Sir Ralph Riley (23 October 1924 – 27 August 1999) was a British geneticist. He was born in Scarborough,

North Yorkshire North Yorkshire is the largest ceremonial county (lieutenancy area) in England, covering an area of . Around 40% of the county is covered by national parks, including most of the Yorkshire Dales and the North York Moors. It is one of four cou ...

in 1924 and served in the army during the Second World War. After the war he studied Botany at

Sheffield University , mottoeng = To discover the causes of things , established = – University of SheffieldPredecessor institutions: – Sheffield Medical School – Firth College – Sheffield Technical School – University College of Sheffield , type = Pu ...

, followed by a two-year PhD study in genetics. He was then recruited by the Plant Breeding Institute (PBI) at Cambridge to study the introduction of useful variation into the wheat crop from its wild relatives. Two years later in 1954 Riley became the founder and first Head of the Cytogenetics Department at the PBI. His target was to increase the wheat gene pool by making the variation in wild relatives available to wheat breeders. In 1957, he discovered the method of doing so by finding the Ph gene. This gene controlled the pairing between the chromosomes of wheat and wild relatives of wheat and soon he was able to demonstrate the cytogenetic ways by which useful genes, such as those that confer novel disease resistances, could be transferred into wheat from a host of wild species. This discovery of the Ph gene allowed the first "genetic engineering" and his methods have since been used around the world in all major cereal breeding programmes. In 1972, he became Director of the PBI and during his six years as Director strove to improve production in UK arable agriculture, developing fundamental research programmes on breeding and introducing plant molecular biology in the UK. He thus ensured that the PBI's pre-eminence in the application of science to plant breeding. During his directorship wheat yields increased from four tonnes per hectare to 6 t/ha, due in large part to the improved PBI varieties. This was particularly vital at a time when the UK needed to be less reliant on North American imports. In 1978 Ralph Riley left the PBI to become Secretary (chief executive) of the Agriculture and Food Research Council (now the Biotechnology and Biological Sciences Research Council), where he served as Secretary for seven years and Deputy-Chairman for a further two. He was elected as a

Fellow of the Royal Society Fellowship of the Royal Society (FRS, ForMemRS and HonFRS) is an award granted by the judges of the Royal Society of London to individuals who have made a "substantial contribution to the improvement of natural knowledge, including mathematic ...

in 1967. He was awarded the

William Bate Hardy Prize The William Bate Hardy Prize is awarded by the Cambridge Philosophical Society. It is awarded once in three years “for the best original memoir, investigation or discovery by a member of the University of Cambridge in connection with Biological S ...

in 1969 and the

Royal Medal The Royal Medal, also known as The Queen's Medal and The King's Medal (depending on the gender of the monarch at the time of the award), is a silver-gilt medal, of which three are awarded each year by the Royal Society, two for "the most important ...

in 1981. and the

Wolf Foundation Prize The Wolf Prize is an international award granted in Israel, that has been presented most years since 1978 to living scientists and artists for ''"achievements in the interest of mankind and friendly relations among people ... irrespective of natio ...

in Agriculture in 1986. He was knighted for his services to science in 1984. He married Joan Norrington in 1949; they had two daughters. He died in Cambridge on 27 August 1999.

References

{{DEFAULTSORT:Riley, Ralph 1924 births 1999 deaths People from Scarborough, North Yorkshire Royal Medal winners English geneticists Fellows of the Royal Society Foreign associates of the National Academy of Sciences Knights Bachelor Plant geneticists Wolf Prize in Agriculture laureates