Richard Heymons (29 May 1867 – 1 December 1943) was a German

zoologist Zoology ()The pronunciation of zoology as is usually regarded as nonstandard, though it is not uncommon. is the branch of biology that studies the animal kingdom, including the structure, embryology, evolution, classification, habits, and ...

and

entomologist Entomology () is the scientific study of insects, a branch of zoology. In the past the term "insect" was less specific, and historically the definition of entomology would also include the study of animals in other arthropod groups, such as arach ...

. He studied in

Humboldt University of Berlin The Humboldt University of Berlin (german: link=no, Humboldt-Universität zu Berlin, abbreviated HU Berlin) is a public university, public research university in the central borough of Mitte in Berlin, Germany. The university was established ...

from 1886 to 1891 and provided overall direction of the Institute of Zoology at the higher educational farm in Berlin from 1915 to 1935. Heymons actively studied tongue worms (

Pentastomida The Pentastomida are an enigmatic group of parasitic arthropods commonly known as tongue worms due to the resemblance of the species of the genus ''Linguatula'' to a vertebrate tongue; molecular studies point to them being degenerate crustaceans. ...

) and his collection of these animals is held in the

Museum für Naturkunde The Natural History Museum (german: Museum für Naturkunde) is a natural history museum located in Berlin, Germany. It exhibits a vast range of specimens from various segments of natural history and in such domain it is one of three major muse ...

Berlin. He also coined the name

Chelicerata The subphylum Chelicerata (from New Latin, , ) constitutes one of the major subdivisions of the phylum Arthropoda. It contains the sea spiders, horseshoe crabs, and arachnids (including harvestmen, scorpions, spiders, solifuges, ticks, and mite ...

for arachnids and their relatives.

Publications

Monographs

* ''The development of the female reproductive organs of Phyllodromica (Blatta) germanica L.'' 1891 * ''On the origin of the sex cells in insects'' . 1893 * ''The segmentation of the insect body'' . 1895. (2010, ) * ''The embryonic development of dermaptera (

earwig Earwigs make up the insect order Dermaptera. With about 2,000 species in 12 families, they are one of the smaller insect orders. Earwigs have characteristic cerci, a pair of forcep-like pincers on their abdomen, and membranous wings fol ...

s ) and orthoptera (

locust Locusts (derived from the Vulgar Latin ''locusta'', meaning grasshopper) are various species of short-horned grasshoppers in the family Acrididae that have a swarming phase. These insects are usually solitary, but under certain circumst ...

s ) – with special reference to cotyledon formation'' . 1895. (2009, ) * ''Principles of development and physique of Odonata (

Odonata Odonata is an order of flying insects that includes the dragonflies and damselflies. Members of the group first appeared during the Triassic, though members of their total group, Odonatoptera, first appeared in Late Carboniferous. The two co ...

) and Ephemeris (

Mayflies Mayflies (also known as shadflies or fishflies in Canada and the upper Midwestern United States, as Canadian soldiers in the American Great Lakes region, and as up-winged flies in the United Kingdom) are aquatic insects belonging to the order ...

)'' . 1896 * ''A contribution to the evolutionary history of the Insecta apterygota (flying

insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (head, thorax and abdomen), three pairs ...

s )'' . 1896 * ''On the morphology of the abdominal appendages in insects'' . 1896 * ''On the organization and development of Bacillus rossii ( Bacillus rossius ) Fabr'' . 1897 * ''Contributions to the

morphology Morphology, from the Greek and meaning "study of shape", may refer to: Disciplines *Morphology (archaeology), study of the shapes or forms of artifacts *Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies, ...

and evolutionary history of the rhynchotes'' . 1899 * ''Biological observations on an Asian solifugae (

Pholcidae The Pholcidae are a family of araneomorph spiders. The family contains over 1,800 individual species of pholcids, including those commonly known as cellar spider, daddy long-legs spider, carpenter spider, daddy long-legger, vibrating spi ...

) together with contributions to the systematics of the same'' . 1901 * ''The evolutionary history of the scolopendres (

Scolopendra ''Scolopendra'' (through Latin from Greek , ''skolopendra'') is a species-rich genus of large tropical centipedes of the family Scolopendridae. Description The genus ''Scolopendra'' contains many species of centipedes found across the world' ...

)'' . 1901. (2011, ) * ''The abdominal appendages of

dragonflies A dragonfly is a flying insect belonging to the infraorder Anisoptera below the order Odonata. About 3,000 extant species of true dragonfly are known. Most are tropical, with fewer species in temperate regions. Loss of wetland habitat threaten ...

and their larvae'' . 1904 * ''A placenta in an insect (

Hemimerus ''Hemimerus'' is a genus of earwig Earwigs make up the insect order Dermaptera. With about 2,000 species in 12 families, they are one of the smaller insect orders. Earwigs have characteristic cerci, a pair of forcep-like pincers on th ...

)'' . 1909

Essays

* ''Contribution to the systematics and morphology of Tongue Worms (Pentastomida)'' . In: ''Zoological Gazette'' , 1922, 55, pp. 154–167. * ''About the dorsal organ of the pentastomida.'' In: ''Meeting reports of the Society of Friends of Natural Sciences in Berlin.'' 1926, pp. 22–24. * ''Pentastomida.'' In: W. Kükenthal, T. Krumbach (eds.): ''Handbook of Zoology; a natural history of the animal kingdom.'' 3 (1). Walter de Gruyter, Berlin 1926, pp. 69–128. * ''Order: Tongue Worms, Pentastomida (Linguatula).'' In: P. Brohmer, P. Ehrmann, G. Ulmer (eds.): ''The animal kingdom of Central Europe.'' 3 (1). Quelle & Meyer, Leipzig 1928, p. xx. * ''Pentastomida of the German Limnological Sunda Expedition.'' In: ''Archive for Hydrobiology.'' 1930. Supplement Volume 3, pp. 193–198. * ''A contribution to the knowledge of the pentastomids of Australia and neighboring areas.'' In: ''Journal of parasitology.'' 4, 1932, pp. 409–430. * ''A New Pentastomida from the Philippines.'' – Zoological Indicator 97, 1932, 295–299. * ''About an abnormality in a pentastomida (Armillifer moniliformis Diesing).'' Meeting reports of the Society of Friends of Natural Sciences in Berlin 3, 1932, 287–290. * ''About the occurrence of Tongue Worms in birds.'' – Ornithological Monthly Reports 41, 1933, 75–76. * with H. Graf Vitzthum: ''New and little-known pentastomids from America.'' Zoological Indicator 109, 1935, 150–158. * ''Pentastomida.'' In: ''Dr. HG Bronn's classes and orders of the animal kingdom.'' Fifth volume. IV Division, 1st Book. Academic Publishing Company, Leipzig 1935, pp. 1–268. * with H. Graf Vitzthum: ''Contributions to the systematics of pentastomids.'' Journal of Parasitics 8, 1936, 1–103. * ''The one from Dr. Edm. Dartevelle in the Belgian Congo State collected Pentastomida.'' In: ''Revue de Zoologie et de Botanique Africaines.'' 33, 1940, pp. 122–123. * ''Contributions to the systematics of the pentastomids II. Some remarkable pentastomids from Lacertilia.'' Journal of Parasitics 10, 1939, 675–690. * ''Contributions to the systematics of the Pentastomids III. Pentastomids with spirally curved body shapes.'' Journal of Parasitics 11, 1939, 77–94. * ''On the habits of the Pentastomida found in crocodiles'' . Report of the meeting of the Society of Friends of Natural Sciences in Berlin, 1940, pp. 253–269. * ''Contributions to the systematics of the pentastomids IV. To the knowledge of the Sambonidae'' . In: ''Journal of Parasitics'' , 12, 1941, pp. 317–329. * ''Contributions to the systematics of the pentastomids V. The type specimens of Diesingia megastoma'' . In: ''Journal of Parasitics'' , 12, 1941, pp. 330–339. * ''Contributions to the systematics of the pentastomids. VI. The species of the genus Alofia in comparison with Sebekia. I. Overview of the species'' . In: ''Journal of Parasitics'' , 12, 1941, pp. 419–432. * ''The canine nasal worms (Linguatula serrata Froelich), its hosts and relationships to European wildlife, its origin and practical importance based on our current knowledge.'' In: ''Journal of parasitology.'' 12, 1942, pp. 607–638. * ''Pentastomida.'' In: Institut des Parcs Nationaux du Congo Belge. Exploration du Parc National Albert. Mission GF de Witte 43, 1943, pp. 3–4

Sources

* https://www.idref.fr/138918112 * https://www.hu-berlin.de/de/ * https://www.uni-goettingen.de/de/geschichte/71626.html * https://www.biodiversitylibrary.org/creator/3857#/titles * https://www.leopoldina.org/members/members directory/member/3822/ * https://www.jstor.org/stable/41768001?seq=1 {{DEFAULTSORT:Heymons, Richard 1867 births 1943 deaths 20th-century German zoologists 19th-century German zoologists