Sciuromorpha ("squirrel-like") is a rodent
clade
A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...
that includes several different rodent families. It includes all members of the
Sciuridae (the squirrel family) as well as the
mountain beaver species.
Traditionally, the term has been defined on the basis of the shape of the
infraorbital canal. A
sciuromorphous zygomasseteric system
The zygomasseteric system (or zygomasseteric structure) in rodents is the anatomical arrangement of the masseter muscle of the jaw and the zygomatic arch of the skull. The anteroposterior or propalinal (front-to-back) motion of the rodent jaw ...
is characterized by attachment of the lateral
masseter muscle along the side of the
rostrum. Unlike
hystricomorphous
The zygomasseteric system (or zygomasseteric structure) in rodents is the anatomical arrangement of the masseter muscle of the jaw and the zygomatic arch of the skull. The anteroposterior or propalinal (front-to-back) motion of the rodent jaw ...
and
myomorphous rodents, the medial masseter muscle does not pass through the infraorbital canal. Among extant rodents, only the families Sciuridae,
Castoridae,
Heteromyidae, and
Geomyidae are truly sciuromorphous. Some authorities would exclude the Geomyidae and Heteromyidae from that list due to the attachment of the medial masseter directly behind the
zygomatic arch.
Carleton and Musser (2005) redefined the rodent suborders on
morphological and
molecular
A molecule is a group of two or more atoms held together by attractive forces known as chemical bonds; depending on context, the term may or may not include ions which satisfy this criterion. In quantum physics, organic chemistry, and bioche ...
grounds. They defined the Sciuromorpha as including three families, Sciuridae,
Aplodontiidae, and
Gliridae. Of these, only the Sciuridae are truly sciuromorphous. The Aplodontiidae are protrogomorphous and the Gliridae are myomorphous. The connection between Aplodontiidae and Sciuridae has been proposed numerous times in the past. The two families have been united into a common
infraorder (Sciurida) or
superfamily
SUPERFAMILY is a database and search platform of structural and functional annotation for all proteins and genomes. It classifies amino acid sequences into known structural domains, especially into SCOP superfamilies. Domains are functional, str ...
(Sciuroidea). It has long been suggested that
dormice (Gliridae) are not particularly related to the
Myomorpha, and their zygomasseteric structure has been termed "pseudomyomorphy". The connection between squirrels and dormice has been almost exclusively suggested through genetic studies, and to a lesser degree via the
fossil rodent ''
Reithroparamys''.
Current families
The suborder Sciuromorpha contains 307 living species in 61 genera and three families. At least three extinct families are also recognised.
* †
Allomyidae
*
Aplodontiidae – mountain beaver
* †
Mylagaulidae
*
Sciuridae – squirrels, chipmunks, marmots, flying squirrels, etc.
* †
Reithroparamyidae
*
Gliridae – dormice
Historical families
*
Castoridae – beavers
*
Geomyidae – pocket gophers
*
Heteromyidae – pocket mice, kangaroo rats, and kangaroo mice
*
Pedetidae – springhares
*
Anomaluridae - anomalures
References
* Carleton, M. D. and G. G. Musser (2005). "Order Rodentia". pp. 745–752 in ''Mammal Species of the World A Taxonomic and Geographic Reference'' (D. E. Wilson and D. M. Reeder eds.). Baltimore: Johns Hopkins University Press.
External links
*
{{Taxonbar, from=Q724477
Rodent taxonomy
Mammal suborders
Extant Eocene first appearances
Taxa named by Johann Friedrich von Brandt