
A reef is a ridge or
shoal
In oceanography, geomorphology, and geoscience, a shoal is a natural submerged ridge, bank, or bar that consists of, or is covered by, sand or other unconsolidated material and rises from the bed of a body of water to near the surface. It ...
of rock,
coral
Corals are marine invertebrates within the class Anthozoa of the phylum Cnidaria. They typically form compact colonies of many identical individual polyps. Coral species include the important reef builders that inhabit tropical oceans and secre ...
or similar relatively stable material, lying beneath the surface of a natural body of water.
Many reefs result from natural,
abiotic processes—
deposition of sand,
wave erosion planing down rock outcrops, etc.—but there are also reefs such as the
coral reef
A coral reef is an underwater ecosystem characterized by reef-building corals. Reefs are formed of colonies of coral polyps held together by calcium carbonate. Most coral reefs are built from stony corals, whose polyps cluster in groups.
C ...
s of tropical waters formed by
biotic
Biotics describe living or once living components of a community; for example organisms, such as animals and plants.
Biotic may refer to:
*Life, the condition of living organisms
*Biology, the study of life
* Biotic material, which is derived from ...
processes dominated by corals and
coralline algae
Coralline algae are red algae in the order Corallinales. They are characterized by a thallus that is hard because of calcareous deposits contained within the cell walls. The colors of these algae are most typically pink, or some other shade of r ...
, and
artificial reef
An artificial reef is a human-created underwater structure, typically built to promote marine life in areas with a generally featureless bottom, to control erosion, block ship passage, block the use of trawling nets, or improve surfing
S ...
s such as shipwrecks and other anthropogenic underwater structures may occur intentionally or as the result of an accident, and sometimes have a designed role in enhancing the physical complexity of featureless sand bottoms, to attract a more diverse assemblage of organisms. Reefs are often quite near to the surface, but not all definitions require this.
Earth's largest coral reef system is the
Great Barrier Reef
The Great Barrier Reef is the world's largest coral reef system composed of over 2,900 individual reefs and 900 islands stretching for over over an area of approximately . The reef is located in the Coral Sea, off the coast of Queensland, ...
in Australia, at a length of over .
Biotic

There is a variety of biotic reef types, including
oyster reefs and
sponge reefs, but the most massive and widely distributed are tropical
coral reef
A coral reef is an underwater ecosystem characterized by reef-building corals. Reefs are formed of colonies of coral polyps held together by calcium carbonate. Most coral reefs are built from stony corals, whose polyps cluster in groups.
C ...
s.
Although corals are major contributors to the framework and bulk material comprising a coral reef; the organisms most responsible for reef growth against the constant assault from ocean waves are
calcareous
Calcareous () is an adjective meaning "mostly or partly composed of calcium carbonate", in other words, containing lime or being chalky. The term is used in a wide variety of scientific disciplines.
In zoology
''Calcareous'' is used as an a ...
algae, especially, although not entirely,
coralline algae
Coralline algae are red algae in the order Corallinales. They are characterized by a thallus that is hard because of calcareous deposits contained within the cell walls. The colors of these algae are most typically pink, or some other shade of r ...
.
The preferred substrate for oyster larvae is the shells of oysters so they tend to settle on adult oysters and thereby develop layers building upwards, eventually forming a fairly massive hard stony calcium carbonate structure on which other reef organisms like
sponges
Sponges, the members of the phylum Porifera (; meaning 'pore bearer'), are a basal animal clade as a sister of the diploblasts. They are multicellular organisms that have bodies full of pores and channels allowing water to circulate throug ...
and
seaweeds can grow, and provide a habitat for mobile benthic organisms.
These biotic reef types take on additional names depending upon how the reef lies in relation to the land, if any. Reef types include
fringing reefs,
barrier reefs, and
atoll
An atoll () is a ring-shaped island, including a coral rim that encircles a lagoon partially or completely. There may be coral islands or cays on the rim. Atolls are located in warm tropical or subtropical oceans and seas where corals can ...
s. A fringing reef is a reef that is attached to an island. A barrier reef forms a calcareous barrier around an island, resulting in a
lagoon
A lagoon is a shallow body of water separated from a larger body of water by a narrow landform, such as reefs, barrier islands, barrier peninsulas, or isthmuses. Lagoons are commonly divided into '' coastal lagoons'' (or ''barrier lagoons' ...
between the shore and the reef. An atoll is a ring reef with no land present. The reef front (ocean side) is a high energy locale, whereas the internal lagoon will be at a lower energy with fine grained sediments.
Mounds
One definition distinguishes reefs from mounds as follows: both are considered to be varieties of organosedimentary buildups – sedimentary features, built by the interaction of organisms and their environment, that have synoptic relief and whose biotic composition differs from that found on and beneath the surrounding
sea floor
The seabed (also known as the seafloor, sea floor, ocean floor, and ocean bottom) is the bottom of the ocean. All floors of the ocean are known as 'seabeds'.
The structure of the seabed of the global ocean is governed by plate tectonics. Most of ...
. Reefs are held up by a macroscopic skeletal framework. Coral reefs are an example of this kind.
Corals
Corals are marine invertebrates within the class Anthozoa of the phylum Cnidaria. They typically form compact colonies of many identical individual polyps. Coral species include the important reef builders that inhabit tropical oceans and secre ...
and calcareous algae grow on top of one another and form a three-dimensional framework that is modified in various ways by other organisms and inorganic processes. By contrast, mounds lack a macroscopic skeletal framework. Mounds are built by microorganisms or by organisms that don't grow a skeletal framework. A microbial mound might be built exclusively or primarily by
cyanobacteria. Examples of
biostromes formed by cyanobacteria occur in the
Great Salt Lake in
Utah
Utah ( , ) is a state in the Mountain West subregion of the Western United States. Utah is a landlocked U.S. state bordered to its east by Colorado, to its northeast by Wyoming, to its north by Idaho, to its south by Arizona, and to its ...
, United States, and in
Shark Bay
Shark Bay ( Malgana: ''Gathaagudu'', "two waters") is a World Heritage Site in the Gascoyne region of Western Australia. The http://www.environment.gov.au/heritage/places/world/shark-bay area is located approximately north of Perth, on the ...
on the coast of
Western Australia
Western Australia (commonly abbreviated as WA) is a state of Australia occupying the western percent of the land area of Australia excluding external territories. It is bounded by the Indian Ocean to the north and west, the Southern Ocean to ...
.
Cyanobacteria do not have skeletons, and individuals are microscopic. Cyanobacteria can encourage the precipitation or accumulation of calcium carbonate to produce distinct sediment bodies in composition that have relief on the seafloor. Cyanobacterial mounds were most abundant before the evolution of shelly macroscopic organisms, but they still exist today;
stromatolites are microbial mounds with a laminated internal structure.
Bryozoans and
crinoid
Crinoids are marine animals that make up the Class (biology), class Crinoidea. Crinoids that are attached to the sea bottom by a stalk in their adult form are commonly called sea lilies, while the unstalked forms are called feather stars or coma ...
s, common contributors to marine sediments during the
Mississippian
Mississippian may refer to:
* Mississippian (geology), a subperiod of the Carboniferous period in the geologic timescale, roughly 360 to 325 million years ago
*Mississippian culture, a culture of Native American mound-builders from 900 to 1500 AD ...
, for instance, produced a very different kind of mound. Bryozoans are small and the skeletons of crinoids disintegrate. However, bryozoan and crinoid meadows can persist over time and produce compositionally distinct bodies of sediment with depositional relief.
The
Proterozoic
The Proterozoic () is a geological eon spanning the time interval from 2500 to 538.8million years ago. It is the most recent part of the Precambrian "supereon". It is also the longest eon of the Earth's geologic time scale, and it is subdivided ...
Belt Supergroup contains evidence of possible
microbial mat and dome structures similar to stromatolite and chicken reef complexes.
Geologic
Rocky reefs are underwater outcrops of rock projecting above the adjacent unconsolidated surface with varying relief. They can be found in depth ranges from
intertidal to deep water, and provide a substrate for a large range of sessile benthic organisms, and shelter for a large range of mobile organisms.
Structures

Ancient reefs buried within
stratigraphic
Stratigraphy is a branch of geology concerned with the study of rock layers ( strata) and layering (stratification). It is primarily used in the study of sedimentary and layered volcanic rocks.
Stratigraphy has three related subfields: lithostr ...
sections are of considerable interest to
geologist
A geologist is a scientist who studies the solid, liquid, and gaseous matter that constitutes Earth and other terrestrial planets, as well as the processes that shape them. Geologists usually study geology, earth science, or geophysics, alt ...
s because they provide paleo-environmental information about the location in
Earth's history. In addition, reef structures within a sequence of
sedimentary rock
Sedimentary rocks are types of rock that are formed by the accumulation or deposition of mineral or organic particles at Earth's surface, followed by cementation. Sedimentation is the collective name for processes that cause these particle ...
s provide a discontinuity which may serve as a trap or conduit for
fossil fuels or mineralizing fluids to form
petroleum
Petroleum, also known as crude oil, or simply oil, is a naturally occurring yellowish-black liquid mixture of mainly hydrocarbons, and is found in geological formations. The name ''petroleum'' covers both naturally occurring unprocessed crude ...
or
ore deposits.
Corals, including some major extinct groups
Rugosa and
Tabulata, have been important reef builders through much of the
Phanerozoic
The Phanerozoic Eon is the current geologic eon in the geologic time scale, and the one during which abundant animal and plant life has existed. It covers 538.8 million years to the present, and it began with the Cambrian Period, when anima ...
since the
Ordovician
The Ordovician ( ) is a geologic period and system, the second of six periods of the Paleozoic Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period million years ago (Mya) to the start of the Silurian Period Mya. ...
Period. However, other organism groups, such as calcifying algae, especially members of the red algae (
Rhodophyta), and molluscs (especially the
rudist bivalves during the
Cretaceous
The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of ...
Period) have created massive structures at various times. During the
Cambrian
The Cambrian Period ( ; sometimes symbolized Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million years ag ...
Period, the conical or tubular skeletons of
Archaeocyatha, an extinct group of uncertain affinities (possibly sponges), built reefs. Other groups, such as the Bryozoa, have been important interstitial organisms, living between the framework builders. The corals which build reefs today, the
Scleractinia
Scleractinia, also called stony corals or hard corals, are marine animals in the phylum Cnidaria that build themselves a hard skeleton. The individual animals are known as polyps and have a cylindrical body crowned by an oral disc in which a mo ...
, arose after the
Permian–Triassic extinction event
The Permian–Triassic (P–T, P–Tr) extinction event, also known as the Latest Permian extinction event, the End-Permian Extinction and colloquially as the Great Dying, formed the boundary between the Permian and Triassic geologic periods, a ...
that wiped out the earlier rugose corals (as well as many other groups), and became increasingly important reef builders throughout the
Mesozoic
The Mesozoic Era ( ), also called the Age of Reptiles, the Age of Conifers, and colloquially as the Age of the Dinosaurs is the second-to-last era of Earth's geological history, lasting from about , comprising the Triassic, Jurassic and Creta ...
Era. They may have arisen from a rugose coral ancestor. Rugose corals built their skeletons of
calcite
Calcite is a carbonate mineral and the most stable polymorph of calcium carbonate (CaCO3). It is a very common mineral, particularly as a component of limestone. Calcite defines hardness 3 on the Mohs scale of mineral hardness, based on scra ...
and have a different symmetry from that of the scleractinian corals, whose skeletons are
aragonite
Aragonite is a carbonate mineral, one of the three most common naturally occurring crystal forms of calcium carbonate, (the other forms being the minerals calcite and vaterite). It is formed by biological and physical processes, including pre ...
. However, there are some unusual examples of well-preserved aragonitic rugose corals in the
Late Permian. In addition, calcite has been reported in the initial post-larval calcification in a few scleractinian corals. Nevertheless, scleractinian corals (which arose in the middle Triassic) may have arisen from a non-calcifying ancestor independent of the rugosan corals (which disappeared in the late Permian).
Artificial
An artificial reef is a human-created underwater structure, typically built to promote
marine life
Marine life, sea life, or ocean life is the plants, animals and other organisms that live in the salt water of seas or oceans, or the brackish water of coastal estuaries. At a fundamental level, marine life affects the nature of the plane ...
in areas with a generally featureless bottom, to control erosion, block ship passage, block the use of
trawling
Trawling is a method of fishing that involves pulling a fishing net through the water behind one or more boats. The net used for trawling is called a trawl. This principle requires netting bags which are towed through water to catch different spec ...
nets,
or improve
surfing.
Many reefs are built using objects that were built for other purposes, for example by sinking oil rigs (through the
Rigs-to-Reefs program),
scuttling ships, or by deploying
rubble or
construction debris. Other artificial reefs are purpose built (e.g. the
reef balls
Reef Ball Foundation, Inc. is a 501(c)(3) non-profit organization that functions as an international environmental non-governmental organization. The foundation uses reef ball artificial reef technology, combined with coral propagation, transpla ...
) from
PVC or concrete. Shipwrecks become artificial reefs on the seafloor. Regardless of construction method, artificial reefs generally provide stable hard surfaces where
algae
Algae ( , ; : alga ) are any of a large and diverse group of photosynthetic, eukaryotic organisms. The name is an informal term for a polyphyletic grouping that includes species from multiple distinct clades. Included organisms range from ...
and invertebrates such as
barnacles, corals, and
oyster
Oyster is the common name for a number of different families of salt-water bivalve molluscs that live in marine or brackish habitats. In some species, the valves are highly calcified, and many are somewhat irregular in shape. Many, but not a ...
s attach; the accumulation of attached marine life in turn provides intricate structure and food for
assemblages of fish.
See also
*
*
*
*
References
Sources
* Shears N.T. (2007) Biogeography, community structure and biological habitat types of subtidal reefs on the South Island West Coast, New Zealand. ''Science for Conservation 281''. p 53. Department of Conservation, New Zealand
External links
''Reef Rescue'' - Smithsonian Ocean PortalCoral Reefs of the Tropics: facts, photos and movies from
The Nature Conservancy
The Nature Conservancy (TNC) is a global environmental organization headquartered in Arlington, Virginia. it works via affiliates or branches in 79 countries and territories, as well as across every state in the US.
Founded in 1951, The Natu ...
NOAA Photo LibraryReef Environmental Education Foundation NOS Data Explorer- A portal to obtain NOAA National Ocean Service data
Atolls – Distribution, Development and Architecture
{{Authority control
*
Stratigraphy
Coastal geography
Coastal and oceanic landforms