Richard L. Hoffman
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Richard Lawrence Hoffman (September 25, 1927 – June 10, 2012) was an American
zoologist Zoology ( , ) is the scientific study of animals. Its studies include the structure, embryology, classification, habits, and distribution of all animals, both living and extinct, and how they interact with their ecosystems. Zoology is one ...
known as an international expert on
millipedes Millipedes (originating from the Latin , "thousand", and , "foot") are a group of arthropods that are characterised by having two pairs of jointed legs on most body segments; they are known scientifically as the class Diplopoda, the name derive ...
, and a leading authority on the
natural history Natural history is a domain of inquiry involving organisms, including animals, fungi, and plants, in their natural environment, leaning more towards observational than experimental methods of study. A person who studies natural history is cal ...
of
Virginia Virginia, officially the Commonwealth of Virginia, is a U.S. state, state in the Southeastern United States, Southeastern and Mid-Atlantic (United States), Mid-Atlantic regions of the United States between the East Coast of the United States ...
and the
Appalachian Mountains The Appalachian Mountains, often called the Appalachians, are a mountain range in eastern to northeastern North America. The term "Appalachian" refers to several different regions associated with the mountain range, and its surrounding terrain ...
. He was a biology professor at Virginia's
Radford College Radford College is an independent school, Anglican, coeducational day school, located in Bruce, Australian Capital Territory, Australia. Founded in 1984, the college is named after Bishop Lewis Bostock Radford. It has a non-selective enrolmen ...
for almost thirty years, and curator of invertebrates at the
Virginia Museum of Natural History The Virginia Museum of Natural History is the state's natural history museum located in Martinsville, Virginia founded in 1984. The museum has several different award-winning publications, is affiliated with the Smithsonian Institution, and has ...
for another twenty years. He co-founded the Virginia Natural History Society, described over 400 species of millipedes, and produced more than 480 scientific publications. He is commemorated in the scientific and/or common names of over 30 animal species, including the valley and ridge salamander (''Plethodon hoffmani'') and Hoffman's dwarf centipede (''Nannarrup hoffmani'').


Biography

Richard Hoffman was born in
Clifton Forge, Virginia Clifton Forge is a town in Alleghany County, Virginia, United States which is part of the greater Roanoke Region. The population was 3,555 at the 2020 census. The Jackson River flows through the town, which as a result was once known as Ja ...
, on September 25, 1927. His father was a railroad machinist whose parents had emigrated from Germany in the late 1800s, and his mother was from a Virginia farming family. Hoffman's scientific publications began at the age of 16, and by the age of 24 he had 29 publications. He attended the
University of Virginia The University of Virginia (UVA) is a Public university#United States, public research university in Charlottesville, Virginia, United States. It was founded in 1819 by Thomas Jefferson and contains his The Lawn, Academical Village, a World H ...
until 1950, where he was influenced by Horton H. Hobbs, Jr. a crustacean biologist and recently hired professor. Unable to complete a freshman math class, Hoffman did not complete a degree at Virginia. Rather, in 1951 he was accepted into graduate school at
Cornell University Cornell University is a Private university, private Ivy League research university based in Ithaca, New York, United States. The university was co-founded by American philanthropist Ezra Cornell and historian and educator Andrew Dickson W ...
, who accepted his publication record as equivalent to passing a math class. When not working in the field or at Cornell, Hoffman worked as a chemist at the Radford Arsenal. He completed his a M.S. in
entomology Entomology (from Ancient Greek ἔντομον (''éntomon''), meaning "insect", and -logy from λόγος (''lógos''), meaning "study") is the branch of zoology that focuses on insects. Those who study entomology are known as entomologists. In ...
in 1959, and went on to earn a PhD in Zoology from
Virginia Tech The Virginia Polytechnic Institute and State University, commonly referred to as Virginia Tech (VT), is a Public university, public Land-grant college, land-grant research university with its main campus in Blacksburg, Virginia, United States ...
in 1960, during which he revised the genus '' Cambarincola'', a group of leech-like
oligochaete Oligochaeta () is a subclass of soft-bodied animals in the phylum Annelida, which is made up of many types of aquatic and terrestrial worms, including all of the various earthworms. Specifically, oligochaetes comprise the terrestrial megadril ...
worms that are
commensal Commensalism is a long-term biological interaction (symbiosis) in which members of one species gain benefits while those of the other species neither benefit nor are harmed. This is in contrast with mutualism, in which both organisms benefit f ...
on
crayfish Crayfish are freshwater crustaceans belonging to the infraorder Astacidea, which also contains lobsters. Taxonomically, they are members of the superfamilies Astacoidea and Parastacoidea. They breathe through feather-like gills. Some spe ...
. His doctoral advisor at Virginia Tech was Dr. Perry C. Holt. After completing his PhD, Hoffman worked as professor of biology at
Radford College Radford College is an independent school, Anglican, coeducational day school, located in Bruce, Australian Capital Territory, Australia. Founded in 1984, the college is named after Bishop Lewis Bostock Radford. It has a non-selective enrolmen ...
(now known as Radford University) from 1960 to 1988. Over this 28-year period, Hoffman taught a number of courses including introductory biology and zoology,
invertebrate zoology Invertebrate zoology is the subdiscipline of zoology that consists of the study of invertebrates, animals without a backbone (a structure which is found only in fish, amphibians, reptiles, birds and mammals). Invertebrates are a vast and very dive ...
, entomology, herpetology, mammalogy, biogeography, and the history of science. From 1964 to 1970 Hoffman edited ''The Radford Review'', the college's scholarly journal. Hoffman made dramatic improvements to the collections of the Radford Natural History Museum. From 1989 to 2009, Hoffman worked as Curator of Recent Invertebrates at the
Virginia Museum of Natural History The Virginia Museum of Natural History is the state's natural history museum located in Martinsville, Virginia founded in 1984. The museum has several different award-winning publications, is affiliated with the Smithsonian Institution, and has ...
. Hoffman died on June 10, 2012, at
Carilion Roanoke Memorial Hospital Carilion Roanoke Memorial Hospital (CRMH) is a private teaching hospital in Roanoke, Virginia, United States. With 703 beds, Carilion Roanoke Memorial Hospital is one of the largest hospitals in the state. It is part of Carilion Clinic. The re ...
in
Roanoke, Virginia Roanoke ( ) is an Independent city (United States), independent city in Virginia, United States. It lies in Southwest Virginia, along the Roanoke River, in the Blue Ridge Mountains, Blue Ridge range of the greater Appalachian Mountains. Roanok ...
, following heart surgery. He was 84. His contributions to science and education were formally recognized by the
Virginia General Assembly The Virginia General Assembly is the legislative body of the Commonwealth of Virginia, the oldest continuous law-making body in the Western Hemisphere, and the first elected legislative assembly in the New World. It was established on July 30, ...
in 2013.


Research

Hoffman was recognized as the world's leading authority of
millipedes Millipedes (originating from the Latin , "thousand", and , "foot") are a group of arthropods that are characterised by having two pairs of jointed legs on most body segments; they are known scientifically as the class Diplopoda, the name derive ...
of his time. He authored or co-authored at least 380 scientific papers on millipedes, and described over 400 species and subspecies, as well as over 200
genera Genus (; : genera ) is a taxonomic rank above species and below family as used in the biological classification of living and fossil organisms as well as viruses. In binomial nomenclature, the genus name forms the first part of the binomial s ...
. In 1958, Hoffman co-authored with Ralph V. Chamberlin a checklist of millipedes of North America, the first such work since 1893, which represented an approximate 600% increase in species recorded. Four decades later, Hoffman's 1999 "Checklist of the millipeds of North and Middle America" built upon on his previous work with Chamberlin, as well as checklists of the Caribbean and Central America by Chamberlin and Loomis respectively, to include over 2,100 species. Hoffman founded and edited the journal ''Myriapodologica'' from 1978 to 2007. He is commemorated in the names of two families, six genera, 19 species, and two subspecies of millipedes, as well as two species of
centipede Centipedes (from Neo-Latin , "hundred", and Latin , "foot") are predatory arthropods belonging to the class Chilopoda (Ancient Greek , ''kheilos'', "lip", and Neo-Latin suffix , "foot", describing the forcipules) of the subphylum Myriapoda, ...
. He coined new names for anatomical structures, including the terms " ozadene and "
ozopore An ozopore is the opening of a defensive gland present in some arthropods, notably in millipedes of the order Polydesmidaocellarium" for the patches of ocelli on each side of the head, also known as ocular fields or "eyes". He also published on fossil millipedes, describing extinct groups and producing a "classic summary" of the millipede fossil record for the ''
Treatise on Invertebrate Paleontology The ''Treatise on Invertebrate Paleontology,'' published from 1953–2007 by the Geological Society of America and the University of Kansas, then 2009–present by the University of Kansas Paleontological Institute, is a definitive multi-authore ...
''. In 2007, a
festschrift In academia, a ''Festschrift'' (; plural, ''Festschriften'' ) is a book honoring a respected person, especially an academic, and presented during their lifetime. It generally takes the form of an edited volume, containing contributions from the h ...
consisting of papers from 41 authors on four continents was assembled in honor of Hoffman's 80th birthday. Hoffman also wrote over 70 papers on the natural history and conservation of reptiles and amphibians. The valley and ridge salamander (''
Plethodon hoffmani ''Plethodon'' is a genus of salamanders in the family Plethodontidae. They are commonly known as woodland salamanders. All members of the genus are endemic to North America (Canada and the United States). They have no aquatic larval stage. In som ...
)'' is named in his honor. Hoffman also studied insects,
mollusks Mollusca is a phylum of protostomic invertebrate animals, whose members are known as molluscs or mollusks (). Around 76,000 extant species of molluscs are recognized, making it the second-largest animal phylum after Arthropoda. The num ...
,
crustaceans Crustaceans (from Latin meaning: "those with shells" or "crusted ones") are invertebrate animals that constitute one group of Arthropod, arthropods that are traditionally a part of the subphylum Crustacea (), a large, diverse group of mainly aquat ...
,
arachnids Arachnids are arthropods in the class Arachnida () of the subphylum Chelicerata. Arachnida includes, among others, spiders, scorpions, ticks, mites, pseudoscorpions, harvestmen, camel spiders, whip spiders and vinegaroons. Adult arachnids ...
, and other invertebrates, and co-founded ''The Insects of Virginia'', a series of occasional publications which ran from 1969 to 2006. Nine insects, three crustaceans, a
nematode The nematodes ( or ; ; ), roundworms or eelworms constitute the phylum Nematoda. Species in the phylum inhabit a broad range of environments. Most species are free-living, feeding on microorganisms, but many are parasitic. Parasitic worms (h ...
, an
annelid The annelids (), also known as the segmented worms, are animals that comprise the phylum Annelida (; ). The phylum contains over 22,000 extant species, including ragworms, earthworms, and leeches. The species exist in and have adapted to vario ...
, and a genus of mites have been named after him.


Influences

Roble and Mitchell write that Hoffman's meeting with Hobbs as an undergraduate was the "most significant event in shaping Richard's future career", from whom he learned how to be a good scientist, and whom Hoffman later considered his only true mentor. Hoffman also expressed admiration for the late Dutch biologist C. A. W. Jeekel, whose early correspondence and subsequent friendship affected Hoffman's approach to taxonomy. Hoffman wrote that Jeekel's 1971 ''Nomenclator Generum et Familiarum Diplopodorum'' "set the standard for milliped systematics in the era that is now ending" and that Jeekel was somewhat a role model, and "the only person who I conceded to be my master in our field of interest". Despite working with Chamberlin early in his career, Hoffman later described the period as "under the malign influence of R.V. Chamberlin, an exemplar of minimal taxonomy", while Hoffman favored rich descriptions synthesizing previous relevant literature rather than scant, isolated descriptions.


Major works

* * * * * * * * *


Eponymous taxa

The following is a selection of taxa named after Hoffman. * Hoffmaneumatidae Golovatch, 1978 (a family of millipedes in the order
Chordeumatida Chordeumatida (from the Greek word for "sausage") is a large order of millipedes containing more than 1,400 species. Also known as sausage millipedes, they are found nearly worldwide. Chordeumatida is the largest order in the superorder Nemato ...
) **'' Hoffmaneuma'' Golovatch, 1978 (a millipede genus in the family Hoffmaneumatidae) * Hoffmanobolidae Shelley, 2001 (a
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...
family of millipedes the order
Spirobolida Spirobolida is an Order (biology), order of "round-backed" millipedes containing approximately 500 species in 12 family (biology), families. Its members are distinguished by the presence of a "pronounced Suture (anatomy), suture that runs "verti ...
) *'' Hoffmanides'' Kraus, 1966 (a millipede genus in the family Odontopygidae) *'' Hoffmanina'' Jeekel, 1968 (a millipede genus in the family
Paradoxosomatidae Paradoxosomatidae, the only family in the suborder Paradoxosomatidea (also known as Strongylosomatidea), is a family (biology), family of flat-backed millipedes in the order Polydesmida. Containing nearly 200 genera and 975 species , it is one o ...
) *'' Hoffmanodesmus'' Schubart, 1962 (a millipede genus in the family Chelodesmidae) *'' Andrognathus hoffmani'' Shear & Marek 2009 (a millipede in the family
Andrognathidae Andrognathidae is a family of millipede in the order Platydesmida. There are about 17 genera and more than 30 described species in Andrognathidae. Genera These 17 genera belong to the family Andrognathidae: * ''Andrognathus'' Cope, 1869 * ''Baz ...
) *''
Nannarrup hoffmani ''Nannarrup'' is a genus of soil centipedes in the family Mecistocephalidae. This genus contains only three species, including the type species '' Nannarrup hoffmani''. Also known as Hoffman's dwarf centipede, ''N. hoffmani'' was discovered in ...
'' Foddai, Bonato, Pereira & Minelli, 2003 (a soil centipede discovered in New York City's
Central Park Central Park is an urban park between the Upper West Side and Upper East Side neighborhoods of Manhattan in New York City, and the first landscaped park in the United States. It is the List of parks in New York City, sixth-largest park in the ...
) *''
Plethodon hoffmani ''Plethodon'' is a genus of salamanders in the family Plethodontidae. They are commonly known as woodland salamanders. All members of the genus are endemic to North America (Canada and the United States). They have no aquatic larval stage. In som ...
'' Highton, 1972 (a salamander in the family
Plethodontidae Plethodontidae, or lungless salamanders, are a family of salamanders. With over 500 species, lungless salamanders are by far the largest family of salamanders in terms of their diversity. Most species are native to the Western Hemisphere, from B ...
) *'' Eucarlia hoffmani'' Golovatch & Korsós, 1992 *'' Chaleponcus vilici'' Enghoff, 2014


See also

*
Ecology of the Appalachian Mountains The Appalachian Mountains, often called the Appalachians, are a mountain range in eastern to northeastern North America. The term "Appalachian" refers to several different regions associated with the mountain range, and its surrounding terrain ...
*
Peaks of Otter salamander The Peaks of Otter salamander (''Plethodon hubrichti'') is a species of salamanders in the family Plethodontidae. It is endemic to the Peaks of Otter area in the Blue Ridge Mountains of Virginia. It is a montane salamander found at elevations abo ...
, which Hoffman had a role in discovering *
Environment of Virginia The natural environment of Virginia encompasses the physical geography and biology of the U.S. state of Virginia. Virginia has a total area of , including of water, making it the 35th- largest state by area. Forests cover 65% of the state, wetlan ...


References


External links


Myriapodological works by Richard L. Hoffman
compiled by the Centre International de Myriapodologie *
''Myriapodologica'' collected papers
{{DEFAULTSORT:Hoffman, Richard Lawrence 1927 births 2012 deaths American entomologists American herpetologists American people of German descent Cornell University College of Agriculture and Life Sciences alumni University of Virginia alumni Virginia Tech alumni Radford University faculty People from Clifton Forge, Virginia People from Martinsville, Virginia Myriapodologists Scientists from Virginia