Rainar Alarik Hakulinen (9 October 1918 – 29 December 1991) was a Finnish
lichenologist and schoolteacher. He was noted as an expert on the lichen family
Candelariaceae
''Candelariaceae'' is a family of lichen-forming fungi in the order Candelariales. It contains seven genera and about 73 species. The family was circumscribed by Finnish lichenologist Rainar Hakulinen in 1954 to contain the type genus
In bio ...
, and was known for his numerous
phytogeographical publications about the
boreal
Boreal may refer to:
Climatology and geography
*Boreal (age), the first climatic phase of the Blytt-Sernander sequence of northern Europe, during the Holocene epoch
*Boreal climate, a climate characterized by long winters and short, cool to mild ...
and
arctic
The Arctic ( or ) is a polar region located at the northernmost part of Earth. The Arctic consists of the Arctic Ocean, adjacent seas, and parts of Canada ( Yukon, Northwest Territories, Nunavut), Danish Realm ( Greenland), Finland, Iceland ...
lichens of Finland,
Northern Norway
Northern Norway ( nb, Nord-Norge, , nn, Nord-Noreg; se, Davvi-Norga) is a geographical region of Norway, consisting of the two northernmost counties Nordland and Troms og Finnmark, in total about 35% of the Norwegian mainland. Some of the la ...
, and
Russian Karelia
East Karelia ( fi, Itä-Karjala, Karelian: ''Idä-Karjala''), also rendered as Eastern Karelia or Russian Karelia, is a name for the part of Karelia that since the Treaty of Stolbova in 1617 has remained Eastern Orthodox under Russian supremacy. ...
.
Biography
Hakulinen was born in on 9 October 1918. At the time, this area was part of Finland, but is now in the
Republic of Karelia
The Republic of Karelia (russian: Респу́блика Каре́лия, Respublika Kareliya; ; krl, Karjalan tašavalta; ; fi, Karjalan tasavalta; vep, Karjalan Tazovaldkund, Ludic: ''Kard’alan tazavald''), also known as just Karelia (ru ...
, a part of Russia. Hakulinen served in the
Finnish Army
The Finnish Army ( Finnish: ''Maavoimat'', Swedish: ''Armén'') is the land forces branch of the Finnish Defence Forces. The Finnish Army is divided into six branches: the infantry (which includes armoured units), field artillery, anti-aircra ...
in its
war against Russia in 1941–1944, spending a lot of his time there on the frontier at the
Svir River
The Svir (, Veps language, Veps: , Karelian language, Karelian/Finnish language, Finnish: ) is a river in Podporozhsky District, Podporozhsky, Lodeynopolsky District, Lodeynopolsky, and Volkhovsky District, Volkhovsky districts in the north-eas ...
. He spent several months in
military hospital
A military hospital is a hospital owned and operated by a military. They are often reserved for the use of military personnel and their dependents, but in some countries are made available to civilians as well. They may or may not be located on a ...
s to heal his wounds.
In his school years, Hakulinen knew the lichenologist
Veli Räsänen
Veli Johannes Paavo Bartholomeus Räsänen (24 August 1888 – 16 July 1953) was a Finnish lichenologist.
Early life and education
Veli Räsänen was born on 24 August 1888 in Simo, Finland. Räsänen's brother was the linguist Martti Räs� ...
, who encouraged his early interest in lichens. Räsäsen had lived in the area for years and had studied its lichen flora quite thoroughly; Hakulinen accompanied him on field trips. Hakulinen maintained his interest after entering the
University of Helsinki
The University of Helsinki ( fi, Helsingin yliopisto, sv, Helsingfors universitet, abbreviated UH) is a public research university located in Helsinki, Finland since 1829, but founded in the city of Turku (in Swedish ''Åbo'') in 1640 as the ...
, and took courses in geography and biology.
He received a
magister degree
A magister degree (also magistar, female form: magistra; from la, magister, "teacher") is an academic degree used in various systems of higher education.
The magister degree arose in medieval universities in Europe and was originally equal to th ...
in 1948.
In 1954, he published his
Ph.D.
A Doctor of Philosophy (PhD, Ph.D., or DPhil; Latin: or ') is the most common degree at the highest academic level awarded following a course of study. PhDs are awarded for programs across the whole breadth of academic fields. Because it is a ...
dissertation on the
taxonomy
Taxonomy is the practice and science of categorization or classification.
A taxonomy (or taxonomical classification) is a scheme of classification, especially a hierarchical classification, in which things are organized into groups or types. ...
of the genus ''
Candelariella
''Candelariella'' is a genus of bright yellow, ocher, or greenish yellow crustose or squamulose lichens in the family Candelariaceae. Members of the genus are commonly called eggyolk lichens, goldspeck lichens, or yolk lichens. The genus was ci ...
'', and he established the family
Candelariaceae
''Candelariaceae'' is a family of lichen-forming fungi in the order Candelariales. It contains seven genera and about 73 species. The family was circumscribed by Finnish lichenologist Rainar Hakulinen in 1954 to contain the type genus
In bio ...
.
After finishing university, Hakulinen worked as a
secondary school teacher, first at
Isokyrö
Isokyrö (; sv, Storkyro) is a municipality of Finland. It is located in the South Ostrobothnia region, from Vaasa. The municipality has a population of () and covers an area of of which is water. The population density is .
In the name of t ...
, but soon later in
Hämeenlinna
Hämeenlinna (; sv, Tavastehus; krl, Hämienlinna; la, Tavastum or ''Croneburgum'') is a city and municipality of about inhabitants in the heart of the historical province of Tavastia and the modern province of Kanta-Häme in the south of ...
, where he settled permanently. He studied the distribution of lichens in Finland, visiting widely throughout the country, and published floristic compilations and treatments of various lichen groups with
distribution maps. In 1963 he published a reference work titled ''Jäkäläkasvio'' that compiled all of the Finnish macrolichens, and common names;
it gave the distributions of the lichens more accurately than any predecessor works.
Other research interests included the growth rate of lichens, and the ecology of lichens growing on small islands and lakeshores. After his mentor Räsäsen died in 1953, he took over the editorship of the
exsiccata series (sets of dried
herbarium
A herbarium (plural: herbaria) is a collection of preserved plant specimens and associated data used for scientific study.
The specimens may be whole plants or plant parts; these will usually be in dried form mounted on a sheet of paper (calle ...
specimens) ''Lichenotheca Fennica a Museo Kupoioensi ëdita''; Hakulinen added 500 numbers to the 800 already distributed by Räsäsen.
Hakulinen published his last scientific paper in 1968, and essentially ceased lichenological activity after that. His applications for professorship at the
Universities of Turku and
Oulu
Oulu ( , ; sv, Uleåborg ) is a city, municipality and a seaside resort of about 210,000 inhabitants in the region of North Ostrobothnia, Finland. It is the most populous city in northern Finland and the fifth most populous in the country after ...
were rejected, and he moved on to other interests. He published three books (1972, 1978, 1986) on the history of his home parish Kurkijoki, and two school history books.
In his obituary of Hakulinen, his colleague
Teuvo Ahti
Teuvo ("Ted") Tapio Ahti (born 1934) is a Finnish botanist and lichenologist. He has had a long career at the University of Helsinki that started in 1963, and then following his retirement in 1997, at the Botanical Museum of the Finnish Museum ...
describes him as "an important link in maintaining continuity between the splendid Finnish tradition of lichenology from
William Nylander
William Andrew Michael Junior Nylander Altelius (born 1 May 1996) is a Canadian-born Swedish professional ice hockey right winger for the Toronto Maple Leafs of the National Hockey League (NHL). Nylander was selected by the Maple Leafs in the ...
through
E. A. Vainio and V. Rasanen to the present". Hakulinen died at his home on 29 December 1991.
The lichen ''
Candelariella hakulinenii
''Candelariella'' is a genus of bright yellow, ocher, or greenish yellow crustose or squamulose lichens in the family Candelariaceae. Members of the genus are commonly called eggyolk lichens, goldspeck lichens, or yolk lichens. The genus was cir ...
'' is named in his honour.
Selected publications
Hakulinen's 40 lichenological publications are listed in Ahti's obituary.
Some of his major works include:
*
*
*
*
See also
*
:Taxa named by Rainar Hakulinen
References
{{DEFAULTSORT:Hakulinen, Rainar
1918 births
1991 deaths
Finnish lichenologists
Scientists from Vyborg
Finnish military personnel of World War II
Finnish schoolteachers
20th-century Finnish scientists