Richard Lawrence Hoffman (September 25, 1927 – June 10, 2012) was an American
zoologist
Zoology ()The pronunciation of zoology as is usually regarded as nonstandard, though it is not uncommon. is the branch of biology that studies the animal kingdom, including the structure, embryology, evolution, classification, habits, and ...
known as an international expert on
millipedes
Millipedes are a group of arthropods that are characterised by having two pairs of jointed legs on most body segments; they are known scientifically as the class Diplopoda, the name derived from this feature. Each double-legged segment is a resu ...
Virginia
Virginia, officially the Commonwealth of Virginia, is a state in the Mid-Atlantic and Southeastern regions of the United States, between the East Coast of the United States, Atlantic Coast and the Appalachian Mountains. The geography an ...
and the
Appalachian Mountains
The Appalachian Mountains, often called the Appalachians, (french: Appalaches), are a system of mountains in eastern to northeastern North America. The Appalachians first formed roughly 480 million years ago during the Ordovician Period. The ...
. He was a biology professor at Virginia's
Radford College
Radford College is an independent school, Anglican, coeducational day school, located in Bruce, Australian Capital Territory, Australia.
Founded in 1984, the college is named after Bishop Lewis Bostock Radford. It has a non-selective enrolme ...
for almost thirty years, and curator of invertebrates at the
Virginia Museum of Natural History
The Virginia Museum of Natural History is the state's natural history museum located in Martinsville, Virginia founded in 1984. The museum has several different award-winning publications, is affiliated with the Smithsonian Institution, and has ...
for another twenty years. He co-founded the Virginia Natural History Society, described over 400 species of millipedes, and produced more than 480 scientific publications. He is commemorated in the scientific and/or common names of over 30 animal species, including the
valley and ridge salamander
The valley and ridge salamander (''Plethodon hoffmani'') is a species of salamander in the family Plethodontidae endemic to the Appalachian Mountains in the eastern United States.
Etymology
The species is named after biologist Richard Hoffman o ...
Clifton Forge, Virginia
Clifton Forge is a town in Alleghany County, Virginia, United States which is part of the greater Roanoke Region. The population was 3,555 at the 2020 census. The Jackson River flows through the town, which as a result was once known as Jac ...
, on September 25, 1927. His father was a railroad machinist whose parents had emigrated from Germany in the late 1800s, and his mother was from a Virginia farming family. Hoffman's scientific publications began at the age of 16, and by the age of 24 he had 29 publications. He attended the
University of Virginia
The University of Virginia (UVA) is a public research university in Charlottesville, Virginia. Founded in 1819 by Thomas Jefferson, the university is ranked among the top academic institutions in the United States, with College admission ...
until 1950, where he was influenced by
Horton H. Hobbs, Jr.
Horton Holcombe Hobbs Jr. (March 29, 1914 – March 22, 1994) was an American taxonomist and carcinologist, specialising in freshwater decapods. He was also a capable artist, musician, cook and botanist.
Hobbs was born in Alachua County, Florid ...
a crustacean biologist and recently hired professor. Unable to complete a freshman math class, Hoffman did not complete a degree at Virginia. Rather, in 1951 he was accepted into graduate school at
Cornell University
Cornell University is a private statutory land-grant research university based in Ithaca, New York. It is a member of the Ivy League. Founded in 1865 by Ezra Cornell and Andrew Dickson White, Cornell was founded with the intention to ...
, who accepted his publication record as equivalent to passing a math class. When not working in the field or at Cornell, Hoffman worked as a chemist at the
Radford Arsenal
Radford Army Ammunition Plant (RFAAP) is an ammunition manufacturing complex for the U.S. military with facilities located in Pulaski and Montgomery Counties, Virginia. The primary mission of the RFAAP is to manufacture propellants and explosives ...
. He completed his a M.S. in
entomology
Entomology () is the scientific study of insects, a branch of zoology. In the past the term "insect" was less specific, and historically the definition of entomology would also include the study of animals in other arthropod groups, such as arach ...
in 1959, and went on to earn a PhD in Zoology from
Virginia Tech
Virginia Tech (formally the Virginia Polytechnic Institute and State University and informally VT, or VPI) is a Public university, public Land-grant college, land-grant research university with its main campus in Blacksburg, Virginia. It also ...
in 1960, during which he revised the genus '' Cambarincola'', a group of leech-like
oligochaete
Oligochaeta () is a subclass of animals in the phylum Annelida, which is made up of many types of aquatic and terrestrial worms, including all of the various earthworms. Specifically, oligochaetes comprise the terrestrial megadrile earthworm ...
worms that are
commensal
Commensalism is a long-term biological interaction ( symbiosis) in which members of one species gain benefits while those of the other species neither benefit nor are harmed. This is in contrast with mutualism, in which both organisms benefit fr ...
on
crayfish
Crayfish are freshwater crustaceans belonging to the clade Astacidea, which also contains lobsters. In some locations, they are also known as crawfish, craydids, crawdaddies, crawdads, freshwater lobsters, mountain lobsters, rock lobsters, ...
. His doctoral advisor at Virginia Tech was Dr. Perry C. Holt.
After completing his PhD, Hoffman worked as professor of biology at
Radford College
Radford College is an independent school, Anglican, coeducational day school, located in Bruce, Australian Capital Territory, Australia.
Founded in 1984, the college is named after Bishop Lewis Bostock Radford. It has a non-selective enrolme ...
(now known as Radford University) from 1960 to 1988. Over this 28-year period, Hoffman taught a number of courses including introductory biology and zoology,
invertebrate zoology Invertebrate zoology is the subdiscipline of zoology that consists of the study of invertebrates, animals without a backbone (a structure which is found only in fish, amphibians, reptiles, birds and mammals).
Invertebrates are a vast and very ...
, entomology, herpetology, mammalogy, biogeography, and the history of science. From 1964 to 1970 Hoffman edited ''The Radford Review'', the college's scholarly journal. Hoffman made dramatic improvements to the collections of the Radford Natural History Museum. From 1989 to 2009, Hoffman worked as Curator of Recent Invertebrates at the
Virginia Museum of Natural History
The Virginia Museum of Natural History is the state's natural history museum located in Martinsville, Virginia founded in 1984. The museum has several different award-winning publications, is affiliated with the Smithsonian Institution, and has ...
.
Hoffman died on June 10, 2012 at
Carilion Roanoke Memorial Hospital
Carilion Roanoke Memorial Hospital (CRMH) is a private teaching hospital in Roanoke, Virginia, USA. With 703 beds, Carilion Roanoke Memorial Hospital is one of the largest hospitals in the state. It is part of Carilion Clinic.
The region's onl ...
in
Roanoke, Virginia
Roanoke ( ) is an independent city in the U.S. state of Virginia. At the 2020 census, the population was 100,011, making it the 8th most populous city in the Commonwealth of Virginia and the largest city in Virginia west of Richmond. It is ...
following heart surgery. He was 84. His contributions to science and education were formally recognized by the
Virginia General Assembly
The Virginia General Assembly is the legislative body of the Commonwealth of Virginia, the oldest continuous law-making body in the Western Hemisphere, the first elected legislative assembly in the New World, and was established on July 30, 1 ...
in 2013.
Research
Hoffman was recognized as the world's leading authority of
millipedes
Millipedes are a group of arthropods that are characterised by having two pairs of jointed legs on most body segments; they are known scientifically as the class Diplopoda, the name derived from this feature. Each double-legged segment is a resu ...
of his time. He authored or co-authored at least 380 scientific papers on millipedes, and described over 400 species and subspecies, as well as over 200
genera
Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial ...
. In 1958, Hoffman co-authored with Ralph V. Chamberlin a checklist of millipedes of North America, the first such work since 1893, which represented an approximate 600% increase in species recorded. Four decades later, Hoffman's 1999 "Checklist of the millipeds of North and Middle America" built upon on his previous work with Chamberlin, as well as checklists of the Caribbean and Central America by Chamberlin and Loomis respectively, to include over 2,100 species. Hoffman founded and edited the journal ''Myriapodologica'' from 1978 to 2007. He is commemorated in the names of two families, six genera, 19 species, and two subspecies of millipedes, as well as two species of
centipede
Centipedes (from New Latin , "hundred", and Latin , " foot") are predatory arthropods belonging to the class Chilopoda (Ancient Greek , ''kheilos'', lip, and New Latin suffix , "foot", describing the forcipules) of the subphylum Myriapoda, an ...
. He coined new names for anatomical structures, including the terms " ozadene and "
ozopore
An ozopore is the opening of a defensive gland present in some arthropods, notably in millipedes of the order Polydesmidaocellarium" for the patches of ocelli on each side of the head, also known as ocular fields or "eyes". He also published on fossil millipedes, describing extinct groups and producing a "classic summary" of the millipede fossil record for the ''
Treatise on Invertebrate Paleontology
The ''Treatise on Invertebrate Paleontology'' (or ''TIP'') published by the Geological Society of America and the University of Kansas Press, is a definitive multi-authored work of some 50 volumes, written by more than 300 paleontologists, and co ...
''. In 2007, a
festschrift
In academia, a ''Festschrift'' (; plural, ''Festschriften'' ) is a book honoring a respected person, especially an academic, and presented during their lifetime. It generally takes the form of an edited volume, containing contributions from the ...
consisting of papers from 41 authors on four continents was assembled in honor of Hoffman's 80th birthday.
Hoffman also wrote over 70 papers on the natural history and conservation of reptiles and amphibians. The
valley and ridge salamander
The valley and ridge salamander (''Plethodon hoffmani'') is a species of salamander in the family Plethodontidae endemic to the Appalachian Mountains in the eastern United States.
Etymology
The species is named after biologist Richard Hoffman o ...
(''
Plethodon hoffmani
''Plethodon'' is a genus of salamanders in the family Plethodontidae. They are also known as woodland salamanders or, more rarely, slimy salamanders. All members of the genus are endemic to North America (Canada and USA). They have no aquatic lar ...
)'' is named in his honor. Hoffman also studied insects,
mollusks
Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000 extant species of molluscs are recognized. The number of fossil species is esti ...
,
crustaceans
Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean gr ...
,
arachnids
Arachnida () is a class of joint-legged invertebrate animals (arthropods), in the subphylum Chelicerata. Arachnida includes, among others, spiders, scorpions, ticks, mites, pseudoscorpions, harvestmen, camel spiders, whip spiders and vine ...
, and other invertebrates, and co-founded ''The Insects of Virginia'', a series of occasional publications which ran from 1969 to 2006. Nine insects, three crustaceans, a nematode, an
annelid
The annelids (Annelida , from Latin ', "little ring"), also known as the segmented worms, are a large phylum, with over 22,000 extant species including ragworms, earthworms, and leeches. The species exist in and have adapted to various ecol ...
, and a genus of mites have been named after him.
Influences
Roble and Mitchell write that Hoffman's meeting with Hobbs as an undergraduate was the "most significant event in shaping Richard's future career", from whom he learned how to be a good scientist, and whom Hoffman later considered his only true mentor. Hoffman also expressed admiration for the late Dutch biologist C. A. W. Jeekel, whose early correspondence and subsequent friendship affected Hoffman's approach to taxonomy. Hoffman wrote that Jeekel's 1971 ''Nomenclator Generum et Familiarum Diplopodorum'' "set the standard for milliped systematics in the era that is now ending" and that Jeekel was somewhat a role model, and "the only person who I conceded to be my master in our field of interest". Despite working with Chamberlin early in his career, Hoffman later described the period as "under the malign influence of R.V. Chamberlin, an exemplar of minimal taxonomy", while Hoffman favored rich descriptions synthesizing previous relevant literature rather than scant, isolated descriptions.
Major works
*
*
*
*
*
*
*
*
*
Eponymous taxa
The following is a selection of taxa named after Hoffman.
* Hoffmaneumatidae Golovatch, 1978 (a family of millipedes in the order
Chordeumatida
Chordeumatida (from the Greek word for "sausage") is a large order of millipedes containing some 1200 species with a nearly worldwide distribution. Also known as "sausage millipedes," they possess around 30 body segments behind the head (includi ...
)
**'' Hoffmaneuma'' Golovatch, 1978 (a millipede genus in the family Hoffmaneumatidae)
*
Hoffmanobolidae
Hoffmanobolidae is a family of millipedes belonging to the order Spirobolida
Spirobolida is an order of "round-backed" millipedes containing approximately 500 species in 12 families. Its members are distinguished by the presence of a "pronou ...
Shelley, 2001 (a
monotypic
In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispec ...
family of millipedes the order
Spirobolida
Spirobolida is an Order (biology), order of "round-backed" millipedes containing approximately 500 species in 12 family (biology), families. Its members are distinguished by the presence of a "pronounced Suture (anatomy), suture that runs "verti ...
)
*'' Hoffmanides'' Kraus, 1966 (a millipede genus in the family
Odontopygidae
Odontopygidae is a family of millipedes belonging to the order Spirostreptida
Spirostreptida is an order (biology), order of long, cylindrical millipedes. There are approximately 1000 described species, making Spirostreptida the third largest ...
)
*'' Hoffmanina'' Jeekel, 1968 (a millipede genus in the family
Paradoxosomatidae
Paradoxosomatidae, the only family in the suborder Paradoxosomatidea (also known as Strongylosomatidea), is a family of flat-backed millipedes in the order Polydesmida. Containing nearly 200 genera and 975 species , it is one of the largest fa ...
)
*'' Hoffmanodesmus'' Schubart, 1962 (a millipede genus in the family
Chelodesmidae
Chelodesmidae, is a millipede family of order Polydesmida. The family includes 219 genera. Two new genera were described in 2012.
Genera
A
*'' Achromoporus''
*'' Afolabina''
*'' Alassodesmus''
*'' Allarithmus''
*'' Alocodesmus''
*''Alys ...
Andrognathidae
Andrognathidae is a family of millipede in the order Platydesmida. There are about 17 genera and more than 30 described species in Andrognathidae.
Genera
These 17 genera belong to the family Andrognathidae:
* '' Andrognathus'' Cope, 1869
* '' ...
)
*''
Nannarrup hoffmani
''Nannarrup hoffmani'', commonly known as Hoffman's dwarf centipede, is a species of centipede that was discovered in New York City's Central Park
Central Park is an urban park in New York City located between the Upper West Side, Upper West ...
'' Foddai, Bonato, Pereira & Minelli, 2003 (a
soil centipede
Centipedes (from New Latin , "hundred", and Latin , "foot") are predatory arthropods belonging to the class Chilopoda (Ancient Greek , ''kheilos'', lip, and New Latin suffix , "foot", describing the forcipules) of the subphylum Myriapoda, an a ...
discovered in New York City's
Central Park
Central Park is an urban park in New York City located between the Upper West Side, Upper West and Upper East Sides of Manhattan. It is the List of New York City parks, fifth-largest park in the city, covering . It is the most visited urban par ...
)
*''
Plethodon hoffmani
''Plethodon'' is a genus of salamanders in the family Plethodontidae. They are also known as woodland salamanders or, more rarely, slimy salamanders. All members of the genus are endemic to North America (Canada and USA). They have no aquatic lar ...
'' Highton, 1972 (a salamander in the family
Plethodontidae
Plethodontidae, or lungless salamanders, are a family of salamanders. Most species are native to the Western Hemisphere, from British Columbia to Brazil, although a few species are found in Sardinia, Europe south of the Alps, and South Korea. In ...
Ecology of the Appalachian Mountains
The Appalachian Mountains, often called the Appalachians, (french: Appalaches), are a mountain range, system of mountains in eastern to northeastern North America. The Appalachians first formed roughly 480 million years ago during the Ordovicia ...
*
Peaks of Otter salamander
The Peaks of Otter salamander (''Plethodon hubrichti'') is a species of salamanders in the family Plethodontidae. It is endemic to the Peaks of Otter area in the Blue Ridge Mountains of Virginia. It is a montane salamander found at elevation ...
, which Hoffman had a role in discovering
*
Environment of Virginia
The natural environment of Virginia encompasses the physical geography and biology of the U.S. state of Virginia. Virginia has a total area of , including of water, making it the 35th- largest state by area. Forests cover 65% of the state, wetla ...