''Rioceras'' is an extinct genus of
orthocerid
Orthocerida is an order of extinct Orthoceratoid cephalopods also known as the Michelinocerida that lived from the Early Ordovician () possibly to the Late Triassic (). A fossil found in the Caucasus suggests they may even have survived until ...
cephalopods of the family
Baltoceratidae
Baltoceratidae is an extinct family of orthoconic cephalopods belonging to the subclass Nautiloidea endemic to what would be Asia, Australia, Europe, North America, and South America during the Ordovician living from about 480–460 mya, exist ...
that lived in what would be North America during the Early
Ordovician
The Ordovician ( ) is a geologic period and System (geology), system, the second of six periods of the Paleozoic Era (geology), Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period million years ago (Mya) to the start ...
from 480—472
mya
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, existing for approximately 8 million years.
[R. H. Flower. 1964. The Nautiloid Order Ellesmeroceratida (Cephalopoda). State Bureau of Mines and Mineral Resources, New Mexico Institute of Mining and Technology, Memoir 12:234]
Taxonomy
''Rioceras'' was named by Flower (1964). Its type is ''Rioceras nondescriptum''. It was assigned to ''
Baltoceratidae
Baltoceratidae is an extinct family of orthoconic cephalopods belonging to the subclass Nautiloidea endemic to what would be Asia, Australia, Europe, North America, and South America during the Ordovician living from about 480–460 mya, exist ...
'' by Flower (1964) and Hook and Flower (1977),
[S. C. Hook and R. H. Flower. 1977. Late Canadian (Zones J, K) cephalopod faunas from Southwestern United States. New Mexico Bureau of Mines and Mineral Resources 32:1-56] but reassigned to the Ellesmeroceratidae by Kröger et al. (2007)[B. Kröger, M. S. Beresi, and E. Landing. 2007. Early orthoceratoid cephalopods from the Argentine Precordillera (Lower-Middle Ordovician). Journal of Paleontology 81(6):1266-1283]
Whether Rioceras is assigned to the Baltoceratidae and therefore to the Orthocerida or to the Ellesmeroceratidae depends on the definition of each. Flower[ placed Rioceras in the Baltoceratidae when he proposed the genus since it lacks diaphragms in the siphuncle which for him was a defining character of the family. Kröger][ removed Rioceras to the Ellesmeroceratidae on the basis of its inwardly thicked connecting rings which produce concave segments, a defining character of that family as emended. Flower did point out in his original diagnosis][ that Rioceras has moderately thick connecting rings "of ellesmeroceratoid aspect"
]
Morphology
The shell is usually long, and may be straight ("orthoconic
An orthocone is an unusually long straight shell of a nautiloid cephalopod.; During the 18th and 19th centuries, all shells of this type were named ''Orthoceras'', creating a wastebasket taxon, but it is now known that many groups of nautiloids d ...
") or gently curved. In life, these animals may have been similar to the modern squid
True squid are molluscs with an elongated soft body, large eyes, eight arms, and two tentacles in the superorder Decapodiformes, though many other molluscs within the broader Neocoleoidea are also called squid despite not strictly fitting t ...
, except for the long shell.
References
{{Taxonbar, from=Q7335393
Prehistoric nautiloid genera
Ordovician cephalopods of North America