Red-capped Cardinal
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The red-capped cardinal (''Paroaria gularis'') is a small species of
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...
in the
tanager The tanagers (singular ) comprise the bird family Thraupidae, in the order Passeriformes. The family has a Neotropical distribution and is the second-largest family of birds. It represents about 4% of all avian species and 12% of the Neotrop ...
family
Thraupidae The tanagers (singular ) comprise the bird family Thraupidae, in the order Passeriformes. The family has a Neotropical distribution and is the second-largest family of birds. It represents about 4% of all avian species and 12% of the Neotrop ...
. It is found in South America.


Taxonomy

In 1760 the French zoologist
Mathurin Jacques Brisson Mathurin Jacques Brisson (; 30 April 1723 – 23 June 1806) was a French zoologist and natural philosopher. Brisson was born at Fontenay-le-Comte. The earlier part of his life was spent in the pursuit of natural history; his published wo ...
included a description of the red-capped cardinal in the supplement to his ''Ornithologie'' based on a specimen collected in "America". He used the French name ''Le cardinal d'Amérique'' and the Latin name ''Cardinalis americanis''. The two stars (**) at the start of the paragraph indicates that Brisson based his description on the examination of a specimen. Although Brisson coined Latin names, these do not conform to the
binomial system The binomial system ( es, Sistema binominal) is a voting system that was used in the legislative elections of Chile between 1989 and 2013. From an electoral system point of view, the binomial system is in effect the D'Hondt method with an ope ...
and are not recognised by the
International Commission on Zoological Nomenclature The International Commission on Zoological Nomenclature (ICZN) is an organization dedicated to "achieving stability and sense in the scientific naming of animals". Founded in 1895, it currently comprises 26 commissioners from 20 countries. Orga ...
. When in 1766 the Swedish naturalist
Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, ...
updated his ''
Systema Naturae ' (originally in Latin written ' with the ligature æ) is one of the major works of the Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the system, now known as binomial nom ...
'' for the twelfth edition he added 240 species that had been previously described by Brisson in his ''Ornithologie''. One of these was the red-capped cardinal. Linnaeus included a terse description, coined the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''Tanagra gularis'' and cited Brisson's work. The specific name ''gularis'' is Latin for "of the throat". The
type locality Type locality may refer to: * Type locality (biology) * Type locality (geology) See also * Local (disambiguation) * Locality (disambiguation) {{disambiguation ...
has been restricted to French Guiana. This species is now placed in the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
''
Paroaria ''Paroaria'', the red-headed cardinals or cardinal-tanagers (as they are not close to the Cardinalidae), are a genus of tanagers. They were until recently placed in the family Emberizidae. Five or six species are placed here. They are all ver ...
'' that was introduced by the French ornithologist
Charles Lucien Bonaparte Charles Lucien Jules Laurent Bonaparte, 2nd Prince of Canino and Musignano (24 May 1803 – 29 July 1857), was a French naturalist and ornithologist. Lucien and his wife had twelve children, including Cardinal Lucien Bonaparte. Life and career ...
in 1832. The genus ''Paroaria'' is classified in the
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Thraupidae The tanagers (singular ) comprise the bird family Thraupidae, in the order Passeriformes. The family has a Neotropical distribution and is the second-largest family of birds. It represents about 4% of all avian species and 12% of the Neotrop ...
, unlike the cardinals proper which are in the
Cardinalidae Cardinalidae (often referred to as the "cardinal-grosbeaks" or simply the "cardinals") is a family of New World-endemic passerine birds that consists of cardinals, grosbeaks, and buntings. It also includes several birds such as the tanager-lik ...
. The masked cardinal was formerly considered
conspecific Biological specificity is the tendency of a characteristic such as a behavior or a biochemical variation to occur in a particular species. Biochemist Linus Pauling stated that "Biological specificity is the set of characteristics of living organ ...
with the red-capped cardinal. The masked cardinal markedly differs from the red-capped cardinal by the black ear-coverts and the red (not black) lower throat and upper chest. Additionally, its lower mandible is typically whitish, but this is not entirely consistent, as it occasionally is pale flesh-coloured. This distinctive
taxon In biology, a taxon ( back-formation from '' taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular n ...
was determined a different
species In biology, a species is the basic unit of Taxonomy (biology), classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of ...
, as suggested by the lack of
hybridization Hybridization (or hybridisation) may refer to: *Hybridization (biology), the process of combining different varieties of organisms to create a hybrid *Orbital hybridization, in chemistry, the mixing of atomic orbitals into new hybrid orbitals *Nu ...
with ''P. g. gularis'' in the limited area of overlap in southern Venezuela. There are two subspecies: * ''P. g. gularis'' (Linnaeus, 1766) (eastern Colombia to eastern
Peru , image_flag = Flag of Peru.svg , image_coat = Escudo nacional del Perú.svg , other_symbol = Great Seal of the State , other_symbol_type = Seal (emblem), National seal , national_motto = "Fi ...
and through the
Guianas The Guianas, sometimes called by the Spanish loan-word ''Guayanas'' (''Las Guayanas''), is a region in north-eastern South America which includes the following three territories: * French Guiana, an overseas department and region of France * ...
to central
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
) * ''P. g. cervicalis'' Sclater, PL, 1862 (north east
Bolivia , image_flag = Bandera de Bolivia (Estado).svg , flag_alt = Horizontal tricolor (red, yellow, and green from top to bottom) with the coat of arms of Bolivia in the center , flag_alt2 = 7 × 7 square p ...
and south west
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...


Description

The adult red-capped cardinal is long and weighs about . The
nominate subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. Not all spec ...
has a crimson head, blackish lores and ocular region, and shiny black upperparts, apart from a white partial collar extending up the neck sides from the white underparts. The throat is black, extending to a point on the upper chest. The upper mandible is black, while the lower is pale flesh-coloured. The legs are dark grey (almost black) and the
iris Iris most often refers to: *Iris (anatomy), part of the eye *Iris (mythology), a Greek goddess * ''Iris'' (plant), a genus of flowering plants * Iris (color), an ambiguous color term Iris or IRIS may also refer to: Arts and media Fictional ent ...
is brownish-orange. In pattern the juvenile resemble the adults, but the upperparts are dusky-brown, the head is deep brownish-buff (darker on the cap), the bill is entirely black and the iris is pale, dull creamy-yellow. The song is a variable, often repeated series of ''suweet-chu'' notes, and the call is a sharp ''chep''.


Distribution and habitat

It occurs in lowlands of the
Guianas The Guianas, sometimes called by the Spanish loan-word ''Guayanas'' (''Las Guayanas''), is a region in north-eastern South America which includes the following three territories: * French Guiana, an overseas department and region of France * ...
, Venezuela, eastern Colombia, eastern
Ecuador Ecuador ( ; ; Quechuan languages, Quechua: ''Ikwayur''; Shuar language, Shuar: ''Ecuador'' or ''Ekuatur''), officially the Republic of Ecuador ( es, República del Ecuador, which literally translates as "Republic of the Equator"; Quechuan ...
, eastern
Peru , image_flag = Flag of Peru.svg , image_coat = Escudo nacional del Perú.svg , other_symbol = Great Seal of the State , other_symbol_type = Seal (emblem), National seal , national_motto = "Fi ...
, northern and eastern
Bolivia , image_flag = Bandera de Bolivia (Estado).svg , flag_alt = Horizontal tricolor (red, yellow, and green from top to bottom) with the coat of arms of Bolivia in the center , flag_alt2 = 7 × 7 square p ...
and the
Amazon basin The Amazon basin is the part of South America drained by the Amazon River and its tributaries. The Amazon drainage basin covers an area of about , or about 35.5 percent of the South American continent. It is located in the countries of Boli ...
in
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
. In Brazil it is, except for populations in the relatively open lowlands of north-eastern
Roraima Roraima (, ) is one of the 26 states of Brazil. Located in the country's North Region, it is the northernmost and most geographically and logistically isolated state in Brazil. It is bordered by the state of Pará to the southeast, Amazonas ...
and along the
Branco River The Branco River ( pt, Rio Branco; Engl: ''White River'') is the principal affluent of the Rio Negro from the north. Basin The river drains the Guayanan Highlands moist forests ecoregion. It is enriched by many streams from the Tepui highlands ...
and lower Rio Negro, apparently largely absent from the regions north of the northern bank of the Amazon River, but these regions are generally very poorly known and its presence cannot be discounted, especially in the section between the lower Rio Negro and
Rio Jari The Jari River, or Jary River ( pt, Rio Jari), is a northern tributary of the Amazon River on the border between the states of Pará and Amapá in northeastern Brazil. It is in the most downstream regions of the Amazon Basin and borders the Guiana ...
(the river on the border between
Pará Pará is a state of Brazil, located in northern Brazil and traversed by the lower Amazon River. It borders the Brazilian states of Amapá, Maranhão, Tocantins, Mato Grosso, Amazonas and Roraima. To the northwest are the borders of Guyana a ...
and
Amapá Amapá () is one of the 26 states of Brazil. It is in the northern region of Brazil. It is the second least populous state and the eighteenth largest by area. Located in the far northern part of the country, Amapá is bordered clockwise by Fr ...
). This is a bird of swamps,
mangrove A mangrove is a shrub or tree that grows in coastal saline or brackish water. The term is also used for tropical coastal vegetation consisting of such species. Mangroves are taxonomically diverse, as a result of convergent evolution in several ...
, várzea and other semi-open areas near water. It is generally common, and even occurs in lightly wooded wet habitats in towns or cities (e.g. Manaus and
Puerto Maldonado Puerto Maldonado () is a city in southeastern Peru in the Amazon rainforest west of the Bolivian border, located at the confluence of the Tambopata and Madre de Dios rivers. The latter river joins the Madeira River as a tributary of the Amazon ...
).


Behaviour

The red-capped cardinal feeds on
insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (head, thorax and abdomen), three pairs ...
s,
rice Rice is the seed of the grass species '' Oryza sativa'' (Asian rice) or less commonly '' Oryza glaberrima'' (African rice). The name wild rice is usually used for species of the genera '' Zizania'' and ''Porteresia'', both wild and domestica ...
and fruit. It is usually found in conspicuous pairs or family groups. If there is a well-marked breeding season, it extends over much of the year. The red-capped cardinal has been recorded to breed at least from June to September in northern South America but at
Sacha Lodge Sasha is a unisex name which originated in Eastern and Southern European countries as the shortened version of Alexander and Alexandra. It is also used as a surname, although very rarely. Alternative spellings include: ( – Russian, Ukrainian ...
at the
Napo River The Napo River ( es, Río Napo) is a tributary to the Amazon River that rises in Ecuador on the flanks of the east Andean volcanoes of Antisana, Sincholagua and Cotopaxi. The total length is . The river drains an area of . The mean annual d ...
(
Ecuador Ecuador ( ; ; Quechuan languages, Quechua: ''Ikwayur''; Shuar language, Shuar: ''Ecuador'' or ''Ekuatur''), officially the Republic of Ecuador ( es, República del Ecuador, which literally translates as "Republic of the Equator"; Quechuan ...
) an adult was seen feeding a fledgling
shiny cowbird The shiny cowbird (''Molothrus bonariensis'') is a passerine bird in the New World family Icteridae. It breeds in most of South America except for dense forests and areas of high altitude such as mountains. Since 1900 the shiny cowbird's range h ...
(a
brood parasite Brood parasites are animals that rely on others to raise their young. The strategy appears among birds, insects and fish. The brood parasite manipulates a host, either of the same or of another species, to raise its young as if it were its ow ...
) in late March. This suggests that breeding can also occur as early as February/March. There is no indication for a difference in breeding seasons between the red-capped and the masked cardinals. They build a shallow open
cup nest A bird nest is the spot in which a bird lays and incubates its eggs and raises its young. Although the term popularly refers to a specific structure made by the bird itself—such as the grassy cup nest of the American robin or Eurasian b ...
, some wide and high outside with a wide by deep cup, in a tree or some other secure spot. In
Ecuador Ecuador ( ; ; Quechuan languages, Quechua: ''Ikwayur''; Shuar language, Shuar: ''Ecuador'' or ''Ekuatur''), officially the Republic of Ecuador ( es, República del Ecuador, which literally translates as "Republic of the Equator"; Quechuan ...
, a nest in the
Cuyabeno Wildlife Reserve The Cuyabeno Wildlife Reserve ( es, Reserva de Producción Faunística Cuyabeno) is the second largest reserve of the 56 national parks and protected areas in Ecuador. It is located in the Putumayo Canton in the Sucumbíos Province and in the ...
was built in the roots of a ''
Macrolobium ''Macrolobium'' is a legume genus in the subfamily Detarioideae. It is a tropical genus with about 80 species. Half occur in Brazil, where they are common in the floodplains of the Amazonian Basin. Members of the genus are used as ornamentals an ...
'' sp. standing in water, while one at
Sacha Lodge Sasha is a unisex name which originated in Eastern and Southern European countries as the shortened version of Alexander and Alexandra. It is also used as a surname, although very rarely. Alternative spellings include: ( – Russian, Ukrainian ...
was above ground in a tree. For nest material, it uses rootlets, thin twigs, and ferns. The
clutch A clutch is a mechanical device that engages and disengages power transmission, especially from a drive shaft to a driven shaft. In the simplest application, clutches connect and disconnect two rotating shafts (drive shafts or line shafts ...
is two or three eggs. These have a background colour varying between whitish and dull olive and have dense dark brown flecks and blotches, heavier at the blunt end. They measure about . This species is parasitised by the
shiny cowbird The shiny cowbird (''Molothrus bonariensis'') is a passerine bird in the New World family Icteridae. It breeds in most of South America except for dense forests and areas of high altitude such as mountains. Since 1900 the shiny cowbird's range h ...
(''Molothrus bonariensis'').


References


Further reading

* ffrench, Richard; O'Neill, John Patton & Eckelberry, Don R. (1991): ''A guide to the birds of Trinidad and Tobago'' (2nd edition). Comstock Publishing, Ithaca, N.Y. * Hilty, Steven L. (2003): ''Birds of Venezuela''.
Christopher Helm Christopher Alexander Roger Helm (born Dundee, 1 February 1937 – 20 January 2007) was a Scottish book publisher, notably of ornithology related titles, including the ''Helm Identification Guides''. Born in Dundee, he was raised in Forfar, ...
, London. * Restall, Robin L.; Rodner, C. & Lentino, M. (2006): ''Birds of Northern South America''.
Christopher Helm Christopher Alexander Roger Helm (born Dundee, 1 February 1937 – 20 January 2007) was a Scottish book publisher, notably of ornithology related titles, including the ''Helm Identification Guides''. Born in Dundee, he was raised in Forfar, ...
, London. (vol. 1). (vol. 2).


External links


Cardinal videos, photos & sounds
on the Internet Bird Collection

VIREO ttp://vireo.acnatsci.org/species_image.php?species=Paroaria+gularis Photo {{Taxonbar, from=Q2668125 red-capped cardinal Birds of the Amazon Basin Birds of Venezuela Birds of the Guianas red-capped cardinal red-capped cardinal Birds of Brazil