of the aforementioned sound of a real roadrunner. In some shorts, the Road Runner makes a noise while sticking his tongue out at Wile E. Coyote, which resembles its actual call. The cartoons rely on a misconception that a roadrunner is much faster than a coyote. In fact, a coyote's fastest sprinting speed is , which is twice that of a roadrunner's at .


Citations


General references

* * * * * * * *


Further reading

*


External links


Distribution map of the Greater RoadrunnerGreater Roadrunner soundsVideo clips of roadrunners in action
{{Authority control Geococcyx, * Articles containing video clips Birds of Mexico