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A claw is a curved, pointed appendage found at the end of a toe or finger in most
amniote Amniotes are tetrapod vertebrate animals belonging to the clade Amniota, a large group that comprises the vast majority of living terrestrial animal, terrestrial and semiaquatic vertebrates. Amniotes evolution, evolved from amphibious Stem tet ...
s (
mammals A mammal () is a vertebrate animal of the class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three middle e ...
, reptiles,
birds Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
). Some
invertebrate Invertebrates are animals that neither develop nor retain a vertebral column (commonly known as a ''spine'' or ''backbone''), which evolved from the notochord. It is a paraphyletic grouping including all animals excluding the chordata, chordate s ...
s such as
beetles Beetles are insects that form the Taxonomic rank, order Coleoptera (), in the superorder Holometabola. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 40 ...
and spiders have somewhat similar fine, hooked structures at the end of the leg or tarsus for gripping a surface as they walk. The pincers of crabs, lobsters and scorpions, more formally known as their chelae, are sometimes called claws. A true claw is made of a hard
protein Proteins are large biomolecules and macromolecules that comprise one or more long chains of amino acid residue (biochemistry), residues. Proteins perform a vast array of functions within organisms, including Enzyme catalysis, catalysing metab ...
called
keratin Keratin () is one of a family of structural fibrous proteins also known as ''scleroproteins''. It is the key structural material making up Scale (anatomy), scales, hair, Nail (anatomy), nails, feathers, horn (anatomy), horns, claws, Hoof, hoove ...
. Claws are used to catch and hold prey in
carnivorous A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour"), is an animal or plant whose nutrition and energy requirements are met by consumption of animal tissues (mainly mu ...
mammals A mammal () is a vertebrate animal of the class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three middle e ...
such as
cats The cat (''Felis catus''), also referred to as the domestic cat or house cat, is a small domesticated carnivorous mammal. It is the only domesticated species of the family Felidae. Advances in archaeology and genetics have shown that the ...
and dogs, but may also be used for such purposes as digging, climbing trees, self-defense and grooming, in those and other species. Similar appendages that are flat and do not come to a sharp point are called nails instead. Claw-like projections that do not form at the end of digits but spring from other parts of the foot are properly named
spur A spur is a metal tool designed to be worn in pairs on the heels of riding boots for the purpose of directing a horse or other animal to move forward or laterally while riding. It is usually used to refine the riding aids (commands) and to ba ...
s.


Tetrapods

In
tetrapod A tetrapod (; from Ancient Greek :wiktionary:τετρα-#Ancient Greek, τετρα- ''(tetra-)'' 'four' and :wiktionary:πούς#Ancient Greek, πούς ''(poús)'' 'foot') is any four-Limb (anatomy), limbed vertebrate animal of the clade Tetr ...
s, claws are made of
keratin Keratin () is one of a family of structural fibrous proteins also known as ''scleroproteins''. It is the key structural material making up Scale (anatomy), scales, hair, Nail (anatomy), nails, feathers, horn (anatomy), horns, claws, Hoof, hoove ...
and consist of two layers. The unguis is the harder external layer, which consists of keratin fibers arranged perpendicular to the direction of growth and in layers at an oblique angle. The subunguis is the softer, flaky underside layer whose grain is parallel to the direction of growth. The claw grows outward from the nail matrix at the base of the unguis and the subunguis grows thicker while travelling across the nail bed. The unguis grows outward faster than the subunguis to produce a curve and the thinner sides of the claw wear away faster than their thicker middle, producing a more or less sharp point. Tetrapods use their claws in many ways, commonly to grasp or kill prey, to dig and to climb and hang.


Mammals

All
carnivora Carnivora ( ) is an order of placental mammals specialized primarily in eating flesh, whose members are formally referred to as carnivorans. The order Carnivora is the sixth largest order of mammals, comprising at least 279 species. Carnivor ...
ns have claws, which vary considerably in length and shape. Claws grow out of the third phalanges of the paws and are made of
keratin Keratin () is one of a family of structural fibrous proteins also known as ''scleroproteins''. It is the key structural material making up Scale (anatomy), scales, hair, Nail (anatomy), nails, feathers, horn (anatomy), horns, claws, Hoof, hoove ...
. Many
predator Predation is a biological interaction in which one organism, the predator, kills and eats another organism, its prey. It is one of a family of common List of feeding behaviours, feeding behaviours that includes parasitism and micropredation ...
y mammals have protractile claws that can partially hide inside the animal's paw, especially the cat family,
Felidae Felidae ( ) is the Family (biology), family of mammals in the Order (biology), order Carnivora colloquially referred to as cats. A member of this family is also called a felid ( ). The 41 extant taxon, extant Felidae species exhibit the gre ...
, almost all of whose members have fully protractible claws. Outside of the cat family, retractable claws are found only in certain species of the
Viverridae Viverridae is a family (biology), family of small to medium-sized feliform mammals, comprising 14 genera with 33 species. This family was named and first described by John Edward Gray in 1821. Viverrids occur all over Africa, in southern Europe, ...
(and the extinct
Nimravidae Nimravidae is an extinct family (biology), family of carnivorans, sometimes known as false saber-toothed cats, whose fossils are found in North America and Eurasia. Not considered to belong to the true cats (family Felidae), the nimravids are gen ...
). A claw that is retractable is protected from wear and tear. Most cats and dogs also have a
dewclaw A dewclaw is a digit – vestigial in some animals – on the foot of many mammals, birds, and reptiles (including some extinct orders, like certain theropods). It commonly grows higher on the leg than the rest of the foot, such that in di ...
on the inside of the front paws. It is much less functional than the other claws but does help the cats to grasp prey. Because the dew claw does not touch the ground, it receives less wear and tends to be sharper and longer. A nail is homologous to a claw but is flatter and has a curved edge instead of a point. A nail that is big enough to bear weight is called a " hoof". (Nevertheless, one side of the cloven-hoof of
artiodactyl Artiodactyls are placental mammals belonging to the order Artiodactyla ( , ). Typically, they are ungulates which bear weight equally on two (an even number) of their five toes (the third and fourth, often in the form of a hoof). The other t ...
ungulate Ungulates ( ) are members of the diverse clade Euungulata ("true ungulates"), which primarily consists of large mammals with Hoof, hooves. Once part of the clade "Ungulata" along with the clade Paenungulata, "Ungulata" has since been determined ...
s may also be called a claw). Every so often, the growth of claws stops and restarts, as does
hair Hair is a protein filament that grows from follicles found in the dermis. Hair is one of the defining characteristics of mammals. The human body, apart from areas of glabrous skin, is covered in follicles which produce thick terminal and ...
. In a hair, this results in the hair falling out and being replaced by a new one. In claws, this results in an abscission layer, and the old segment breaks off. This process takes several months for
human Humans (''Homo sapiens'') or modern humans are the most common and widespread species of primate, and the last surviving species of the genus ''Homo''. They are Hominidae, great apes characterized by their Prehistory of nakedness and clothing ...
thumbnails. Cats are often seen working old unguis layers off on wood or on boards made for the purpose. Ungulates' hooves wear or self-trim by ground contact. Domesticated equids (
horse The horse (''Equus ferus caballus'') is a domesticated, one-toed, hoofed mammal. It belongs to the taxonomic family Equidae and is one of two extant subspecies of ''Equus ferus''. The horse has evolved over the past 45 to 55 mi ...
s,
donkey The donkey or ass is a domesticated equine. It derives from the African wild ass, ''Equus africanus'', and may be classified either as a subspecies thereof, ''Equus africanus asinus'', or as a separate species, ''Equus asinus''. It was domes ...
s and
mule The mule is a domestic equine hybrid between a donkey, and a horse. It is the offspring of a male donkey (a jack) and a female horse (a mare). The horse and the donkey are different species, with different numbers of chromosomes; of the two ...
s) usually need regular trimming by a farrier, as a consequence of reduced activity on hard ground.


Primates

Primate Primates is an order (biology), order of mammals, which is further divided into the Strepsirrhini, strepsirrhines, which include lemurs, galagos, and Lorisidae, lorisids; and the Haplorhini, haplorhines, which include Tarsiiformes, tarsiers a ...
nails consist of the unguis alone, as the subunguis has disappeared. With the evolution of grasping hands and feet, claws are no longer necessary for locomotion, and instead most digits exhibit nails. However, claw-like nails are found in small-bodied callitrichids on all digits except the hallux or big toe. A laterally flattened grooming claw, used for grooming, can be found on the second toe in living strepsirrhines, and the second and third in tarsiers.
Aye-aye The aye-aye (''Daubentonia madagascariensis'') is a long-fingered lemur, a Strepsirrhini, strepsirrhine primate native to Madagascar with rodent-like teeth that perpetually grow and a special thin middle finger that they can use to catch grubs ...
s have functional claws on all other digits except the hallux, including a grooming claw on the second toe. Less commonly known, a grooming claw is also found on the second pedal digit of night monkeys (''Aotus''), titis (''Callicebus''), and possibly other New World monkeys.


Reptiles

Most reptiles have well-developed claws. Most lizards have toes ending in stout claws. In snakes, feet and claws are absent, but in many boids such as ''
Boa constrictor The boa constrictor (scientific name also ''Boa constrictor''), also known as the common boa, is a species of large, non-venomous, heavy-bodied snake that is frequently kept and bred in captivity. The boa constrictor is a member of the Family (b ...
'', remnants of highly reduced hind-limbs emerge with a single claw as "spurs" on each side of the anal opening. Lizard claws are used as aids in climbing, and in holding down prey in carnivorous species.


Birds

A talon is the claw of a
bird of prey Birds of prey or predatory birds, also known as (although not the same as) raptors, are hypercarnivorous bird species that actively predation, hunt and feed on other vertebrates (mainly mammals, reptiles and smaller birds). In addition to speed ...
, its primary hunting tool. The talons are very important; without them, most birds of prey would not be able to catch their food. Some birds also use claws for defensive purposes. Cassowaries use claws on their inner toe (digit I) for defence and have been known to disembowel people. All birds, however, have claws, which are used as general holdfasts and protection for the tip of the digits. The hoatzin and turaco are unique among
extant Extant or Least-concern species, least concern is the opposite of the word extinct. It may refer to: * Extant hereditary titles * Extant literature, surviving literature, such as ''Beowulf'', the oldest extant manuscript written in English * Exta ...
birds in having functional claws on the thumb and index finger (digits I and II) on the forelimbs as chicks, allowing them to climb trees until the adult plumage with flight feathers develop. However, several birds have a claw- or nail-like structure hidden under the feathers at the end of the hand digits, notably ostriches, emus, ducks, geese and kiwis.


Amphibians

The only
amphibian Amphibians are ectothermic, anamniote, anamniotic, tetrapod, four-limbed vertebrate animals that constitute the class (biology), class Amphibia. In its broadest sense, it is a paraphyletic group encompassing all Tetrapod, tetrapods, but excl ...
s to bear claws are the
African clawed frog The African clawed frog (''Xenopus laevis''), also known as simply xenopus, African clawed toad, African claw-toed frog or the ''platanna'') is a species of African Aquatic animal, aquatic frog of the family Pipidae. Its name is derived from the ...
s. Claws evolved separately in the amphibian and
amniote Amniotes are tetrapod vertebrate animals belonging to the clade Amniota, a large group that comprises the vast majority of living terrestrial animal, terrestrial and semiaquatic vertebrates. Amniotes evolution, evolved from amphibious Stem tet ...
( reptiliomorph) line. However, the hairy frog has claw analogues on its feet; the frog intentionally dislocates the tips of its fingers to unsheathe the sharp points of its last
phalanges The phalanges (: phalanx ) are digit (anatomy), digital bones in the hands and foot, feet of most vertebrates. In primates, the Thumb, thumbs and Hallux, big toes have two phalanges while the other Digit (anatomy), digits have three phalanges. ...
.


Arthropods

The scientifically correct term for the "claw" of an arthropod, such as a lobster or crab, is a chela (plural ''chelae''). Legs bearing a chela are called chelipeds. Chelae are also called pincers.


See also

*
Horse hoof A horse hoof is the lower extremity of each leg of a horse, the part that makes contact with the ground and carries the weight of the animal. It is both hard and flexible. It is a complex structure surrounding the distal Phalanx bones, phalanx o ...
* Dactyly


References


External links


Rat Claws
from the original on {{Date, 2022-04-16), also explains much about mammalian claws in general. Animal anatomy