Ranger's Toad
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The raucous toad (''Sclerophrys capensis''), also known as Ranger's toad, is a species of
toad Toad (also known as a hoptoad) is a common name for certain frogs, especially of the family Bufonidae, that are characterized by dry, leathery skin, short legs, and large bumps covering the parotoid glands. In popular culture (folk taxonomy ...
from
Southern Africa Southern Africa is the southernmost region of Africa. No definition is agreed upon, but some groupings include the United Nations geoscheme for Africa, United Nations geoscheme, the intergovernmental Southern African Development Community, and ...
.Du Preez, L. H., Carruthers, Vincent; A complete guide to the frogs of southern Africa. Pub: Cape Town, South Africa : Struik Nature, 2009


Taxonomy

''Sclerophrys capensis'' is a member of the family
Bufonidae {{Cat main, Toad This category Category, plural categories, may refer to: General uses *Classification, the general act of allocating things to classes/categories Philosophy * Category of being * ''Categories'' (Aristotle) * Category (Kant) * C ...
. This species was described as ''Bufo regularis rangeri'', in 1935, by John Hewitt of the Albany Museum, Grahamstown, from collections made by naturalist Gordon A. Ranger at his farm "Gleniffer" 3 km E of Kei Road, in the
Eastern Cape The Eastern Cape ( ; ) is one of the nine provinces of South Africa. Its capital is Bhisho, and its largest city is Gqeberha (Port Elizabeth). Due to its climate and nineteenth-century towns, it is a common location for tourists. It is also kno ...
province of
South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. Its Provinces of South Africa, nine provinces are bounded to the south by of coastline that stretches along the Atlantic O ...
. Among several other forms described from these collections was the eastern leopard toad, ''Sclerophrys pardalis'', which Ranger distinguished from the raucous toad on behaviour and calls (according to his nephew, Mr Gary Ranger, "Gleniffer" was subdivided in the late 1930s and the portions renamed, including "Hillside" and "Rockdale", where there is a monument to the older Mr Ranger). Hewitt considered the raucous toad, along with most other large toads of Southern Africa, as subspecies of the widespread African toad ''Bufo regularis'' (a species which is now considered absent from Southern Africa, predominantly occurring in subsaharan Africa north of the equator). In 1964, Poynton raised ''Bufo rangeri'' of Hewitt, 1935 to full species status, distinguishing it from co-occurring related forms. More recently, Frost and co-authors (2006) assigned this species to the new genus ''Amietophrynus'', as part of a global revision of amphibian taxonomy based largely on DNA sequence data. Ohler and Dubois (2016) studied the type (and, previously, the only known) specimen of ''Sclerophrys capensis'', collected in the
Cape Province The Province of the Cape of Good Hope (), commonly referred to as the Cape Province () and colloquially as The Cape (), was a province in the Union of South Africa and subsequently the Republic of South Africa. It encompassed the old Cape Co ...
and described by
Johann Jakob von Tschudi Johann Jakob von Tschudi (25 July 1818 – 8 October 1889) was a Switzerland, Swiss naturalist, explorer, and diplomat. He is known for his travels in South America, his scientific contributions to zoology and anthropology, and his diplomatic se ...
in 1838, and discovered that it belonged to the present species, then known as ''Amietophrynus rangeri''. Accordingly, following the
Principle of Priority Priority is a principle in Taxonomy (biology), biological taxonomy by which a valid scientific name is established based on the oldest available name. It is a decisive rule in Botanical nomenclature, botanical and zoological nomenclature to recogn ...
of the
ICZN Code The International Code of Zoological Nomenclature (ICZN) is a widely accepted convention in zoology that rules the formal scientific naming of organisms treated as animals. It is also informally known as the ICZN Code, for its formal author, t ...
, ''
Sclerophrys ''Sclerophrys'' is a genus of "true toads", family Bufonidae, native to Africa and the southern Arabian Peninsula. Originally, all of these species were classified in the genus ''Bufo''. The genus, originally named ''Amietophrynus'', was split du ...
'' replaces ''Amietophrynus'', and ''S. capensis'' replaces ''A. rangeri'', as the valid name for this genus and species, respectively.


Description

Its maximal size, snout-to-vent, is about 120 mm. It is a characteristically robust medium-large-sized toad,
dorsal Dorsal (from Latin ''dorsum'' ‘back’) may refer to: * Dorsal (anatomy), an anatomical term of location referring to the back or upper side of an organism or parts of an organism * Dorsal, positioned on top of an aircraft's fuselage The fus ...
and
lateral Lateral is a geometric term of location which may also refer to: Biology and healthcare * Lateral (anatomy), a term of location meaning "towards the side" * Lateral cricoarytenoid muscle, an intrinsic muscle of the larynx * Lateral release ( ...
skin liberally sprinkled with wart-like protuberances. Colour brown-to-greyish with more-or-less paired dark-brown patches flanking the
medial Medial may refer to: Mathematics * Medial magma, a mathematical identity in algebra Geometry * Medial axis, in geometry the set of all points having more than one closest point on an object's boundary * Medial graph, another graph that repr ...
dorsal line. The dark patches on the head meet in a brown chevron on top of the head behind and between the eyes. The
anterior Standard anatomical terms of location are used to describe unambiguously the anatomy of humans and other animals. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position pro ...
dorsal skin of the head lacks dark brown patches. Like most typical toads, it has a large parotoid gland behind each eye. If injured it will exude whitish spots of venom on the parotoid glands; this can be hazardous to domestic dogs if they bite a toad. The underside is whitish with the gular (throat) area of males infused with yellow with a dark blue-black bar posterior to the gular.


Call

The advertisement call, made by males in the breeding season, is a loud repetitive duck-like quacking. Groups of calling males generally space themselves along the water's edge, or among plants in the water. Males in chorus call antiphonally (with calls from different individuals alternating).


Habits

It is fairly aquatic as toads go, commonly inhabiting ponds and dams and streams, though it seems to prefer running water and accordingly favours fountains and similar water bodies. It typically is shy, but like many toads it will visit houses and other places where insect prey are attracted to lights, mainly outside the breeding season. It then establishes itself in moist, sheltered spots such as behind flowerpots and becomes tame if not molested. It will eat practically any small animal, including small vertebrates, but mainly insects up to the size of crickets.


Range

''S. capensis'' occurs in
South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. Its Provinces of South Africa, nine provinces are bounded to the south by of coastline that stretches along the Atlantic O ...
,
Lesotho Lesotho, formally the Kingdom of Lesotho and formerly known as Basutoland, is a landlocked country in Southern Africa. Entirely surrounded by South Africa, it is the largest of only three sovereign enclave and exclave, enclaves in the world, t ...
,
Eswatini Eswatini, formally the Kingdom of Eswatini, also known by its former official names Swaziland and the Kingdom of Swaziland, is a landlocked country in Southern Africa. It is bordered by South Africa on all sides except the northeast, where i ...
, extreme southern Namibia, and possibly extreme southern
Botswana Botswana, officially the Republic of Botswana, is a landlocked country in Southern Africa. Botswana is topographically flat, with approximately 70 percent of its territory part of the Kalahari Desert. It is bordered by South Africa to the sou ...
,
Mozambique Mozambique, officially the Republic of Mozambique, is a country located in Southeast Africa bordered by the Indian Ocean to the east, Tanzania to the north, Malawi and Zambia to the northwest, Zimbabwe to the west, and Eswatini and South Afr ...
, and
Zimbabwe file:Zimbabwe, relief map.jpg, upright=1.22, Zimbabwe, relief map Zimbabwe, officially the Republic of Zimbabwe, is a landlocked country in Southeast Africa, between the Zambezi and Limpopo Rivers, bordered by South Africa to the south, Bots ...
.


Habitat

''S. capensis'' occurs in
fynbos Fynbos (; , ) is a small belt of natural shrubland or heathland vegetation located in the Western Cape and Eastern Cape provinces of South Africa. The area is predominantly coastal and mountainous, with a Mediterranean climate. The fynbos ...
heathland, grassland, dry thicket forest, savanna, and agricultural land. Breeding takes place in dams, ponds, and pools along slow-forming streams. It shows some preference for permanent water bodies.


References

{{Taxonbar, from=Q25916769, from2=Q1946058 Sclerophrys Frogs of Africa Amphibians of Eswatini Vertebrates of Lesotho Amphibians of Namibia Amphibians of South Africa Taxa named by Johann Jakob von Tschudi Amphibians described in 1838 Taxonomy articles created by Polbot